173,626 research outputs found
Tetracanna octonema Goy 1979
No full textTetracanna octonema Goy, 1979 Distribution in South America: medusa—Atlantic Ocean, Brazil, at 16.48°S 39°W (Goy 1979; Migotto et al. 2002).Published as part of M. P. Oliveira 1,16, S P. Miranda 2, *,, Es W. Mianzan 10,, Ro E. Migotto 11,, Ne B. Nascimento 2,11, Eli Nogueira Júnior 12,, Er Quiñones 13,, Izio Scarabino 14,, Tín Schiariti 10,, Io N. Stampar 15,, Tronolone 2, , Quíria B. & Onio C. Marques 2,11, 2016, Census of Cnidaria (Medusozoa) and Ctenophora from South American marine waters, pp. 1-256 in Zootaxa 4194 (1) on page 144, DOI: 10.11646/zootaxa.4194.1.
Eucheilota foresti Goy 1979
No full textEucheilota foresti Goy, 1979 Remarks: the validity of the species and its record have to be checked. Distribution in South America: medusa—Atlantic Ocean, Argentina, at 35.84°S 56.32°W (Goy 1979; Genzano et al. 2008a).Published as part of M. P. Oliveira 1,16, S P. Miranda 2, *,, Es W. Mianzan 10,, Ro E. Migotto 11,, Ne B. Nascimento 2,11, Eli Nogueira Júnior 12,, Er Quiñones 13,, Izio Scarabino 14,, Tín Schiariti 10,, Io N. Stampar 15,, Tronolone 2, , Quíria B. & Onio C. Marques 2,11, 2016, Census of Cnidaria (Medusozoa) and Ctenophora from South American marine waters, pp. 1-256 in Zootaxa 4194 (1) on page 152, DOI: 10.11646/zootaxa.4194.1.
Microprosthema looensis Goy & Felder 1988
No full textMicroprosthema looensis Goy & Felder, 1988 (Figs. 1–3) Material examined. 1 male (cl 5.2 mm), FLMNH UF 53415, Panama, Bocas del Toro, east of Isla Bastimentos, near Cayo Coral (Coral Key), shallow sand flat with abundance of coral rubble and sponges, depth: 3 m, in crevice of large dead coral block, leg. M. Leray, A. Anker & E.C. Rodriguez Guerra, 23 July 2018 [fcn 18-002]; 1 female (cl 4.7 mm), FLMNH UF 53416, Panama, Isla Mamey near Isla Grande, shallow reef flat with strong current, depth: 1.5 m, under large flat piece of coral rubble, leg. A. Anker, 1 August 2018 [fcn 18-007]. Distribution. USA: Florida Keys (Goy & Felder, 1988); Panama: Bocas del Toro and Isla Mamey near Isla Grande (present study). Remarks. The morphological characters of the two Panamanian specimens agree well with those described by Goy & Felder (1988) for M. looensis: (i) carapace covered with numerous blunt to subacute teeth (Fig. 1A, B); (ii) cervical groove distinct (Fig. 1B); (iii) third pereopod with teeth on both carpus and merus (Fig. 1G); (iv) most of the third pereopod surface pubescent, i.e. covered with fine, hair-like setae (Figs. 1G, 2B; see also Goy & Felder 1988: fig. 7D); and (v) pleonites sculptured and armed with blunt teeth (Fig. 1E). Goy & Felder (1988) described and illustrated the rostrum of the holotype, which has two distal (= pre-orbital) teeth on the dorsal margin and no teeth on the ventral margin. However, in the Panamanian specimens, the rostrum is rather variable: the male has three pre-orbital teeth on the dorsal margin and one tooth on the ventral margin (Fig. 1A, C), whilst the female possesses two small pre-orbital teeth and one small subapical tooth dorsally and two small subapical teeth ventrally (Fig. 1H). According to Goy & Felder’s (1988) illustrations of the holotype, none of the teeth on carapace attains the post-orbital margin, whereas the Panamanian male has two particularly large teeth that reach slightly beyond the post-orbital margin in dorsal view (Fig. 1B). The previously unknown male thoracic sternum (Fig. 1D), pleon (Fig. 1E) and third pereopod (Fig. 1G) are shown to complete the original description of M. looensis. The third pereopod is noticeably stouter in the Panamanian male than in the Panamanian female (Figs. 2A, 3), but not significantly stouter than in the ovigerous female from Florida (holotype, cf. Goy & Feder 1988: fig. 7D). We also noted some rather minor differences in the armature of the third pereopod, for example, between the Panamanian male and the holotype. For instance, in the holotype, the dorsal surface of the merus of the third pereopod bears two adjacent teeth subdistally, whilst in the Panamanian male, it has only one tooth (cf. Fig. 1G and Goy & Felder 1988: fig. 7D). The ventral surface of the merus in both the holotype and the Panamanian male has two large widely spaced teeth, however, in a slight different positions (cf. ibid.). Similarly, in the holotype, the dorsal surface of the third pereopod carpus is armed with a series of small teeth in its proximal half and one larger tooth at about its 0.7 length (Goy & Felder 1988: fig. 7D). In contrast, in the Panamanian male, the configuration of the dorsal carpal dentition is fairly different, with two larger and several smaller teeth, in addition to a stout distal tooth (Fig. 