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    Bolborhinum geotrupoides Laporte 1840

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    Bolborhinum geotrupoides (Laporte, 1840) (Figs. 1, 2, 15–18) Original combination. Bolboceras geotrupoides Laporte, 1840: 104. Type locality: “ CHILE, Talca Prov., Alto Vilches, 1100 m ” (original type locality: “ Chili ”). Type series: neotype at CMNC labeled: “ CHILE, Talca Prov. / AltoVilches, 1100m. / 10–12.XII. 1976 / H. F. Howden” (typeset), b) “H.&A. HOWDEN / COLLEC- TION / Ottawa, Canada ” (typeset with black border), c) “ BOLBOCERAS / GEOTRUPOIDES / LAPORTE ɗ / NEOTYPE / A.B.T. SMITH ” (red label, handwritten and typeset), d) “Southern Neotropical Scarabs / database # AS 2607270 / Bolborhinum geotrupoides / (Laporte, 1840) ɗ / DET: A.B.T. SMITH 2006 ” (typeset). Neotype here designated. See “Designation of lectotypes and neotypes ” section for further statements of qualifying conditions for the designation of this neotype. All reasonable steps were exhausted to trace the original type material of this taxon. Many Laporte type specimens have been lost, but some are currently housed in the MNHN and in the Museum of Victoria in Abbotsford, Australia (MVMA). The second author searched for type material at the MNHN in May 2005 and Ken Walker (curator of entomology at the MVMA) searched for type material in the MVMA, both without success. We can only conclude that the type material has been lost. The specimen selected as the neotype matches the original description of Laporte (1840) and falls within the original type locality. Synonym. Bolboceras binasutum Fairmaire and Germain, 1861: 2 (emended from Bolboceras binasutus). New Synonymy. Type locality: “Chilensia.” Type series: lectotype at MNHN labeled a) “ 1683 ” (handwritten), b) “ Bolboceras / binasutus ” (handwritten by Fairmaire), c) “MUSEUM PARIS / Collection Léon Fairmaire / 1906 ” (typeset), d) “ TYPE ” (red label, typeset), e) “ BOLBOCERAS / BINASUTUM / FAIRMAIRE & GERMAIN / LECTOTYPE ɗ / A.B.T. SMITH ” (red label, handwritten and typeset), f) “Southern Neotropical Scarabs / database # AS 2609492 / Bolborhinum geotrupoides / (Laporte, 1840) / DET: A.B.T. SMITH 2007 ” (typeset). Lectotype here designated. Synonym: Bolboceras distinguendum Fairmaire and Germain, 1861: 2 (emended from Bolboceras distinguendus). Type locality: “Chilensia.” Type series: lectotype male at MNHN labeled a) “ 1622 ” (handwritten), b) “ Bolboceras / distinguendus ” (handwritten by Fairmaire), c) “MUSEUM PARIS / Collection Léon Fairmaire / 1906 ” (typeset), d) “ TYPE ” (red label, typeset), e) “ BOLBOCERAS / DISTINGUENDUM / FAIRMAIRE & GERMAIN / LECTOTYPE ɗ / A.B.T. SMITH ” (red label, handwritten and typeset), f) “Southern Neotropical Scarabs / database # AS 2609493 / Bolborhinum geotrupoides / (Laporte, 1840) / DET: A.B.T. SMITH 2007 ” (typeset). Lectotype here designated. Specimens examined. 343 specimens were examined from AUPC, CMNC, CNCI, FMNH, GACC, HAHC, INHS, JEBC, JMEC, MEUC, MGAC, MNHN, MNNC, SRTC, UCCC, UMCE, VMDC. Diagnosis. Length 14.5–22.0 mm. Color generally reddish-brown. The number, shape, and placement of the male clypeal horns are diagnostic as follows: head dorsally with two medial horns that are connected by an elevated carina (Fig. 15); the apical horn is more robust and located at the clypeal apex, the basal horn is smaller and located immediately behind the first. The two horns could be interpreted as a single bifid horn (in small individuals the second clypeal horn is replaced by a pair of small tubercles located on each side of the midline). Male pronotum with deep and wide anterior excavation occupying almost two thirds of pronotum; surface smooth, generally impunctate; posterosuperior ridge transverse, slightly arcuate anteriorly, sometimes with lateral edge anteriorly projected in larger specimens. Female similar to male, except in the trapezoidal shape and smaller length of the head in comparison with the male; dorsal surface medially with a small, weakly elevated tubercle and a pair of short lateral carinae that are convergent distally. Clypeus anteriorly truncate, elevated. Pronotum with weak depression located immediately behind the apical margin; disc with fine furrow and sparse punctures on the dorsal surface; laterally with punctures dense to sparse, distributed irregularly across the surface. Distribution (Fig. 56): ARGENTINA (2). Neuquén (1): Lago Tromén (1). Río Negro (1): Pilcaniyeu (1). CHILE (341). Región Metropolitana de Santiago (3): El Monte (1); Naltagua (2). VI Región del General Bernardo O’Higgins (1): Alto Huemul, San Fernando (1). VII Región del Maule (84): Curicó Maritimo (1); El Coigo (1); Fundo El Radal (2); Fundo Malcho (5); La Balsa (4); La Mina (17); Laguna del Maule (2); Lara Bullileo (4); Linares (7); Parque Inglés (2); Parral (1); Potrero Grande (2); Reserva Nacional Radal 7 Tazas (3); Vilches Alto (33). VIII Región del Biobío (190): Abanico (2); Antuco (1); Atacalco (12); Chiguayante (50); Coihueco (1); Cordillera de Chillán (8); Fundo El Castillo (1); Fundo El Roble (1); La Invernada (10); Las Trancas (54); Los Angeles (5); Los Lleuques (10); Nonquén (1); Piedras Comadres (10); Puente Marchant (8); Puente Torrealba (1); Ralco (1); Recinto (4); Río Ñuble (1); Shangri-la (7); Yungai (1); region record only (1). IX Región de La Araucanía (59): Angol (7); Cunco (1); Laguna Galletue (3); Liucura (1); Lonquimay (15); Malalcahuello (1); Marimenuco (7); Molco (1); Nahuelbuta (2); Parque Nacional Nahuelbuta, Pehuenco (1); Parque Nacional Villarrica, Puesco (5); Temuco (3); Traiguen (1); Villa Portales (10); Volcán Lonquimay (1). XIV Región de Los Ríos (1): Valdivia (1). Country record only (3). Temporal data: January (60), February (26), March (8), April (5), September (5), October (24), November (89), December (123). Remarks: Identification of this species is complicated because of the variable head armature of males. The second clypeal horn can be either prominent or absent, and when present, this horn is either bifurcate or simple. In individuals where the second clypeal horn is absent, there is in its place a pair of distinctly separated tubercles behind the first clypeal horn (Figs. 17–18). This variation is not as extreme as the male dimorphism recognized by Howden (1979) in some species of the Australian genus Blackburnium Boucomont, but it is enough to cause confusion and probably lead to the failure to recognize B. geotrupoides and B. binasutum as synonyms until now. We examined over 200 male specimens of this species and found the male armature of B. geotrupoides, B. binasutum, and B. distinguendum to fall within our concept of this somewhat variable species. Therefore the latter two names are here synonymized under B. geotrupoides.Published as part of Mondaca, José & Smith, Andrew B. T., 2008, A revision of the southern South American genus Bolborhinum Boucomont (Coleoptera: Geotrupidae: Bolboceratinae), pp. 1-48 in Zootaxa 1794 on pages 10-11, DOI: 10.5281/zenodo.18260

    Sous demande de l'Assemblée, M. de La Porte remet sur le bureau le mémoire et le billet du roi, mais il s'oppose à lecture publique de ce dernier, lors de la séance du 21 juin 1791

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    Aubergeon de Murinais Guy Joseph d', Regnaud de Saint-Jean d'Angély Michel Louis Etienne, Briois de Beaumetz Bon-Albert, Defermon des Chapelières Jacques, Moreau Etienne-Vincent, Lameth Charles Malo, comte de, Beauharnais Alexandre François, vicomte de, Laporte Arnaud. Sous demande de l'Assemblée, M. de La Porte remet sur le bureau le mémoire et le billet du roi, mais il s'oppose à lecture publique de ce dernier, lors de la séance du 21 juin 1791. In: Archives Parlementaires de 1787 à 1860 - Première série (1787-1799) Tome XXVII - Du 6 juin au 5 juillet 1791. Paris : Librairie Administrative P. Dupont, 1887. pp. 377-378

    Dendropaemon (Sulcopaemon) quadratus Laporte 1832

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    39. Dendropaemon (Sulcopaemon) quadratus (Laporte, 1832) (Figs. 39, 104–105, 132, 153, 160) Enicotarsus Quadratus Laporte 1832, Ann. Soc. Ent. Fr. 1: 403 (original description) Enicotarsus quadratus: Castelnau 1840, Hist. Nat. Ins. 2: 83 (diagnosis, distribution) Dendropemon quadratus: Harold 1869, Cat. Col. IV: 1020 (mentioned as synonym) Dendropemon smaragdinus Waterhouse 1891, Ann. Mag. Nat. Hist. 6 8: 56 (original description) new synonymy Dendropemon quadratus: Felsche 1909, Deut. Ent. Zeit. 1909: 757 (comment) Dendropemon quadratus: Gillet 1911, Col. Cat. 38: 88 (mentioned as synonym) Dendropemon smaragdinus: Gillet 1911, Col. Cat. 38: 88 (catalogue) Dendropaemon planus Olsoufieff 1924, Insecta 13: 123 (original description) new synonymy Dendropaemon quadratus: Olsoufieff 1924, Insecta 13: 124 (monograph) Dendropaemon quadratus: Pessôa & Lane 1936, Rev. Biol. Hygiene 7: 89 (identification key, comment) Dendropaemon planus: Blut 1939, Arch. Naturg. (N.F.) 8: 293 (mentioned as synonym) Dendropaemon smaragdinus: Blut 1939, Arch. Naturg. (N.F.) 8: 293 (monograph) Dendropaemon quadratus: Blut 1939, Arch. Naturg. (N.F.) 8: 294 (monograph) Dendropaemon smaragdinus Chevrolati Blut 1939, Arch. Naturg. (N.F.) 8: 295 (original description) new synonymy Dendropemon planum: Blackwelder 1944, U.S. Nat. Mus. Bull. 185: 211 (checklist) Dendropemon quadratum: Blackwelder 1944, U.S. Nat. Mus. Bull. 185: 211 (mentioned as synonym) Dendropemon smaragdinum: Blackwelder 1944, U.S. Nat. Mus. Bull. 185: 211 (checklist) Dendropemon smaragdinum chevrolati: Blackwelder 1944, U.S. Nat. Mus. Bull. 185: 211 (checklist) Dendropaemon quadratum: Edmonds 1972, Univ. Kansas Sc. Bull. 49: 851 (comment taxonomy) Dendropaemon (D.) smaragdinus: Edmonds 1972, Univ. Kansas Sc. Bull. 49: 851 (comment taxonomy) Dendropaemon planus: Arnaud 1982, Rev. Fr. Ent. (N.S.) 4: 117 (lectotype designation) Dendropaemon smaragdinum chevrolati: Vaz-de-Mello 2000, Hac. Proy. CYTED: 192 (faunistic) Dendropaemon smaragdinum smaragdinum: Vaz-de-Mello 2000, Hac. Proy. CYTED: 192 (faunistic) Dendropaemon planus: Vitolo 2000, Rev. Acad. Colomb. Cienc. 24: 599 (identification key) Dendropaemon (D.) quadratus: Arnaud 2002, Col. Monde 28: 15 (mention) Dendropaemon (D.) smaragdinus: Arnaud 2002, Col. Monde 28: 15 (mention) Dendropaemon (D.) smaragdinus: Vaz-de-Mello & Génier 2009, Col. Bull. 63: 365 (biology) Type locality. Jatahy, Etat de Goyaz. Diagnosis. The small size (less than 10.0 mm), with distinct metallic markings combined with the emarginate anterior edge of clypeus on external side of each clypeal tooth will place this species in the quadratus complex. It differs from all other species in the quadratus complex by the two-(instead of three-) segmented meso-and metatarsi. Description. Male neotype (Fig. 39). Body. Body small, length 8.5 mm, maximum width 4.5 mm; body subrectangular in dorsal view; dorsum largely flat. Color. Dorsal surface dark brown to black, glossy, with green metallic sheen; head black along anterior edge of clypeus, metallic green on remaining surface; pronotum with green metallic sheen except on small areas along anteromedian carina and adjacent to lateral fossae; elytra with uniform green metallic sheen; ventrum black; pygidium with green metallic sheen; legs dark reddish brown to black. Head. Clypeus broadly arcuate, anterior portion slightly upturned; clypeal teeth triangular; clypeal median emargination broadly u-shaped, clypeal edge emarginate on external side of each clypeal tooth, clypeal teeth ventral surface lacking carina, clypeal margin ill-defined, bordered posteriorly by a more or less regular row of punctures, clypeal surface with transverse blunt rugulae; clypeogenal suture well-defined, bluntly carinate internally; genal surface with fine blunt transverse rugulae laterally, minutely punctate internally, transversely tumescent; clypeofrontal carina low, more than 6 times wider than high, slightly arcuate in dorsal view, simply carinate, clypeofrontal carina apical edge slightly trilobate in frontal view; eyes small in dorsal view, interocular ratio 4.9. Pronotum. Pronotum transverse in dorsal view, pronotal width/length ratio 1.4; disc of pronotum minutely punctate throughout, with a fine longitudinal sulcus on posterior two-third; pronotal anterior margin wider and flat lateral to eyes; anterior portion with a medially sinuous sharp carina transversely tuberculate medially; anterior angles surface with fine tubercles laterally and few short longitudinal rugulae internally, slightly but distinctly sulcate along posterior edge of anterior margin; lateral fossae rounded and bordered anteriorly by a sharp carina; lateral portions unmodified; pronotal basal fossae very small and more or less rounded; posterior margin well-defined, lacking crenulation and setae. Elytra. Elytra approximately as long as wide in dorsal view, elytral combined width/length ratio 1.0; elytral base lacking distinct margin, simply convex; elytral striae 1–4 very wide basally and tapering toward apex, distinctly more impressed basally, elytral striae 5 similar to 4 on disc, strial punctures ill-defined throughout, stria 1 well-defined apically, going straight to elytral apical margin; interstriae slightly convex, minutely punctate throughout, surface glossy. Thoracic sterna. Proepisternal carina reduced, present along coxal insertion only; metasternal median lobe flat, ventral