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Ankylocythere barbouri Villalobos Figueroa & Hobbs 1974, new species
No full textAnkylocythere barbouri Villalobos Figueroa & Hobbs, 1974 1974 Ankylocythere barbouri, new species.—Villalobos Figueroa & Hobbs: 2, 7–9, 16, 17, Fig. 1, 3a–e (D, H, R, T). 2014a Ankylocythere barbouri Villalobos & Hobbs, 1974.—Mestre, Monrós & Mesquita-Joanes: 931, 946 (C).Published as part of Williams, Bronwyn W. & Weaver, Patricia G., 2018, A historical review of the taxonomy and classification of Entocytheridae (Crustacea: Ostracoda: Podocopida), pp. 1-129 in Zootaxa 4448 (1) on page 27, DOI: 10.11646/zootaxa.4448.1.1, http://zenodo.org/record/144449
Huellas de ceniza. Lic.Enrique Villalobos
No full textLa reseña realizada por el autor Enrique Villalobos, de la novela Huellas de ceniza muestra una perspectiva critica de la sociedad costarricense a partir del proceso de globalización.The review by author Enrique Villalobos, of the novel Huellas de ceniza shows a critical perspective of the Costa Rican society from the globalization process.Universidad NacionalCentro de Estudios Generale
Tzotzilthelphusa villarosalensis Villalobos & Álvarez, 2013, n. sp.
No full textTzotzilthelphusa villarosalensis n. sp. (Figs. 6, 7) Type material. Holotype: male, cb 31.2 mm, cl 20.9 mm; ditch along highway 101 near Villa de las Rosas (16 º 21 ' 0.5 " N, 92 º 23 ' 59.07 " W; 1152 m), Municipio de Villa de las Rosas, Chiapas, Mexico; 3 Dec. 2007; coll. F. Álvarez, J.L. Villalobos, A. Botello, C. Hernández; CNCR 26192. Allotype: female, cb 24.8 mm, cl 16.8 mm,; same locality as holotype, 30 Apr 2009; coll. F. Álvarez, J.L. Villalobos, G. Armendáriz, E. Torres; CNCR 25385. Paratypes: 4 males, cb 32.5 -21.0 mm, cl 21.0- 18.7 mm; 3 females, cb 21.9 - 19.7 mm, cl 15.0- 13.4 mm; same locality, date and collectors as allotype; CNCR 27339. Description. In dorsal view carapace moderately oval, widest in middle (cb/cl = 1.63); dorsal surface convex, punctated, regions faintly indicated, cephalic and branchial somewhat elevated; pair of gastric pits barely distinct, oblique, narrowly elongated; cervical grooves wide, shallow, arched, not reaching anterolateral margin (Fig. 6 a). Postfrontal lobes and median groove discernible. Frontal region curved downwards to frontal border; superior frontal border absent, inferior one smooth, projected, divided into 2 low convex lobes, separated by rounded medial depression (Fig 6 d). Superior orbital margin continuous with frontal border; inferior orbital margin marked by row of rounded granules; exorbital angle flat. Anterolateral margin smooth along anterior third, granulated on branchial region, straight through posterior half. Posterior margin concave. Eyes with cornea normally faceted, pigmented, filling orbits. Epistome narrow; epistomial tooth low, triangular, deflexed, borders smooth. Suborbital and subhepatic regions of carapace smooth, punctated; pterygostomial region pubescent around mouthparts. Third maxilliped with ischium rectangular, merus with external margin widely rounded, ratio exopod/ischium 0.9 (Fig. 6 c). Aperture of efferent branchial channel subquadrate. Chelipeds different in shape and size. Major cheliped with merus subtriangular in cross section, short, thick, shorter than carapace length, dorsal surface smooth; internal dorsal margin with row of acute granules; external margin delimited by rounded granules on proximal two-thirds. Carpus smooth with small, low tubercles on internal margin. Palm compressed, high (length/high = 1.3); fingers longer than palm, stout, smooth, not gaping, with low, triangular teeth on cutting edges (Fig. 6 b). Thoracic sternites of third maxilliped and first pereiopod completely fused, except for short fissures at lateral edges of sternum; sternal sulci s 4 /s 5, s 5 /s 6 and s 6 /s 7 distinct, just failing to reach midline of thoracic sternum; sternal sulcus s 7 /s 8 reaching midline. All abdominal segments free. Telson subtriangular, lateral margins straight, tip rounded. First male gonopod as described under genus diagnosis. Etymology. The specific name is derived from "Villa de las Rosas" the name of the closest town to the type locality in Chiapas, Mexico. Remarks. Tzotzilthelphusa villarosalensis n. sp. has a gonopod with the distal third bent laterally, but without torsion. The apical cavity of the gonopod is oblique to the principal axis of the gonopod due to the elevation of the mesial surface. The gonopod morphology of the new species is similar to that of Typhlopseudothelphusa hyba, raising an interesting problem because that species was placed in Typhlopseudothelphusa due to its external morphology, which is very similar to that of T. mocinoi. However, considering the gonopod morphology T. hyba could be placed in Tzotzilthelphusa n. gen., a problem that will have to be addressed in the future. Geographically, T. hyba and T. villarosalensis n. sp. are only 40 km apart to the south of San Cristóbal de las Casas.Published as part of Villalobos, José Luis & Álvarez, Fernando, 2013, Two new genera and three new species of freshwater crabs (Crustacea: Pseudothelphusidae: Potamocarcinini) from Chiapas, Mexico, pp. 457-470 in Zootaxa 3599 (5) on page 467, DOI: 10.11646/zootaxa.3599.5.3, http://zenodo.org/record/21881
ÁLVAREZ CONDE, Enrique;Figueruelo BURRIEZA, Ángela ; NUÑO GOMEZ , Laura (Dir.) ; CANCIO ÁLVAREZ, Ma. Dolores (Coord.), Estudios Interdisciplinares sobre Igualdad, Madrid, Iustel, 2011. 798 pp.