1G). However, all these differences can be interpreted as intraspecific variation, which indeed seems to be common in the genus Microprosthema (Goy & Martin 2013). Goy & Felder (1988) described and illustrated the mandible of the holotype of M. looensis. The mandibular palp of the holotype is subdivided, according to the authors, into two distinct articles, the peduncular article and the expanded terminal article (Goy & Felder 1988: fig. 6B, C). However, the presence of a two-articulated palp in the mandible of M. looensis contradicts previous observations made in various other stenopodidean taxa, the vast majority of which are characterised by the presence of a three-articulated palp (Goy 2010: fig. 65.6; but see Quintal & Goy 2019: fig. 2F). Therefore, one of the mandibles of the Panamanian male was dissected and its palp illustrated (Fig. 1F) for comparison with that of the holotype. Interestingly, the mandibular palp of the male shows the typical three-articulated condition, as described for most other species of Microprosthema and Stenopodidea in general. Thus the biarticulated mandibular palp of the holotype of M. looensis may be an aberrant condition of that specimen or at least may be viewed as untypical for the species. The colouration of M. looensis was briefly described by Goy & Felder (1988) as “Carapace and abdomen [= pleon] whitish tan; antennae, telson, uropods, and all appendages white. Eggs are pale green”. This description was possibly based on a superficial observation of the living shrimp against a solid black background. The carapace, pleon and third pereopods of both Panamanian specimens are pale pink with a slight yellowish tinge (Figs. 2, 3), which is due to the presence of numerous, scattered, small, red chromatophores, visible only under a dissecting microscope or with a digital camera zoom. The tail fan, antennular and antennal flagella, and all other appendages (first and second pereopods, ambulatory pereiopods) are white to semi-translucent with a pale yellow tinge (Figs. 2, 3). Microprosthema is represented by five other species in the western Atlantic, in addition to M. looensis. These species are: M. granatense Criales, 1997, M. inornatum Manning & Chace, 1990, M. manningi Goy & Felder, 1988, M. semilaeve (von Martens, 1872) and M. tortugasensis Goy & Martin, 2013 (Goy & Felder 1988; Criales 1997; Goy & Martin 2013). Although De Grave et al. (2016) questioned the presence of M. inornatum in the western Atlantic, based on a single record from a deep-water locality (63–110 m) in the Gulf of Mexico (Goy & Martin 2013), Dr. Joseph W. Goy (pers. comm.) confirmed the identity of the specimen in question (USNM 1541992) as M. inornatum. Microprosthema looensis is the only western Atlantic species with the carapace densely covered with spines, the third pereopods pubescent, and the pleon sculptured and armed with small scattered spines (see also key in Goy & Martin 2013). It is also the second species reported from Panama, together with M. semilaeve, which is much more common on shallow sand flats with abundant coral rubble (De Grave & Anker 2017).Published as part of Ferreira, Luciane Augusto De Azevedo, Leray, Matthieu & Anker, Arthur, 2020, New findings of the stenopodidean shrimp Microprosthema looensis Goy & Felder 1988 (Decapoda: Stenopodidea: Spongicolidae), pp. 445-450 in Zootaxa 4729 (3) on pages 445-449, DOI: 10.11646/zootaxa.4729.3.11, http://zenodo.org/record/363274
Tabulatur des Tanzes
No full textC. Schmidt-GoyBildunterschrift: „Tabulatur des Tanzes“, „Die sanfte oder Fliederblüten=Weis“, „Die strenge oder historisch=plastisch= / mimisch=dekorative Weis“, „Die wilde oder expressionistische Weis" sowie "946“.Herstellungsangaben: "C. Schmidt-Goy“Es handelt sich um einen Zeitungsauschnitt aus: Die Jugend, Jg. 22, H. 49, 1917, S. 946
Inventaire de la céramique moderne et contemporaine
No full textIn : GOY (C.) dir. – Belfort (90), Parking de la MAT. Du couvent des capucins à l’hôpital militaire. Rapport de diagnostic, Dijon : Inrap Grand Est su
Inventaire de la céramique moderne et contemporaine
No full textIn : GOY (C.) dir. – Belfort (90), Parking de la MAT. Du couvent des capucins à l’hôpital militaire. Rapport de diagnostic, Dijon : Inrap Grand Est su
Rudolf Goy, Die Ueberlieferung der Werke Hugos von St. Viktor, Monographien zur Geschichte des Mittelalters, Band 14, 1976
No full textM. C. Rudolf Goy, Die Ueberlieferung der Werke Hugos von St. Viktor, Monographien zur Geschichte des Mittelalters, Band 14, 1976. In: Revue des Sciences Religieuses, tome 53, fascicule 2, 1979. p. 191
S. Cantrelle, C. Goy et C. Munier, dir., Histoire d'un quartier de Montbéliard (Doubs). Le bourg Saint-Martin (Xllle-XXe s.)