ridge absent. Legs. Profemur posterior surface slightly but distinctly convex and glabrous internally, posterointernal margin rather thick, evenly developed, internal edge rather narrow, with a contiguous row of setae along anterointernal edge and few scattered long setae on anterior half, remaining surface smooth. Protibia with four teeth on lateral edge; internal basal angle lobate; anterior surface lacking aligned row of setae internally, with few aligned and isolated setiferous punctures only, surface coarsely microsculptured between punctures; posterior surface with irregular rugose punctures externally to median carina, surface finely and irregularly microsculptured between punctures, with a single interrupted setal row along lateral teeth. Mesofemur angularly produced on anterointernal edge apically. Mesotibia rather short, gradually widening toward apex in anterior view; anteroapical edge slightly sinuate in anterior view, anteroapical row of setae complete; apicoanterior edge circularly indented internally; external edge more or less rounded, with several large elongate setiferous punctures. Mesotarsus similar in shape to metatarsus, 2 -segmented, first segment moderately elongate, approximately two times as long as wide at apex. Metafemur internal edge nearly straight and lateral edge arcuate, lacking distinct depressed area anterointernally before apex, apicoposterior edge unmodified, anterior surface with a well-defined sulcus on median half. Metatibia robust, short, slightly widening toward apex in anterior view, anterior surface with distinct row of setae, surface glossy externally with distinct transverse microsculpture internally, metatibial posterior surface concave between internal and lateral edge, with transverse microsculpture. Metatarsus 2 -segmented (Fig. 132), first segment moderately elongate, approximately two times as long as wide at apex, with anterointernal carina well defined and almost reaching apical edge. Abdominal sternites. Sternites 3–6 longitudinally slightly convex; sternites 4–6 with 1–3 unaligned rows of setae laterally, glabrous medially; sternite 7 slightly longitudinally concave medially, shorter than segment 6 along midline; pygidium minutely punctate on disc. Male genitalia (Figs. 104–105). Parameres produced into a lobe laterally; surface smooth, glossy apically. Measurements (9 males, 8 females). Length: male 7.5–9.5 (8.3 ± 0.6), female 6.5 –10.0 (8.6 ± 1.2) mm. Primary type data. Enicotarsus quadratus Laporte (Fig. 153). Male neotype (MNHN) present designation: [Trinidade/ (Goyaz)/ Ch.Pujol]; [WORLD / SCARAB./ DATABASE/ WSD00016803]; [NEOTYPE / Enicotarsus / quadratus / Laporte, 1832 / dés. Génier & Arnaud, 2014] red card; [Dendropaemon / quadratus / Laporte/ dét. F. Génier, 2009] handwritten. Dendrop[a]emon smaragdinus Waterhouse . Holotype male (BMNH): [Type] disc with red border; [67 45]; [TYPE] red card; [Dendropemon / smaragdinus,/ (Type) Waterh.] handwritten; [WORLD / SCARAB./ DATABASE/ WSD00016804]; [HOLOTYPE / Dendrop[a]emon/ smaragdinus / Waterhouse, 1891] red card. Dendropaemon planus Olsoufieff . Lectotype female (MNHN): [Jatahy/ (GOYAZ)] green card; [MUSÉUM PARIS / 1936 / COLL. A. BOUCOMONT] green card; [Typus] red card; [Dendrop. planus n.sp. / det. G. OLSOUFIEFF] partly handwritten; [WORLD / SCARAB./ DATABASE/ WSD00016805]; [Dendropaemon / planus Ols. / LECTOTYPE / P. ARNAUD DET 1982] partly handwritten, red border. Dendropaemon smaragdinus chevrolati Blut . Holotype female (MTD): [Enicotarsus / quadratus / de Lap Bresil / type] handwritten; [Brasilien] green card; [Coll. C. Felsche/ Kauf 20, 1918] green card; [♀]; [Dendropaemon / s. Chevrolati ♀/ Brasilien.— Type.] handwritten, black border; [viridis / Perty/ Paraguay] purple border; [WORLD / SCARAB./ DATABASE/ WSD0016807]; [HOLOTYPE / Dendropaemon / smaragdinus chevrolati / Blut, 1939] red card; [Dendropaemon ♀/ quadratus / (Laporte, 1832)/ dét. Génier & Arnaud, 2009]. Material examined. [NO DATA]: -., coll. [anonymous]— 1 male (holotype of D. smaragdinus Waterhouse) (BMNH); ARGENTINA: MISIONES, Dos de Mayo, elev. 500 m (27 ° 1 ' 39 ''S, 54 ° 40 ' 3 ''W), vii. 1977, coll. R. Foerster— 1 female (CPFA); Loreto, (27 ° 18 ' 22 ''S, 55 ° 32 ' 10 ''W), i. 1956, coll. F.H. Walz— 1 male (BDGC); same locality, [no date], coll. A. Ogloblin— 1 female (CEMT); same locality, i. 1955, coll. F.H. Walz— 1 female (CMNC); same locality, xi. 1958, coll. Martínez— 1 female, 2 males (CMNC); same locality, i. 