No full textVillalobos, WV (reprint author), Univ Talca, Talca, Chile
Alvarezius zongolicae Moreno-Juárez & Villalobos & Álvarez 2022, n. comb.
No full textAlvarezius zongolicae (Alvarez, Villalobos & Moreno, 2012) n. comb. Pseudothelphusa zongolicae Alvarez, Villalobos & Moreno, 2012, figs. 2, 3. — Villalobos et al., 2019: 157 (Table 1). — Álvarez et al., 2020: 10 (Supplementary material, Table 1). Type material. Holotype: CNCR 26641; 1 ♂, CL 18.1 mm, CW 30.0 mm; spring at Choapa (18°43’32”N, 96°57’07”W, 1053 m asl), Municipality of Zongolica, Veracruz, Mexico; colls. E. Moreno, O. Trejo; 19 April 2011. Paratype: CNCR 26642; 1 ♂, CL 17.7 mm, CW 28.7 mm; same locality as holotype; 12 June 2011. Other material examined. CNCR26643, 4♂, 1♀,CL 15.8–22.8mm,CW 25.5–37.5 mm, Macuilca (18°38’49”N, 96°56’58”W, 528 m asl), Municipality of Zongolica, Veracruz, Mexico, coll. E. Moreno & O. Trejo, 11 June 2011; CNCR 29494, 5 ♂, CL 8.9–19.9 mm CW 13.6–32.7 mm, spring of water, Tepeplampa (18°33’41”N, 96°56’51”W, 891 m asl), Municipality of Zongolica, Veracruz, Mexico, coll. E. Moreno & S. Rodríguez, 12 December 2012; CNCR 29495, 6 ♂, 1 ♀, CL 11.2–23.3 mm, CW 12.5–40.6 mm, stream near Tepetlampa (18°33’10”N, 96°56’30”W, 110 m asl), Municipality of Zongolica, Veracruz, Mexico; colls. E. Moreno & S. Rodríguez; 11 December 2012; CNCR 35458, 2 ♂, 2 ♀, CL 18.1–21 mm, CW 22.4–36.2 mm, same locality as holotype, coll. E. Moreno, A. Soto & R. Rodríguez, 10 October 2019; CNCR 36478, 4 ♂, 4 ♀, CL 7.8–9.8 mm, CW 10.6–15.6 mm, spring inside cave, Cintalapa (18°38’12”N, 96°54’0.4”W, 1050 m asl), Municipality of Zongolica, Veracruz, Mexico; colls. E. Moreno & L. Gómez; 3 April 2022. CNCR 36479; 2 ♂, spring in Acahualco, Cuahutilica (18°36’19.6”N, 96°56’26”W, 1169 m asl), Municipality of Zongolica, Veracruz, Mexico; colls. E. Moreno & L. Gómez, 3 April 2022; CNCR 36480, 1 ♂, CL 23.2 mm, CW 39.9 mm, waterfall on the road, Coapa-Pinopa (18°37’16.2”N, 96°56’21.5”W, 1063 m asl), Municipality of Zongolica, Veracruz, Mexico, coll. E. Moreno & L. Gómez, 3 April 2022; CNCR 36481, 3 ♂, CL 20.7–24.2 mm, CW 34.9–41.4 mm, Acahualco Cave, Cuahutilica (18°36’16”N, 96°56’09”W, 1,111 m asl), Municipality of Zongolica, Veracruz, Mexico, coll. E. Moreno & L. Gómez, 3 April 2022. Description. Carapace transversely oval, dorsal surface slightly convex, smooth, finely punctate, regions faintly marked, except for moderately swollen mesobranchial region (Fig. 5a). Superior frontal border present, incomplete towards middle portion, formed by incomplete row of granules, in frontal view inclined towards median groove (Fig. 5b). Inferior frontal border complete, thick, slightly rounded in dorsal view, bilobed in frontal view, extending laterally to form superior margin of orbits. Median groove deep, narrow, arising above inferior frontal border, extending posteriorly beyond postfrontal lobes. Postfrontal lobes low, well defined. Cervical grooves wide, shallow, straight towards margin of carapace, ill- defined, slightly arching towards central portion of carapace, not reaching anterolateral margin. Walking legs typical of genus. Chelipeds asymmetrical. Major chela right, palm somewhat swollen, movable finger half-length of chela, fingers not gaping; both fingers with irregular arrangement of teeth of varying sizes, both fingers curved inwards (Fig. 5c). Third maxilliped with merus showing complete rounded external margin, ischium rectangular, ratio ischium/exopod 0.85 (Fig. 5d). G1 moderately slender, distal half of principal axis showing mesial torsion of caudo-marginal projection. In