No full textGarrigou Grandchamp Pierre. S. Cantrelle, C. Goy et C. Munier, dir., Histoire d'un quartier de Montbéliard (Doubs). Le bourg Saint-Martin (Xllle-XXe s.). In: Revue archéologique du Centre de la France, tome 40, 2001. pp. 309-310
Spongicola liosomatus Quintal & Goy 2019, sp. nov.
No full textSpongicola liosomatus sp. nov. (Figs. 1–4) Material examined. HOLOTYPE: Venezuela. Caribbean Sea, R / V Dr. Fridtof Nansen, stn 477, northern Blanquilla Island, 11°54’N, 64°17’W, 135–160 m, 2.VI.1988, bottom trawling net, male (cl 5.2 mm) inside hexactinellid sponge (USNM 1573503). PARATYPES: Venezuela. Caribbean Sea, R / V Fridtof Nansen, stn 477, northern Blanquilla Island, 11°54’N, 64°17’W, 135–160 m, 2. VI.1988, bottom trawl net, 2 males (cl 3.8mm; cl 5.4 mm) inside hexactinellid sponge (USNM 1573504); 1 female (cl 4.7 mm) inside hexactinellid sponge (MMM-Crus-0300). Diagnosis. Small commensal spongicolid shrimp associated with sponges, with slightly depressed body bearing few carapacial spines. Rostrum with dorsal margin slightly upturned toward distal end, bearing 2 to 4 dorsal teeth and small projection at posterior end, ventrally with distal tooth. Carapace without postrostral submedian spine, hepatic spine present in line with antennal spine, 3 or 4 anterolateral spines, 4 to 6 pterygostomian spinules. Abdominal terga entirely smooth, pleura of first to fifth somites broadly rounded without marginal spines. Mandibular palp 2-segmented. First and second pereiopods cutting edges with membranous ridge, palm inflated; chela of third pereiopod with serrated upper margin, lower margin glabrous. Description. (Holotype, male). Rostrum (Figs. 1, 2A) compressed, triangular at base, reaching slightly past end of second segment of antennular peduncle; dorsally upturned toward distal end, bearing 2 dorsal teeth and small projection at posterior end, ventrally with distal tooth. Carapace (Figs. 1, 2A) subcylindrical, surface glabrous; cervical groove absent; postrostral submedian teeth absent; antennal tooth small, acuminate; hepatic tooth small; 3 anterolateral spines; pterygostomial angle produced, rounded with 6 spinules. Abdomen (Figs. 1, 2A) smooth; first to sixth terga glabrous; pleura of first to fifth somites broadly rounded without marginal spines; no spiniform process on sixth somite. First somite with distinct transverse carina anteriorly flowing into transverse groove. Second and third somites with faint transverse grooves about midlength. Telson (Figs 2A, 4A) lanceolate equally long as broad; with 2 longitudinal carinae on dorsal surface provided with 2 or 3 spines; 2 more spines between bases of carinae; 1 dorsal spine present on each side of base of telson; lateral margin with 3 or 4 spines on each side; posterior margin fringed with long setae (not shown) and provided with 3 spines, 2 subterminal flanking large median terminal. Eyes (Figs. 1, 2A, D, E) comparatively large, eyestalk covering proximal one-third of antennal scale in dorsal view; with 4 spinules on inner margin, 2 spinules on dorsal surface; cornea darkly pigmented, semi-globular, about same width as eyestalk. Antennular peduncle (Figs. 2A, B) with basal article more than twice as long as second segment, third segment about 0.75 second segment. Stylocerite small, acuminate, reaching near distal margin of third segment, large spine on inner margin reaching middle of second segment. Second segment with distolateral margin bearing long spine. Outer antennular flagellum slightly longer than inner flagellum, proximal portion bearing 7 paired aesthetascs, no setae present. Antenna (Fig. 2C) short. Basicerite stout, bearing distolateral spine, ventrolateral margin unarmed; antennal scale semicircular, about twice as long as broad; outer margin slightly concave, with 6 teeth, terminal one largest, dorsal surface with 2 distinct longitudinal carinae. Antennal flagellum shorter than carapace. Mandible (Fig. 2F) with 2-segmented palp; incisor and molar processes fused bearing few teeth. Maxillule (Fig. 2G) with simple, slender endopod tapering distally; coxal endite suboval, with submarginal row of setae on outer surface; basal endite moderately broad, truncated distally with several setae. Maxilla (Fig. 2H) with curved, slender endopod; coxal and basal