1954, coll. F.H. Walz— 1 male (CMNC); BRAZIL: [unspecified locality], [no date], coll. [anonymous]— 1 male (CPFA); GOIÁS, Jataí, (17 ° 53 'S, 51 ° 43 'W), [no date], coll. [anonymous]— 1 female (lectotype of D. planus Olsoufieff) (MNHN); Rio Verde, (17 ° 47 ' 50 ''S, 50 ° 54 '0''W), [no date], coll. [anonymous]— 1 female, 1 male (MTD); Trindade, (16 ° 38 ' 52 ''S, 49 ° 29 ' 53 ''W), [no date], coll. Ch. Pujol— 1 male (neotype) (MNHN); same locality, [no date], coll. [anonymous]— 1 female (ZMHB); same locality, [no date], coll. Ch. Pujol— 1 female (paralectotype of D. planus Olsoufieff) (MNHN); [unspecified locality], [no date], coll. [anonymous]— 1 female (holotype of D. smaragdinus chevrolati Blut) (MTD); MATO GROSSO DO SUL, UNESP Farm [=Fazenda Experimental da Universidade Estadual Paulista, câmpus de Ilha Solteira], Selvíria, (20 ° 20 ' 2.1 ''S, 51 ° 24 ' 26.3 ''W), 18.i. 2008, coll. F. Oikawa— 1 female (MEFEIS); [NO DATA]: -., coll. [anonymous]— 1 female, 1 male (MNHN, MNRJ); PARAGUAY: ITAPÚA, Puerto Cantera, (27 ° 13 ' 44 ''S, 55 ° 36 ' 9 ''W), xii. 1956, coll. F.H. Schade— 1 female (CMNC); same locality, xi. 1956, coll. Walz— 1 male (CMNC). Natural history. A specimen was collected in a pitfall trap baited with Sus scrofa dung. Remarks. Females differ in having the anterior pronotal carina straighter and set very close to the anterior margin, the carina is only finely tuberculate medially. The abdominal sternite 7 is also nearly flat medially. Variation, as usual, occurs in the extent of the metallic marking on head and pronotum. The greenish tinge is rather uniform in the sample studied. The extent and density of the fine tubercles and rugulae on the pronotal anterior angles vary, in some specimens the sculpturing is finer and denser and set on a slightly larger area. Nomenclature and taxonomy. The type of D. quadratus is presume destroyed (see comment under D. ater on Laporte’s collection). Historically, the status of this species has remain unsettled. In the first review of the group, Castelnau (1840) simply copied the description he made in 1832 omitting the Latin diagnosis. Harold (1869), in his catalogue, considered D. quadratus as a synonym of D. viridis. Felsche (1909) briefly commented Harold’s synonymy and suggest that D. quadratus cannot be synonymous with D. viridis as the meso- and metatarsi are three segmented and are only two segments in D. viridis. Felsche stated that the third segment is present but barely visible based on a specimen studied from Chevrolat’s collection and labeled as “ Enicotarsus quadratus de Cast.- Type ”. Olsoufieff (1924) discuss again the situation and concluded that D. quadratus is indeed a valid species, but do not attempt to settle the issue. Pessôa and Lane (1936) essentially repeat Olsoufieff’s observations and questionably leave D. quadratus as a synonym of D. viridis. Finally, Blut (1939) concluded that the status of D. quadratus remain uncertain and is the first author to use the measurements given by Laporte in the original description to suggest that D. quadratus cannot be a synonym of D. viridis which is distinctly larger. Blut also rightly point out that the specimen from Chevrolat’s collection possess only two tarsal segments and not three as stated by previous authors. Furthermore, Blut suggest that D. quadratus body size would be similar to those of D. refulgen s. However, in the end, Blut conclude that the name D. quadratus is to be forgotten as he considered the name to be in litteris. The purpose of this work is to address all taxonomic problems pertaining to the genus and therefore we designate a neotype for Laporte’s species D. quadratus of which the original description meet requirements for being valid. In order to remain as conservative as possible we designate a male specimen from the Museum national d’Histoire naturelle that comply with the original description and has precise locality data. This specimen matches approximately the size given by Laporte and the very rudimentary description. It also matches the historical specimen from Chevrolat’s collection. This specimen differs from the original description by the “[pronotum] très-fortement ponctué avec des espaces lisses”, which could correspond to certain Coprophanaeoides species. We however, interpret the heavy punctation as being the sculpturing of the lateral pronotal declivities. Following the designation of a neotype for D. quadratus the following nomenclatural acts can now be implemented (the second epithet being valid): 1) D. smaragdinus Waterhouse, 1891 = D. quadratus Laporte, 1832, new synonymy. The holotype of D. smaragdinus is a very small individual of D. quadratus, aside being slightly teneral and having the median lobe of the metasternum slightly more convex no differences can be found. 