mesial view (Fig. 5e), marginal suture evident, arching distally, disappearing under caudo-marginal projection; lobe of caudo-marginal projection (Cmp) axe-shaped, large, about half length of gonopod, cephalic border smooth; caudo-marginal distal crest straight, spinulated at cephalic curvature. In cephalic view (Fig. 5f), mesial process (Mp) oriented laterally, slightly inclined proximally, tapering laterally, ending in two acute tips, superior margin closing apex cavity, extending towards internal portion of apex cavity forming central crest; caudo-marginal projection with single large, axe-shaped lobe, extending proximally slightly beyond half-length of gonopod, cephalic margin armed with small, acute teeth; crest of caudo-marginal projection becoming less thick cephalically; conical prominence (Cp) on caudal angle of lateral crest bearing apical setae. In lateral view (Fig. 5g), mesial process with two tips oriented laterally; caudo-marginal projection (Cmp) appearing as large tongue-shaped lobe; conical prominence (Cp) emerging from lateral crest. In caudal view (Fig. 5h), caudal surface showing torsion of gonopod; mesial process (Mp) subtriangular; conical prominence (Cp) of lateral crest evident; lobe of caudo-marginal projection (Cmp) elongate, slender, extending parallel to main axis of gonopod; marginal suture straight, visible along proximal two thirds of gonopod. In apical view (Fig. 5i), apex cavity approximately rectangular, field of spines (Sf) on lateral side of cavity, extending to conical prominence (Cp); central crest of apex cavity sharp, slightly undulated; distal crest of lateral surface straight; mesial process (Mp) subtriangular, distal margin rounded; lobe of caudo-marginal projection (Cmp) as undulated sheet, cephalic margin with minute spines; opening of sperm channel caudal. Distribution. Mexico, Veracruz, in the surroundings of the Municipality of Zongolica. Type locality. Spring at Choapa (18°43’32”N, 96°57’07”W; 1,053 m asl), Municipality of Zongolica, Veracruz, Mexico (CNCR 26641). Remarks. The term “conical projection” of the latero-caudal portion of the distal border of the apical cavity from the original description by Álvarez et al. (2012) is now changed to “conical lobe”, since the former suggests the fusion of two plates in a single apical process (Hobbs 1942, 1945; Smalley 1964); however, in this case the structure develops directly from the lateral surface.Published as part of Moreno-Juárez, Eric G., Villalobos, José Luis & Álvarez, Fernando, 2022, Two new genera and one new species of freshwater crabs of the subfamily Pseudothelphusinae (Decapoda: Brachyura: Pseudothelphusidae) from southwestern Mexico, pp. 24-36 in Zootaxa 5200 (1) on pages 32-33, DOI: 10.11646/zootaxa.5200.1.2, http://zenodo.org/record/725183
Ptychophallus costaricensis Villalobos 1974
No full textPtychophallus costaricensis Villalobos, 1974 (Figs. 16, 17, 80) Ptychophallus costaricensis Villalobos, 1974: 197, figs. 1–4.— Rodríguez 1992: 184.— Campos & Lemaitre 1999: 560 (map).— Villalobos Hiriart & Álvarez 2008: 297 (in list). Ptychophallus coastaricensis [sic]— Ng et al. 2008: 176 (in list). Material examined. COSTA RICA. Cartago Province, Atlantic drainage: 1 male, USNM 1089241, Tapantí National Park, unnamed tributaries, approximately 9 km (road) NW tunnel, 1400 m, 9.72 °N 83.78 °W, 8–9.vi. 1988, C.M. & O.S. Flint Jr. leg.