endites deeply bilobed; scaphognathite well developed; posterior lobe slightly elongate, suboval terminal margin with very long setae. First maxilliped (Fig. 2I) with endopod unsegmented; basal endite large, subtriangular, with concave mesial margin; exopod well developed; epipod large, distinctly bilobed. Second maxilliped (Fig. 2J) with endopod 7-segmented with coxa and basis fused; ischium short, distal margin sharply pointed; merus and carpus of equal length; propodus cup–shaped; dactylus suboval equal propodal length; podobranch simple; exopod well developed, flagellar. Third maxilliped (Fig. 2K) with 7-segmented endopod; basis short; ischium elongate, distal margin with 3 spines; merus half ischial length, outer margin with small midlength spine, distal end with 1 large, 1 small spine; carpus with well developed distal grooming apparatus, equal propodal length; dactylus tapering distally; exopod rudimentary. First pereiopod (Fig.3A) slender, glabrous, chelate; fingers with membranous ridge on cutting edge. Dactylus 0.33 times length of palm distally ending with setal brush; palm slightly inflated also with setal brush. Carpus longest segment, about 1.5 times as long as chela. Grooming apparatus lacking. Merus about 0.8 times length of carpus. Ischium shorter than merus. Second pereiopod (Fig. 3B) glabrous, slightly longer than third pereiopod, overreaching antennal scale with carpus. Finger cutting edges composed of membranous ridges; dactylus and palm ending in setal brushes; palm inflated, twice dactylar length. Carpus, merus equal in length; ischium 0.5 meral length. Third pereiopod (Fig. 3C) chela large, strongly compressed laterally, almost equal to carapace length; ventral margin glabrous; dorsal margin distally serrate with 7 teeth; fingers distally curving inward, crossing, each ending in sharp point, opposable margin of movable finger with prominent median tooth fitting to large opposing concavity; cutting edge distally armed with 4 small teeth. Carpus triangular, dorsomesial distal angle produced into subtriangular lobe ending in 2 large teeth. Merus 0.7 length of palm, with 1 distal tooth dorsally, ventromedially with large tooth, distal end with large tooth. Ischium 0.5 meral length, strong distodorsal tooth. Fourth pereiopod (Figs. 3D, F) long, slender, sparsely setose; dactylus short, biunguiculate, ventral margin with small accessory tooth at base of ventral unguis; propodus 3.0 times dactylar length, with 12 movable spinules on ventrodistal margin; carpus 1.2 length of propodus, with ventromedial, ventrodistal spinules; merus 0.8 carpal length, glabrous; ischium distinctly shorter than merus. Fifth pereiopod (Fig. 3E) similar to fourth, with only ventrodistal carpal spinule, 8 movable ventrodistal spinules on propodus. Gill formula typical for genus (cf. Saito & Komai 2008: table 1). First pleopods unilobed, small, 2-segmented, feebly setose. Second to fifth pleopods bilobed, each fringed with long setae. Uropods (Fig. 4A) with protopod unarmed. Exopod with 8–10 teeth on outer margin; dorsal surface with 2 longitudinal ridges; inner and distal margins with long setae. Endopod with 10 or 11 teeth on outer margin; dorsal surface with longitudinal ridge; inner and distal margins with long setae. Paratypes similar to holotype. No sexual dimorphism. Rostrum with 2 to 4 dorsal spines; pterygostomian spines vary from 2 to 6 on each side of carapace; telson with 5 or 6 lateral spines. Third pereiopods (Figs. 4B, C) with palm bearing 8 or 9 small teeth on dorsal margin, cutting edge lacking small distal teeth; merus with 5 to 7 teeth on ventral margin. Coloration. Carapace translucent with scarce red spots. Pereiopods translucent with red spots; carpus of fourth and fifth pereiopods with more intense red spots. Antennules with red stripes intercalated with translucent spaces; antennal flagellum similarly banded but starting 1/3 from the base. Distribution. Known only from the type locality, off northern Blanquilla Island, Venezuela, Caribbean Sea, 135– 160 m. Host. No information on the host was available. Etymology. From the Greek “liosomatos” (= smooth-bodied), in reference to the reduced armature on the carapace and abdomen of the new species. Remarks. The present new species is referred to the genus