2) D. planus Olsoufieff, 1924 = D. quadratus Laporte, 1832, new synonymy. The lectotype of D. planus has been studied and except for being a female and having the pronotal green metallic marking less extensive no other differences could be found. 3) D. smaragdinus chevrolati Blut, 1939 = D. quadratus Laporte, 1832, new synonymy. The holotype female of D. smaragdinus chevrolati has been compared to the holotype of D. quadratus and only differs in the extent of the pronotal metallic marking which are absent on most of the disc. The pronotal and cephalic microsculpture are slightly more pronounced. The fine tubercles and rugulae of the pronotal lateral declivities are denser and cover slightly more surface. The pronotal punctures are fine instead of minute laterally posterior to lateral fossae. Some specimens in the material studied show intermediate forms and this variation is here considered intraspecific.Published as part of François Génier & Patrick Arnaud, 2016, Dendropaemon Perty, 1830: taxonomy, systematics and phylogeny of the morphologically most derived phanaeine genus (Coleoptera: Scarabaeidae, Scarabaeinae, Phanaeini) in Zootaxa 4099 (1) on pages 76-79, DOI: 10.11646/zootaxa.4099.1.1, http://zenodo.org/record/26827

    Le caractère substantiel des formalités de publicité préalable à l'adjudication

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    International audience(Civ. 2e, 13 janv. 2022, no 20-18.155, B, D. actu. 28 janv. 2022, obs. C. Bléry ; Rev. prat. rec. 2022. 18, chron. Aude-Alexandre Le Roux et O. Salati ; JCP G 2022, 352, note C. Laporte ; Gaz. Pal. 15 mars 2022, obs. C. Brenner

    Using ISO/IEC 29110 to harness process improvement in very small entities

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    The recently published ISO/IEC 29110 standard Lifecycle profiles for Very Small Entities has at its core a Management and Engineering Guide [1] which are targeted at very small entities (enterprises, organizations, departments or projects) having up to 25 people [2], to assist them unlock the potential benefits of using standards which are specifically designed to address their needs. This paper will outline this new standard and the implementation of a series of pilot project initiative harnessing a set of detailed guidelines known as “Deployment Packages” to assist very small entities in understanding the potential usage of this new software process standard

    Heteroscelis servillii Laporte 1833

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    Heteroscelis servillii Laporte, 1833 (Figs. 1–14) Heteroscelis servillii Laporte, 1833: 86; Stål 1870: 31; Kirkaldy 1909: 30 (syn.). Phyllocheirus servillei: Spinola 1837: 306 Phyllochirus servillei: Amyot & Serville 1843: 80 (emen); Stål 1862: 93; Stål 1870: 31 (syn.); Kirkaldy 1909: 30. Agerrus remipes Stål, 1859: 434; Stål 1862: 93 (syn.); Schouteden 1907: 19; Kirkaldy 1909: 30. Bodetria brenthoides Walker, 1867: 119; Stål 1870: 31 (syn.); Distant 1900: 63; Schouteden 1907: 19; Kirkaldy 1909: 30. Bodetria chrysoclora Walker, 1868: 528; Stål 1870: 31 (syn.); Distant 1900: 63; Schouteden 1907: 19; Kirkaldy 1909: 30. Bodetria indecora Walker, 1868: 528; Stål 1870: 31 (syn.); Distant 1900: 63; Schouteden 1907: 19; Kirkaldy 1909: 30. Heteroscelis brenthoides: Schouteden 1907: 19; Kirkaldy 1909: 30 (syn.). Heteroscelis servillei: Distant 1900: 63; Schouteden 1907: 19; Pirán 1961: 87; Thomas 1992: 46; Grazia & Schwertner 2011: 713; Grazia et al. 2015: 686; Silva et al. 2018: 404; Lupoli 2019: 80; Cazorla 2021: 10; Castro-Huertas et al. 2022: 9; Roell et al. 2023: 28, 29, 47. Phyllocheirus brenthoides: Kirkaldy 1909: 30. Phyllocheirus servillii: Kirkaldy 1909: 30; Pirán, 1961: 87 (syn.). Heteroscelis zischkai Kormilev, 1951: 40; Pirán 1961: 88; Thomas 1992: 46 (syn.). Heteroscelis dureti Pirán, 1954: 19; Thomas 1992: 46 (syn.). Heteroscelis bergi Pirán, 1954: 20; Thomas 1992: 46 (syn.). Heteroscelis carcavalloi Pirán, 1961: 96; Thomas 1992: 46 (syn.). Heteroscelis amazonica Pirán, 1961: 98; Thomas 1992: 46 (syn.). Types examined SURINAM. Agerrus remipes Stål, 1859; female syntype; labels: “Surinam.”, “Stål”, “ Agerrus remipes Stål. Typ. ”, “111”, “TYPUS”, “NHRS—GULI 000096153”; (NHRS). Photo examined (Fig. 3). BRAZIL. Heteroscelis amazonica Pirán, 1961; female holotype; labels: “Ig do Passarinho Manaus. Am. 24.II.55. Costa Leite e N. Cerqueira—cols. 07. 