—San José Province, Pacific drainage: 1 male, paratype, UCR-MZ 707, Dota Canton, Santa María, Finca El Cedral, 23.iv. 1972, R. Saenz, leg.; 6 males (cw 17.0, cl 11.6 – cw 25.8, cl 16.0), paratypes, UCR-MZ 721, Dota Canton, creek approximately 5.0 km of house at El Cedral Farm, road to Naranjo, 5.v. 1972, R. Saenz, V. Ramírez & C. Villalobos leg.; 2 males (cw 23.1, cl 14.0; cw 33.5, cl 20.3), 10 females (cw 18.6, cl 11.8 – cw 25.2, cl 15.2), paratypes, UCR-MZ 720, San Lucas, creek aprox. 5.5 km of house at El Cedral Farm, road to Naranjo, 7.v. 1972, R. Saez, V. Ramírez & C. Villalobos leg.; 6 males (cw 15.5, cl 10.1 – cw 29.1, cl 18.0), 4 females, paratypes, UCR-MZ 722, Naranjo, Naranjo River, Guillermo Solís’ farm, 7.v. 1972, R. Saenz, V. Ramírez & C. Villalobos leg.; 1 male (cw 33.8, cl 21.4), 1 female (cw 28.2, cl 17.5), paratypes, INPA 1844, same data; 1 male (cw 34.8, cl 20.3), 1 female (cw 26.5, cl 16.8), paratypes, TMNH 6408, San Isidro de El General, Los Tubos Creek, 7.iv. 1972, C. Villalobos leg.; 2 males plus 1 G 1, paratypes, UCR-MZ 719, San Isidro de El General, Los Tubos Creek, 1 km W Boquete, Adan Bonilla’s farm, 7.iv. 1972, V. Juárez leg.; 4 males, 2 females, UCR-MZ 742 - 0 1, San Isidro del General, Los Tubos Creek, 1 km W Boquete, Adan Bonilla’s farm, 22.iv. 1972, V. Juárez leg.; 1 male, INPA 1858, same data. Description of gonopod. Straight in mesiocaudal view, apex bent approximately 90 ° in laterocephalic direction. Marginal suture on mesial side, straight; marginal process rounded, produced beyond distal border of apex. Mesial process small, subtriangular, slightly downturned, with 2 short spines on laterocephalic side, proximal spine slightly sharper than distal spine. Lateral process long, strongly bilobed, approximately 70 % of stem length (from proximal opening to caudal border of apex), usually as wide as lateral end of apex, with deep notch in the middle; lateral border with deep median incision, both lobes subequal in size. Distocaudal ridge short, narrow well separated from distal border of lateral process by distinct depression. Apex oblong, narrow, with narrow notch on lateral end. Field of apical spines well developed, facing towards cephalolateral side. Holotype and type locality. Male, cw 44.4, cl 27.1, UCR-MZ 392. Costa Rica, San José Province, San Isidro del General (Pacific drainage). The holotype could not be found in the UCR-MZ holdings. It is a male originally separated from lot CVS-079- 72 (number in C. Villalobos field notes) (Villalobos, 1974), which received the catalogue number UCR-MZ 719. The G 1 of one male paratype from this same lot is illustrated in Figs. 15, 16. Distribution. Ptychophallus costaricensis is distributed on both Atlantic and Pacific drainages of central southern Costa Rica, Cartago and San José Provinces (Fig. 80). Remarks. Ptychophallus costaricensis is very close to P. micracanthus. They differ by the number of spines on the laterocephalic side of the mesial process (two in P. costaricensis [Fig. 15]; one in P. micracanthus [Fig. 34]), and the development of the marginal process (more developed, reaching beyond the caudal border of apex in P. costaricensis [Fig. 16]; less developed, reaching about the caudal border of apex in P. micracanthus [Fig. 33]). Ptychophallus costaricensis occurs only in central-southern Costa Rica, whereas P. micracanthus is distributed in central Panama.Published as part of Magalhães, Célio, Wehrtmann, Ingo S., Lara, Luis Rólier & Mantelatto, Fernando L., 2015, Taxonomy of the freshwater crabs of Costa Rica, with a revision of the genus Ptychophallus Smalley, 1964 (Crustacea: Decapoda: Pseudothelphusidae), pp. 301-344 in Zootaxa 3905 (3) on page 309, DOI: 10.5281/zenodo.23512
Chordodes ruginosus Villalobos, Zanca & Yanez, 2009, n. sp.