Spongicola because of the rudimentary exopod on the third maxilliped and the compressed chelae of the third pereiopods. The presence or absence of an exopod on the third maxillipeds has been considered important in separation of genera within the Spongicoloidae (de Saint Laurent & Cleva 1981; Holthuis 1993; Saito & Takeda 2003; Saito & Komai 2008; Goy 2010). However, the development of the exopod of the third maxilliped has been shown to be variable among species of Microprosthema (Saito & Anker, 2014). Moreover, species of Spongiocaris and Spongicoloides show some variation in the number of branchial exites (de Saint Laurent & Cleva 1981; Garcia Raso 1996) where certain exites can be missing, rudimentary, or well developed in specimens of the same species collected at one location. Differences are also evident in relation in the size and degree of development in some of the spongicoloids. These facts, plus the fragility of the gills and the relatively small number of known specimens, makes the usefulness of branchial formulae in separating species of spongicoloids suspect. A reappraisal of the characters used to separate the genera within Spongicoloidae is necessary (Saito & Takeda 2003; Chen et al. 2016). Spongicola liosomatus sp. nov. appears closest to S. levigatus, widely distributed in the western Pacific (Saito & Komai 2008). These two species lack a postrostral submedian spine on the carapace; have smooth, rounded abdominal pleura; and have an unsegmented endopod on the first maxilliped. The carapacial and abdominal spination shows similarity to S. teres (Komai, 2015) from French Polynesia and the 2-segmented mandibular palp has only been found in Microprosthems looensis from the Florida Keys (Goy & Felder 1988). The new species has inflated palms of the first and second pereiopods as seen in S. inflatus (de Saint Laurent & Cleva 1981). However, S. liosomatus sp. nov. differs from these species and all other members of Spongicola by the unserrated ventral margins of the third pereiopod chela and the membranous ridges present on the cutting edges of the first and second pereiopods. With the transfer of S. cubanicus to the genus Spongiocaris (Saito 2008), S. liosomatus sp. nov. is the first member of Spongicola sensu stricto to be found in the Atlantic Ocean.Published as part of Quintal, Bladimir Rodríguez & Goy, Joseph W., 2019, A new species of the Stenopodidean shrimp Genus Spongicola (Crustacea: Decapoda: Spongicolidae) representing the first record of the genus from the Atlantic Ocean, pp. 393-400 in Zootaxa 4648 (2) on pages 394-399, DOI: 10.11646/zootaxa.4648.2.12, http://zenodo.org/record/335480
Spongiocaris neocaledonensis Goy, 2015, n. sp.
No full text<i>Spongiocaris neocaledonensis</i> n. sp. <p>(Figs. 12–14)</p> <p> Non <i>Spongiocaris yaldwyni</i>.— Goy, 2010: 220, figs. 65.3A, B, 65.4E, 65.12J.</p> <p> <b>Type material</b>. HOLOTYPE: <b>New Caledonia, Norfolk Ridge</b>. SMIB 4, stn DW58, 22°59.8’S, 167°24.2’E, 560 m, 9.III.1989, male, 1 female (MNHN-NA 12000). ALLOTYPE: <b>New Caledonia, Norfolk Ridge</b>. SMIB 4, stn DW58, 22°59.8’S, 167°24.2’E, 560 m, 9.III.1989, female (MNHN-NA 12000). PARATYPES: SMIB 4, stn DW61, 22°59.9’S, 167°22.8’E, 550 m, 10.III.1989, 3 females (1 ov.) (MNHN-NA 12001).— SMIB 4, stn DW63, 22°58.7’S, 167°21.1’E, 520 m, 10.III.1989, 1 female ov. (MNHN-NA 12004).— SMIB 4, stn DW64, 22°55.3’S, 167°16.4’E, 460 m, 10.III.1989, 1 male, (MNHN-NA 12005).— <b>New Caledonia</b>. BIOCAL, stn DW 46, 22°53’S, 167°17’E, 570–610 m, 30.VIII.1985, 1 male, (MNHN-NA 12007).—MUSORSTOM 4, stn CP215, 22°55.7’S, 167°17’E, 485–520 m, 28.IX.1985, 3 females (2 ov.) in <i>Regardella okinseana</i>, (USNM 256952).—BIOGEOCAL, stn CP290, 20°36.91’S, 167°03.34’E, 970– 760 m, 27.IV.1987, 1 female, (MNHN-NA12002). — <b>Loyalty Islands</b>. W. Lifou, N. Baie du Santal, CALSUB, Dive 7, 22°48’S, 167°05’E, 966– 845 m, 25.II.1989, 1 male on gorgonian, (MNHN-NA 12003).