9/59”, “ Heteroscelis amazonica Pirán 1960 ”, “ HOLOTYPUS ”; (MNRJ) (Fig. 4). BRAZIL. Heteroscelis carcavalloi Pirán, 1961; female alotype; labels: “Sítio Vieiralves Manaus—Am. 21.Xll.65. Elias e Rappa—cols. 02. 9/59”, “ Heteroscelis carcavalloi Pirán 1960 ”, “ ALLOTYPUS ”; (MNRJ). BRAZIL. Bodetria brenthoides Walker, 1867; female holotype; labels: “Amazon St. Paul / 60 32” “Type”, “ Holotype ”, “1. Bodetria brenthoides ”; “NHMUK 010592390”; (NHMUK). Photo examined (Fig. 5). BRAZIL. Bodetria chrysoclora Walker, 1868; male holotype; labels: “Braz / 62 57” “Type”, “ Holotype ”, “ Bodetria chrysoclora ”; “NHMUK 010592391”; (NHMUK). Photo examined (Fig. 6). BRAZIL. Bodetria indecora Walker, 1868; male holotype; labels: “Braz / 62 57” “Type”, “ Holotype ”, “ Bodetria indecora ”; “NHMUK 010592393”; (NHMUK). Photo examined (Fig. 7). BOLIVIA. Heteroscelis carcavalloi Pirán, 1961; male holotype; labels: “ HOLOTYPUS ”, “Yungas del Palmar. IV–954. A. Martinez.”, “ Heteroscelis carcavalloi Pirán, 1960.”; (MACN). Photo examined (Fig. 8). BOLIVIA. Heteroscelis zischkai Kormilev, 1951; female alotype; labels: “ BOLIVIA, Region Chaparé, Dep. Cochabamba (400 mtr), 30. VIII. 1951. Dirings.”. Heteroscelis zischkai Kormilev. Museo Argentino de Ciencias Naturales. ”. “ ALLOTYPE ”; (MACN) (Fig. 9). PARAGUAY. Heteroscelis dureti Pirán, 1954; female holotype; labels: “ Heteroscelis dureti Det. Pirán. 952”, “ HOLOTYPUS ”, “ Paraguay, Villarrica, Schade col. 15.IV.46”; (MACN). Photo examined (Fig. 10). ARGENTINA. Heteroscelis bergi Pirán, 1954; male holotype; labels: “Missiones—Argentina. Dep. Concep.— Sta. Maria. M.J. Viana”, “53.484”, “ HOLOTYPUS ”, “ Heteroscelis bergi Det. Pirán. 952”; (MACN). Photo examined (Fig. 11). Other material examined. VENEZUELA, Guayana: (Conuco, Rio Moroco), 3 males, 18/ 19.I.1972 G. & E. Scherer (NMPC); Zulia: (Kasmera, 9.94907, -72.75292), 1 female, 20.IX.1961 F. Fernandez Y. & C. J. Rosales (UFRG); Cx. Panamexicana El Vigia Coloneito: (Km 8.32611, -72.08742), 1 female, 6.I.1955. F. Fernandez Y. & C. J. Rosales (UFRG); Aragua: (El Limón, 10.30589, -67.63212), 1 male, 2.III.1951 F. Fernandes (UFRG); Aragua: (Maracay, 10.23535, -67.59113), 1 female, 2.V.1953 N. Angeles (UFRG). TRINIDAD AND TOBAGO: 2 males, XII.1953 G. u. Helga Frey. (NMPC). BRAZIL, 1 female, 11.IV.1962 Cerq. (UFRG); Pará: (Belém Mocambo, - 1.45583, -48.50444), 1 female, 6.II.1979 P Tadeu (UFRG); (Peixe-boi, -1.10168, -47.27259), 1 male, 12.IV.1977 P Waldir. (UFRG); (Serra Norte Manganês -6.01748, -50.30386), 1 female, 22.X.1984 T. Pimentel; Amazonas: (Manaus, CEPLAC, -2.8905, -59.9576), 1 female, 15.VII.1977 (UFRG); (Serra dos Porcos, 0.416667, -69.366667), 1 male, 25.II.1977 Franklim (INPA); (Manaus, CEPLAC, Am 010 Km 31, -2.8905, -59.9576), 1 male, 7.V.1988 Albuquerque (UFRG); (Manaus, -3.04361, -60.01282), 1 male, 5.V.1976 Paraluppi (UFRG); (Manaus, Br 174 Km 41, Agricultura Geológica -2.6770, -60.04668), 1 male, 15.VI.1981 C.A. Fonseca (INPA); (Itacoatiara, -3.1373, - 58.4418), 1 female, Dirings (MCNZ); Amapá: (Serra do Navio, 1.65803, -52.28195), 1 female, 13.VII.1961 AP. J & B. Bechyné (UFRG); Rio Grande do Norte: (Natal, -5.8101, -35.22674), 1 male, I.1952 M. Alvarenga (UFRG); Pernambuco: (Paudalho, Acerolândia, -7.850, -35.250. 160m), 1 male, II.2016 Grossi & Parizotto (CERPE); (Paudalho, Acerolândia, -7.850, -35.250. 160m), 1 female, XI.2017 Grossi & Parizotto (CERPE); Mato Grosso do Sul: (Bonito, Praia da Figueira -21.2972, -56.5005), 1 female, 01.III.2009 Grossi & Parizotto (DZUP); Rio Grande do Sul: (Cerro Azul -27.7403, -51.5199), 1 female, I.1931 Pe. Buck leg. (MCNZ); ECUADOR, Amazonia: 1 female, II.1998 lgt. Bezdek (NMPC); Napo: (Tiputini Biodiversity Station, 216m, -0.6379, -76.1495), 1 male, 06.II.1999 T.L. Erwin, et al. (USNM); (Tiputini Biodiversity Station, 216m, -0.6379, -76.1495), 1 female, 22.X.1998 T.L. Erwin, et al. (USNM); PERU, Pucalipa: (Rio Ucayali, -8.4084, -74.6058), 1 female, Dirings (MCNZ); BOLIVIA, Cochabamba (Chapare, 400m -16.80922, -65.71926), 1 female, 07.IX.1951 Dirings (MACN). Records retrieved from iNaturalist TRINIDAD AND TOBAGO, San Juan-Laventille: (10.605137, -61.42763), 14.II.2021. BRAZIL, Amazonas: (Manaus, -3.095784, -59.989879), 