No full textChordodes ruginosus n. sp. (Figures 6, 7) Holotype: 1 female [QCAZI 2040], [MLP 5943] Orellana province, Taracoa, Pozo Primavera, Ecuador (1235m) (04º 90´18 ´´ S – 75 º 72´74 ´´ W). J. Vieira col. 21 /07/ 2006. Host: Undetermined Mantids. Etymology. The name (L. ruga, meaning wrinkled) refers to the surface characteristics shown by the simple and tubercle areoles. Description. The body color is dark brown, the anterior tip is tapering. The posterior end is rounded. The body length is 132 mm; the maximal diameter is 1.4 mm. The cuticle contains six areolar types (Figs. 6 A, 6 B): simple, tubercle, thorn, bulging, crowned and circumcluster areoles. Simple areoles are more numerous, low, very slender with a wrinkled surface (like a blackberry) and are perpendicular to the body axis. Tubercle and thorn areoles occur quite scattered on the cuticular surface (Fig. 6 B). Tubercle areoles have a short tubercle. This tubercle is low (9–12 µm high) with a round apex. Thorn areoles, with a robust spine on top (25–29 µm high); originate from a ring-shaped basal structure. Bulging areoles resemble simple areoles but are more elevated (24 µm). They are arranged forming groups of 2 or 3 areoles. Two further types of areoles occur in clusters and are elevated 26–28 µm above the surface, circumcluster areoles (15–18 areoles) that are slender with a slightly smooth surface, surround two crowned areoles with a crown of filaments on top (25–27 µm long) (Fig. 6 B). Along the ventral groove of the body, there are crowned areoles with very long filaments (up to 273 µm) (Fig. 7). Comments. This species belongs in a large group of Chordodes species, in which simple areoles are strongly structured into “blackberry” areoles and clusters of crowned and circumcluster areoles are present (see Schmidt-Rhaesa et al. 2008). Chordodes ruginosus resembles C. staviarskii, concerning the presence of the same six types of areoles (see De Villalobos & Zanca 2005). In C. staviarskii the simple areoles are mostly circular in shape, circumcluster areoles are less numerous and the cuticular structures have an irregular distribution. In C. ruginosus the slender form of the simple areoles, the number and the shape of the circumcluster areoles and the distribution pattern of the cuticular structures justify the description of new species.Published as part of Villalobos, Cristina De, Zanca, Fernanda & Yanez, Alvaro Barragan, 2009, Three new species of Chordodes and new data on hairworms (Nematomorpha) from Ecuador, pp. 37-52 in Zootaxa 2205 on page 44, DOI: 10.5281/zenodo.18970
Sylvathelphusa kalebi Villalobos & Álvarez, 2013, n. sp.
No full textSylvathelphusa kalebi n. sp. (Figs. 2, 3) Type material. Holotype: male, cb 17.3 mm, cl 10.5 mm; allotype: female, cb 14.8 mm, cl 9.6 mm; paratype: male, cb 15.1 mm, cl 9.4 mm; La Pera nature reserve, 12 km NE of Berriozábal (16 ° 49 ' 53 " N, 93 ° 17 ' 42 " W; 1178 m), Municipio de Tuxtla Gutiérrez, Chiapas, Mexico; 12 Sep 2008; colls: K. Zárate, I.A. Córdoba, A.J. Flores, C.A. Pérez, G. Salinas; CNCR 26179. Other material examined (non-types). 5 males, cb 12.0- 15.5 mm, cl 7.8-10.5 mm; 2 females 15.0 mm both, cl 9.2-9.5 mm; same locality as holotype; 27 Nov 2008; colls: J.L. Villalobos, C. Hernández, K. Zárate, A. García, C. Enríquez, E. Torres; CNCR 26181. Description. Small sized crabs (cb <17.3 mm). In dorsal view, carapace oval shaped, widest in the middle; dorsal surface flat, punctated, regions faintly indicated; pair of gastric pits barely distinct, oblique, elongated, close to each other on metagastric region; cervical grooves narrow, shallow, straight, not reaching anterolateral margin of carapace (Fig. 2 a). Postfrontal lobes distinct, low, separated by median groove. Frontal region granulated, depressed with respect to carapace surface; inferior frontal border granulated, projected, bilobed, with deep medial notch; superior frontal border straight, bilobed in dorsal view, lobes defined by row of sharp granules separated by shallow mesial notch (Fig. 2 a). Superior margin of orbit formed by granules, continuous with inferior frontal border; lower orbital margin marked by row of granules; exorbital angle projected, narrow, acute. Lateral margins of carapace serrated, anterior half with sharp granules, posterior half smooth. Posterior margin of carapace slightly concave. Eyes with cornea pigmented, faintly faceted, not filling orbits (Fig. 2 d). Epistome narrow, epistomial tooth low, triangular, slightly deflexed, borders smooth (Fig. 2 c). Suborbital, subhepatic and pterygostomian regions pubescent. Third maxilliped with ratio exopod/ischium 0.47, ischium with outer margin slightly sinuous, inner margin straight, external margin of merus widely curved (Fig. 2 c). Aperture of efferent branchial channel subrectangular. Chelipeds subequal in size and shape; merus subtriangular in cross section, 4.3 times as long as wide, superior internal border with row of conical and acute spines, increasing in size distally, alternating with small spinules. Carpus smooth, prominent and acute subdistal spine