</p> <p> <b>Diagnosis</b>. Small commensal spongicolid shrimp associated with sponges or gorgonians, with slightly depressed body bearing few spines; carapace with distinct cervical groove with 4–10 small spines; small branchiostegal spine present; small supraorbital, antennal, and hepatic spines sometimes present; rostrum moderately long reaching to middle of last antennular peduncle segment with 4–6 dorsal, 0–1 ventral, 0–2 lateral spines; pigment of cornea restricted to submedial ring in starburst pattern; unarmed epistome; endopod of maxillule unarmed; setiferous organ of third maxilliped covering distal half of propodal ventral margin; first pereiopod with well developed setiferous organ; males with spines on meri, propodi of third pereiopods, single forward curving ventral medial spine on fifth abdominal pleomere; females with entire third pereiopods glabrous, lacking ventral medial spine on fifth abdominal somite; third pereiopod ventral propodal margin distally with dense fringe of setal brushes; fourth and fifth pereiopods with carpi distinctly 3-segmented; first to fifth pereiopods lack epipods but bear cicatrices; posterior margin of fifth abdominal pleomere ending in 4 or 5 spines, that of sixth pleomere in 3–9 spines; few dorsal spines on sixth abdominal pleomere; uropodal endopodite with 2 terminal dorsal hairs.</p> <p> <b>Description</b>. (Holotype, male MNHN-Na-12000) Rostrum (Figs. 12 A, B) broad at base, slightly compressed anteriorly; moderately long, reaching past middle of last antennular peduncle segment; dorsal margin with 5 spines followed by blunt tubercle; no ventral or lateral spines.</p> <p>Carapace (Figs.12 A, B) with distinct cervical groove bearing 10 small spines. Small spine at rostral base with 2 smaller spines behind located dorsomesially. Small antennal and branchiostegal spines, large spine at pterygosmial angle. Group of 3 small anterolateral spines present. Ventrolateral angle rounded, posterolateral angle of branchiostegite broadly rounded.</p> <p>Abdominal pleomeres (Fig. 12 A) generally glabrous, lacking carinae. First to fifth pleomeres with rounded pleural margins; sixth pleomere’s pleuron ending in acute point. Posterior margin of fifth pleomere ending in 4 spines, that of sixth in 3 spines. Dorsally, sixth pleomere bears row of 3 medial spines, 2 large submedial and 2 smaller lateral spines. Only fifth pleomere with strong median ventral spine, directed anteriorly.</p> <p>Eyes (Figs. 12 A, B) well developed, of moderate size; cornea globular with black pigment restricted to submedial ring arranged in starburst pattern. No spinules on ophthalmic peduncle.</p> <p>Telson (Fig. 12 C) long, broad subtriangular, with median groove flanked by 2 longitudinal carinae. Telsonal base with pairs of small outer and medial spines. Dorsal surface of carinae with left side bearing 5 spines, right bearing 4. Few setae within median groove and outside carinae. Lateral margins of telson with 7 left, 8 right acute teeth. Posterior margin rounded, setose with 10 small teeth.</p> <p>Uropods (Fig. 12 C) well developed, about as long as telson. Basal segment strong with 2 small teeth posterolaterally. Exopodite broader than ovate endopodite; outer margin of left exopodite with 10 teeth, that of right with 11 teeth; dorsal surface of exopodite with 2 strong ridges. Margins of endopodite unarmed, dorsally with 1 strong ridge, terminating in 2 curved dorsal hairs. Unarmed margins of exopoditess, endopodites with long plumose setae.</p> <p>Epistome (Fig. 12 E) unarmed, rounded anteriorly, with rounded lateral extensions. Labrum normally developed, posterior margin acutely rounded. Paragnath slightly bilobed. Thoracic sternites (Fig. 12 D) narrow with 2 submedian spinules on segments 4, 6, 7, 8 and 1 submedian spinules on segment 5.</p> <p>Antennular peduncle (Fig. 12 B) moderately long, extending to middle of scaphocerite. Basal segment more than 2.0 times length of second segment, third same length as second. Outer margin with small rounded stylocerite, few setae; 2 small spines on distodorsal inner margin. Upper and lower flagella short, but well developed; upper flagellum unarmed, lower flagellum consisting of 22 segments with numerous aesthetascs along proximal to midlength.</p> <p>Antenna (Figs. 12 A, B) with stout basicerite with 2 strong distodorsal spines on outer margin, medial carina with rounded distal extremity, sharp hooked spine ventrolaterally. Carpocerite unarmed, short not exceeding first antennular peduncle segment. Antennal flagellum long, slender, unarmed, extending approximately to tip of telson. Scaphocerite broad, slightly quadrangular, dorsally bearing 2 strong longitudinal carinae; outer margin slightly concave bearing 3 small spines proximally, 5 small teeth distally, larger distal most tooth; inner margin convex, fringed with long plumose setae.