13.VIII.2020. BRAZIL, Espírito Santo: (S„o Gabriel de Palha, -18.906837, -40.478105), II.2014. BRAZIL, S„o Paulo: (Campinas, -22.819691, -47.064677), 16.II.2023. BRAZIL, Rio de Janeiro: (Cabo Frio, -22.886901, -42.026649), 8.I.2017. Diagnosis. Mandibular plates distinctly longer than clypeus, converging apically; head length anterior to the eyes more than 2/3 of the total length of the head (Fig. 2, D); urosternite III tubercle yellow or orange at apex, and surpassing the posterior margin of metasternum (Fig. 2, B). Redescription BODY: General color ranging from brown and dark red to iridescent hues of green, yellow, blue and purple, with or without yellow spots on the pronotum, scutellum and coria (more commonly observed in non-iridescent specimens). HEAD: Labium almost reaching the metasternum (Fig. 2, B); proportion of antennomeres: I<II<III ≅ IV ≅ V. THORAX: Coria longer than scutellum, not reaching the posterior margin of connexival segment VI, uniformly punctured; membrane dark and surpassing posteriorly the abdomen; metapleural evaporatoria surpassing posteriorly the anterior limit of coxae; peritreme reaching or surpassing half the pleural width; apex of peritreme rounded, posterior margin a little more arched than the anterior, both convex (Fig. 2, E); profemora with rudimentary ante-apical spine; foretibial length on average 3.5x larger than the width of the tibial dorsal expansion (Fig. 2, F). ABDOMEN: Posterior angles of urosternites III to VII ending in minute spine, more developed in VII (Fig. 2, B). Male: MEASUREMENTS (n=13): Head length 1.76 ± 0.28 (1.30–1.84); width 1.29 ± 0.07 (1.20–1.50); pronotum length 2.00 ± 0.26 (1.50–2.40); width 4.21 ± 0.26 (3.80–4.50); scutellum length 2.38 ± 0.16 (2.08–2.60); width 2.30 ± 0.13 (2.08–2.56); length of antennomeres: I (n=11) 0.39 ± 0.03 (0.30–0.40), II (n=11) 0.71 ± 0.09 (0.60 –0.90), III (n=11) 0.95 ± 0.10 (0.80–1.04), IV (n=11) 1.03 ± 0.14 (0.72–1.20); V (n=8) 1.09 ± 0.20 (0.72–1.40); length of labiomeres: I (n=8) 0.67 ± 0.08 (0.60–0.80), II (n=8) 0.87 ± 0.07 (0.80–1.00), III (n=8) 0.45 ± 0.07 (0.40–0.60), IV (n=8) 0.68 ± 0.09 (0.60–0.90); width of foretibal expansion (n=10) 0.59 ± 0.07 (0.48–0.72); width of abdomen (n=11) 3.21 ± 0.18 (2.96–3.50); total length 7.04 ± 0.65 (6.10–8.30). GENITALIA: Pygophore (Fig. 12): setae denser on posterolateral angles and on medial portion of dorsal rim; dorsal rim sinuous, medially concave; ventral rim slightly sinuous, medially concave; segment X with surface corrugated and with a membranous longitudinal line; head of parameres lateroventrally directed in posterior view. Phallus (Fig. 13): conjunctival dorsal lobe with one pair of long projections and one single forked central projection, ventral lobes with two projections each. Female: MEASUREMENTS (n=15): Head length (n=11) 2.06 ± 0.38 (1.40–2.60); width (n=11) 1.47 ± 0.20 (1.20– 1.90); pronotum length (n=13) 2.45 ± 0.26 (1.92–2.80); width (n=13) 4.83 ± 0.42 (3.92–5.60); scutellum length 2.88 ± 0.32 (2.24–3.40); width 2.56 ± 0.40 (1.84–3.00); length of antennomeres: I (n=12) 0.42 ± 0.06 (0.30–0.50), II (n=12) 0.79 ± 0.15 (0.56 –1.10), III (n=12) 1.04 ± 0.21 (0.60–1.30), IV (n=8) 1.22 ± 0.39 (0.80–2.00); V (n=5) 1.03 ± 0.18 (0.96–1.30); length of labiomeres: I (n=9) 0.77 ± 0.11 (0.60–0.90), II (n=9) 1.02 ± 0.16 (0.70–1.30), III (n=9) 0.50 ± 0.10 (0.40–0.70), IV (n=9) 0.85 ± 0.14 (0.60–1.00); width of foretibal expansion (n=11) 0.72 ± 0.09 (0.56–0.90); width of abdomen 3.70 ± 0.33 (3.04–4.30); total length (n=8) 8.28 ± 1.05 (6.88–9.50). GENITALIA (Fig. 14): Valvifers VIII posterior margin bisinuous, slightly projected over the basal angles of laterotergites IX; laterotergites VIII wider than long, with spiracles equidistant from the basal angle and the lateral margin; valvulae IX with 1+1 bean-shaped secondary thickenings; vesicular area half the length of the proximal part of the ductus receptaculi; capsula seminalis oval. Distribution: Colombia, Ecuador (new record), Trinidad and Tobago, Venezuela, Guyana, Surinam, French Guyana, Brazil, Peru, Bolivia, Paraguay and Argentina (Fig. 1).Published as part of Sampaio, Vinicius Gomes, Roell, Talita & Campos, Luiz Alexandre, 2023, Revision of Heteroscelis Latreille, 1829 (Heteroptera: Pentatomidae: Asopinae) with the description of a new species from Colombia, pp. 401-438 in Zootaxa 5278 (3) on pages 405-416, DOI: 10.11646/zootaxa.5278.3.1, http://zenodo.org/record/790637

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