on inner side. Chela slender (length/height 3.6), surface smooth, with faint minute teeth on upper and lower rounded borders. Fingers slender, as long as palm, not gaping, tips crossing; both fingers with sharp triangular teeth on cutting edges, becoming smaller distally; dactylus with alternating large and small teeth (Fig. 2 b). Thoracic sternites of third maxillipeds and first pereiopods completely fused, except for small notches at lateral edges of sternum; sternal sulci s 4 /s 5, s 5 /s 6 and s 6 /s 7 distinct, just failing to reach midline of thoracic sternum. All abdominal segments free. Telson subtriangular, narrow; lateral margins distinctly straight, tip rounded. Male gonopod not completely chitinized, latero-mesially compressed, slightly arched in caudal and cephalic views, distal half wider. In mesial view, marginal suture almost straight along most of gonopod, slightly concave in middle (Fig. 3 a). Marginal plate between caudal and mesial surfaces, without marginal setae basally; distal half widening distally, ending in triangular apical projection overreaching mesial process, lying on caudal margin of apical projection, ending in acute distal margin with triangular tubercle (Fig. 3 a). Caudal portion of marginal plate limited by lateral suture, distal third rounded. Mesial surface ending in tongue-like apical projection, distocephalically directed, longer than apical projection of marginal plate, distal margin broad, rounded, sculptured with shallow longitudinal grooves and scattered subacute granules (Fig. 3 a). Mesial process as triangular, stout, acute spine, directed cephalad, forming 90 º angle with respect to principal axis of gonopod (Fig. 3 a). Caudal surface with medial shallow, rounded notch. Lateral suture visible through proximal two thirds, apically separating marginal plate from caudal surface. In cephalic view, mesial process conical, stout, acute (Fig. 3 b). Lateral surface with small triangular distomedial tubercle (Fig. 3 b); distal third of laterocaudal portion forming high, rounded border, with central depression (Fig. 3 b). Distal end of marginal plate subacute. In lateral view, caudal surface with medial constriction (Fig. 3 c). Lateral surface becoming broader caudally, forming on distal third ample plate with high caudal margin, laterally directed (Fig. 3 c). Deep suture at base of apical projection of mesial surface, separating lateral and mesial plates (Fig. 3 c). In caudal view, gonopod slightly arched laterally (Fig. 3 d). Distal half with lateral margin expanded, with low, distal, triangular prominence (Fig. 3 d). In apical view, apical portion of marginal plate ending in acute tip (Fig. 2 e). Mesial surface with broad, rounded distal margin, with few granules (Fig. 2 e). Mesial process conical, stout, acute, directed cephalically (Fig. 2 e). Apical cavity covered by hood formed by distal crest of lateral surface. Opening of spermatic channel located caudally (Fig. 2 e). Color in life. Carapace dark olive or gray, chelipeds white and ambulatory legs light brown, thoracic sternites and abdomen light yellow. Etymology. The species is named after Kaleb Zárate, biologist and speleologist, who first discovered this population. Remarks. This is the first record of a completely terrestrial pseudothelphusid crab in Mexico, although Pseudothelphusa parabelliana Álvarez, 1989, Odontothelphusa maxillipes (Rathbun, 1898), Potamocarcinus magnus (Rathbun, 1895) and Raddaus bocourti (A. Milne-Edwards, 1866), have been observed at considerable distances from the nearest stream. The crabs representing the new species were found in a tropical rainforest NE of the town of Berriozábal, at altitudes ranging from 950 to 1200 m. They were found under limestone in crevices, where the soil is humid and contains abundant organic matter composed of leaf litter in different stages of decomposition. No streams, rivers or other aquatic bodies were observed nearby.Published as part of Villalobos, José Luis & Álvarez, Fernando, 2013, Two new genera and three new species of freshwater crabs (Crustacea: Pseudothelphusidae: Potamocarcinini) from Chiapas, Mexico, pp. 457-470 in Zootaxa 3599 (5) on pages 459-463, DOI: 10.11646/zootaxa.3599.5.3, http://zenodo.org/record/21881
Libro intitulado los problemas de Villalobos :|bq[ue] co[n]tiene dos tractados. El primero es de cuerpos naturales, el segundo es de cosas morales ... ; [y dos dialogos d[e] medicina y el tratado de las tres gra[n]des y vna cancion, y la comedia de Amphytrion]
No full textDatos de autor y título tomados del incipitDatos de publicación tomados del colofón. - Marca tip. en v. de última h.Sign.: A-L6, M8. - Texto a dos col. con apost. marg. - L. got. - Inic. grab.La antep. de la segunda obra con escudo xil. imperial (ocupa la mayor parte de la página)La comedia del Plauto llamada Amphytrio[n], q[ue] traduzia el doctor Villalobos, la qual gloso el en algunos passos obscuros, nuevame[n]te impressa y emendada por el mismo author, h. L-LXXII, con antep. propi
Pseudothelphusa purhepecha Ojeda-Escoto, Villalobos & Álvarez, 2017, n. sp.