</p> <p>Mandible (Figs. 12 F, G) robust, with short, fused molar and incisor processes. Molar surface with 3 small teeth; incisor bearing 2 stout teeth distally followed by 6 small teeth proximally. Palp well developed, 3-segmented. Proximal segment without setae; middle segment setose mesially, on distal outer margin; distal segment densely setose laterally and distally.</p> <p>Maxillule (Fig. 12 H) with slender, unarmed, undivided endopodite. Proximal endite broad, roundly truncated distally with numerous simple and compound spinose setae. Distal endite much narrower, rounded, with setae along its margins.</p> <p>Maxilla (Fig. 12 I) with numerous plumose setae on both lobes of coxal and basal endites, proximal coxal endite broadest. Endopodite long, but robust, not exceeding anterior margin of scaphgnathite, with 16 plumose setae along margins. Scaphognathite long, anterior portion broader than posterior portion, with numerous plumose setae along margin.</p> <p>First maxilliped (Fig. 14 A) with 2-segmented endopodite. Proximal segment 2.0 times longer than distal segment with 10 long plumose along outer margin, 3 simple setae on inner margin. Distal segment tapering with 10 long plumose setae on outer margin, stout terminal spine, long plumose seta on inner margin. Basipodite large, rounded anteriorly and posteriorly, slightly concave near middle, with outer margin bearing dense fringe of setae. Coxopodite bilobed with numerous seate. Exopodite well developed, flagellum arising from unsegmented peduncle with 5 proximal and 20 distodorsal long plumose setae. Moderately large epipod with slender proximal and distal lobes approximately equal in length, small arthrobranch also present.</p> <p>Second maxilliped (Fig. 14 B) with 5-segmented endopodite. Dactylus suboval, slightly longer than broad, with dense fringe of setae along distodorsal margin. Propodus equal to dactylar length, densely setose on dorsal margin, ventral margin unarmed, notched proximally, rounded distally. Carpus triangular, narrowed proximally, equal to propodal length, densely setose on outer margin. Merus long, slightly concave medially, about 4.0 times carpal length with fringe of short setae on inner margin, 4 setae on outer margin. Ischium and basis not fused, both with few setae. Coxa with small epipod, arthrobranch and lateral podobranch. Exopodite long, slender, undivided with 3 proximal and 20 distodorsal long plumose setae.</p> <p>Third maxilliped (Fig. 14 C) with well developed, 5-segmnted endopodite. Dactylus slender, tapering, 2.0 times longer than broad, dorsal margin with 3 simple setae, numerous long simple setae mesially and on inner margin. Propodus 2.0 times dactylar length, 4 outer setae, entire inner margin with dense fringe of setae on distal setiferous organ. Carpus about equal propodal length, merus 2.0 times carpal length, ischium equal meral length; all with few setae on outer margins, densely setose in double row on inner margins. Basis without exopodite, sparsely setose on inner margin. Coxa bearing small epipod, two small arthrobranchs.</p> <p>First pereiopod (Fig. 14 D) small, slender, when stretched reaching past scaphocerite, segments with few simple setae on margins. Dactylus less than 0.5 propodal length. Fingers slightly compressed, with hooked highly chitinized tips, overlapping when closed. Cutting edges rather indistinct with propodus and dactylus bearing chitinous ridge along inner margins. Fingers and distodorsal extremity of palm with small tufts of long setae arranged like brushes. Distoventral carpus and proximoventral propodus with setiferous organ. Carpus longest segment, over 2.0 times propodal length. Merus almost equal carpal length, ischium equal to propodal length. Basis and coxa short, unarmed, no epipod present.</p> <p>Second pereiopod (Fig. 14 E) similar to first, but longer, stronger, segments with few simple setae along margins. Fingers with hooked, heavily chintinized tips, overlapping when closed. Cutting edges entire, chitinized ridges along inner dactylar and prpodal margins. Fingers and distdorsal extremity of palm with small tuft of long setae. Carpus longest segment, about 1.5 times propodal length, merus 0.8 carpal length, ischium 0.3 meral length. Basis and coxa short, unarmed, no epipod present.