No full textPseudothelphusa purhepecha n. sp. (Figs. 6, 7) Pseudothelphusa (Pseudothelphusa) dugesi belliana Rodríguez & Smalley, 1969: 81 fig. 13, pl. 9. Pseudothelphusa belliana Rodríguez, 1982: 135, fig. 88. García-Madrigal & Bastida-Zavala, 1999: 921. Pseudothelphusa belliana form Agua Blanca Villalobos, 2005: 306, pl. 16. Type material. Male holotype, cl 21.9 mm, cb 36.7 mm; 1 female allotype cl 26.0 mm, cb 41.9 mm; El Serpentín in San José Purúa (19°29' N, 100°29' W; 1,900 m asl), Municipality of Jungapeo, Michoacán, Mexico; 1 January 1957; coll. A. Padilla; CNCR 31746. 4 male paratypes, cl 11.4–17.3 mm, cb 17.3–27.9 mm; 4 female paratypes, cl 15.5–24.9 mm, cb 24.4–41.5 mm, same locality, date and collector as holotype; CNCR 31747. All of these individuals, the holotype, allotype, and paratypes, were considered as Pseudothelphusa dugesi belliana and included as part of the material examined by Rodríguez & Smalley (1969: 81, fig. 13a–e), catalog number UNAM 4 25 61. Other material examined. 7 males, cl 13.2–15.0 mm, cb 21.0– 24.2 mm; 11 females, cl 12.4–18.3 mm, cb 19.3–29.2 mm; Agua Blanca (18°57' N, 103°19' W; 500 m asl), Municipality of Coalcomán de Vázquez Pallares, Michoacán, Mexico; 5 July 1986; coll. O. Meade; CNCR 5871. 2 males, cl 20.0, 24.6 mm, cb 32.4, 41.2 mm; 2 females, cl 18.0, 19.8 mm, cb 29.0, 33.7 mm; same locality and date as CNCR 5871; colls. R. Lamothe and L.J. Rangel; CNCR 8362. 2 males, cl 20.0, 24.0 mm, cb 32.4, 41.0 mm; 2 females, cl 18.0, 19.8 mm, cb 29.0, 33.7 mm; same locality as 5871; without collection date; coll. anonymous; CNCR 8816. 7 males, cl 10.8–20.7 mm, cb 16.3– 33.2 mm; 7 females, cl 8.3–19.6 mm, cb 11.7–31.9 mm; Tuxpan, near San José Purúa, Municipality of Zitácuaro, Michoacán, Mexico; 1–2 November 2004; coll. F. Álvarez; CNCR 27310. Description. Dorsal surface of carapace smooth, puntacted, central portion flat (Fig. 6 A). Front without superior border, curved downward to reach inferior frontal border, visible in dorsal view (Fig. 6 D). Front smooth, faintly bilobed. Postfrontal lobes well formed, portion of carapace anterior to postfrontal lobes convex (Fig. 6 D). Median frontal groove shallow, well-marked, separating postfrontal lobes, becoming obsolete little after. Cervical grooves well marked, wide, straight, reaching anterolateral margins. Regions moderately indicated, gastric, branchial, intestinal swollen; cardiac depressed (Fig. 6 A). Anterolateral margin not prominent in lateral view, serrated, with blunt tubercles, 22–23 in males, 15–17 in females, from cervical groove to branchial region; area between orbit, cervical grove with 6–7 rounded granules. Posterior margin straight in males (Fig. 6 A), slightly concave in females. External angle or orbit faintly developed, granulated, with shallow basal notch; internal angle inside orbital cavity with moderate keel. Antennal basal article, separated from anterior frontal margin by narrow hiatus (Fig. 6 D). In frontal view, antennules, antennular fossae concealed by front, in juveniles antennular fossae partially visible; ventrally, antennular fossae wider in middle, inferior frontal margin slightly sinuous; interantennular septum almost completely covered by carapace. Opercular plate of antennal gland ovoidal, with small middle constriction, tuft of setae on external third. Epistome, area surrounding buccal cavity with dense patch of setae; lateral portions with rounded granules; epistomal tooth small, triangular, granulated, with apex directed downwards at same level of interantennular septum, between third maxilliped palps when closing buccal cavity (Fig. 6 D). Opening of efferent branchial channel subquadrangular, ratio wide/length 1.19 (range 1.0 to 1.6; n= 40). Third maxilliped with ischium trapezoidal, slightly longer than wide; merus narrower than ischium, anterior margin rounded with shallow, rounded notch next to insertion of palp (Fig. 6 C); ratio exopod/ischium 0.66 (range 0.57 to 0.77; n= 40). Chelipeds asymmetrical. Merus of larger cheliped with row of blunt tubercles along internal margin, carpus with strong triangular spine on internal margin. Internal surface of chela smooth, globose; fingers gaping, curved inward distally, with low, triangular teeth on cutting edges (Fig. 6 B). Dactylus rather narrow, widely curved, dorsally ornamented with several longitudinal rows of small pits, black granules. Male gonopod moderately strong. In mesial view, distal crest of caudomarginal process curved, higher than lateral crest. Distal lobe of caudomarginal projection ending as triangular tooth, separated from mesial process by V-shaped notch, apex reaching beyond cephalic margin of proximal lobe. Proximal lobe longer than wide, reniform, proximally directed (Fig. 7 A). In cephalic view, distal lobe of caudomarginal projection directed cephalically, as proximal lobe; internal surface swollen, in contact with internal expansion of mesial process (Fig. 7 B). Proximal lobe parallel to principal axis of gonopod; internal surface swollen, cephalic margin straight. Mesial process as triangular laminar expansion; lateral margin ending in two teeth, latero-caudally directed; distal one small, subacute, proximal one larger, strong, acute. Superior border of process slightly concave, extending internally to close apical cavity cephalically, touching internal surface of caudomarginal projection; same border forming central crest of apical cavity to delimit field of spines (Fig. 7 B). Inferior border widely convex. In caudal view, caudal surface of mesial process smooth; distal crest widely concave; caudomarginal crest higher than lateral one (Fig. 7 D). In apical view, cavity U-shaped, opening of sperm channel in caudal position; field of spines wide, close to lateral crest, with about 50 terminal pore setae, some organized in central tuft; mesial surface armed with hook-shaped spines, caudal, lateral surfaces with spinules (Fig. 7 E). Geographical distribution. Known from several localities in the state of Michoacán: Agua Blanca (19°30'04" N, 100°22'22" W; 1,690 m asl), Municipality of Zitácuaro; El Serpentín in San José Purúa (19°29' N, 100°29' W; 1,460 m asl), Municipality of Jungapeo; Río Frío (19°04'15" N, 101°25' W), Río San José (19°03'30" N, 101°26'01" W), and Río Guayabo (19°04'15" N, 101°26'53" W), Municipality of Turicato; Huiramba (19°32'13" N, 101°26' W), Municipality of Huiramba; 1 km from Tres Palos (18°59' N, 101°41' W), and 1 km from Carámicuas (19°02'40" N, 101°38'30" W), Municipality of Ario (García-Madrigal and Bastida-Zavala, 1999). These localities are all part of the Tuxpan River basin, interconnected by small tributaries. The region is characterized by an irregular topography formed principally by the Tuxpan shield and an important number of volcanic cones. The thermal springs in San José Purúa and Agua Blanca are inside aquatic parks, part of the Jungapeo geothermal system (Siebe et al. 2007). Etymology. The species name is taken from P’urhepecha, the prehispanic native culture that developed and established through the highlands of NW Michoacán, it is used as a noun in apposition. Remarks. Few variations were noted among the studied specimens. The shape and number of granules along the anterolateral margin of the carapace varied slightly (22–23 in males, 15–17 in females); the granules vary from rounded to subacute, the latter are more evident in females. The posterior margin of the carapace varies from straight in big adults to slightly concave in smaller crabs, both male and female. The distal tooth of the mesial process of the gonopod appears as a rounded granule in small males, changing to a well-developed tooth in large males. The mesial process of the gonopod with two teeth on the lateral margin distinguishes the new species from the related Pseudothelphusa belliana (Rathbun, 1898: 515, fig. 4). Furthermore, this character relates the new species with P. nayaritae Álvarez & Villalobos, 1994, P. rechingeri Pretzmann, 1965, P. lophophallus Rathbun, 1898, and P. parabelliana Álvarez, 1989. It can be distinguished from all of them by the distal lobe of the caudo-marginal projection which is well formed in contrast to the reduction in size seen in P. parabelliana or the reduction to a series of small teeth seen in the other three species mentioned. The distribution of morphologically related species to P. purhepecha n. sp. does not reflect a close geographical distribution; P. nayaritae, P. rechingeri, and P. lophophallus are distributed the states of Nayarit, Durango and Sinaloa, whereas P. parabelliana occurs in the Los Tuxtlas region, Veracruz. Instead, P. purhepecha n. sp. is geographically close to P. dilatata, distributed in the Balsas River basin.Published as part of Ojeda-Escoto, Juan Carlos, Villalobos, José Luis & Álvarez, Fernando, 2017, Three new species of freshwater crabs of the genus Pseudothelphusa (De Saussure, 1857) (Crustacea: Decapoda: Pseudothelphusidae) from Mexico, pp. 559-571 in Zootaxa 4216 (6) on pages 567-570, DOI: 10.5281/zenodo.24244
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