</p> <p>Third pereiopod (Figs. 12 A) largest, strongest, almost as long as entire body length, with few simple setae along margins of segments. Fingers elongate with heavily chitinized hooked crossing tips, distally bearing small tufts of long setae. Dactylus with distal chitinous ridge along cutting edge, proximally with large sharp tooth, smaller rounded tooth. Propodal cutting edge with distal chitinous ridge, proximally fossa for receiving large dactylar tooth, crowned with large rounded tooth followed by serrate ridge. Propodus longest segment with outer margin distally bearing dense fringe of setal brushes. Carpus less than 0.3 propodal length, narrowing proximally. Merus about 0.8 propodal length with distomedial ridge, 5 blunt dorsal spines, 4 blunt ventral spines. Ischium 0.5 meral length, unarmed. Basis and coxa short, stout, unarmed, no epipod present.</p> <p>Fourth, fifth pereiopods (Figs. 14 I, J) long, slender, fifth slightly longer than fourth, with simple setae along margins. Dactyli triunguiculate, with smaller proximal tooth on accessory spine. Unguis short, curved, clearly separated from dactylar corpus. Propodi about 0.3 carpal lengths with 14 movable spines on ventral margins. Carpi longest segments, divided into 3 subsegments, last 2 terminating in movable spine at distodorsal margin. Meri about as long as carpi, ischia 0.3 times meral length. Bases and coxae short, stout, unarmed, without epipods.</p> <p>First pleopod (Fig. 12 A) uniramous, second to fifth biramous, all lacking appendices, bearing no spinules.</p> <p>Maxillipeds Pereiopods</p> <p> I II III I II III IV V <b>Measurements</b> (mm). Holotype: PCL: 8.3; RCL: 11.8; TL: 31.3; Allotype—PCL: 10.5; RCL: 12.6; TL: 34.0. Paratypes & additional material: PCL: 4.3–11.4; RCL: 6.0–13.4; TL: 14.4–38.2. Eggs in 4 ovigerous females were few in number (2–9) while the allotype bore 62 eggs. All of the eggs were in early stages of embryonic development averaging 2.04 × 1.70 mm in size.</p> <p> <b>Distribution</b>. New Caledonia, Norfolk Ridge; Loyalty Islands in 460– 970 m.</p> <p> <b>Etymology</b>. The name <i>neocaledonensis</i> refers to the species type locality, New Caledonian waters.</p> <p> <b>Coloration</b>. Unknown.</p> <p> <b>Type locality</b>. New Caledonia, Norfolk Ridge, 22°59.8’S, 167°24.2’E, 560 m</p> <p>Allotype (female, MNHN-NA-12000) (Figs.13 A–D). Sexual differences from the holotype include: inflated carapace; broader thoracic sternites with fourth, fifth bearing 2 rounded projections, others unarmed; no ventral median spine on fifth abdominal somite; abdominal pleura expanded, overlapping each other; posterior margin of fifth abdominal somite with 5 spines; sixth abdominal somite with row of 3 dorsal spines, 9 posterior margin spines; third pereiopod completely glabrous; pleopods much larger and more setose.</p> <p>Five paratypes and 5 additional specimens agree well with the holotype and allotype and show the same sexual differences. The spination of the third pereiopods varies in the males (Figs. 14 F–H) with larger males being less spinose and the meral distdorsal ridge ending in 1 or 2 spines. Spination of the carapace is variable with some specimens bearing hepatic, superior and inferior orbital spines and others lacking an antennal spine. Other variation in spination include: 4–6 dorsal, 0–1 ventral and 0–2 lateral spines on the rostrum; 4–10 cervical groove spines; 4– 6 outer margin spines on scaphocerite; 8–16 ventral movable spines on the propodi of the fourth and fifth pereiopods; 4–9 spines on the telson’s median carinae; 5–10 spines on telson’s lateral border; 6–12 posteror teeth on telson; and 8–14 teeth on outer margin of uropodal exopodite. This slight variation in body spination probably reflect allometric growth changes and normal variation in the species.</p> <p> <b>Host</b>. Three females were taken from the hexactinellid sponge <i>Regardella okinoseana</i> Ijima, 1901 at MUSORSTOM 4 station 215 and another female from the same sponge species at BIOCAL station 46. The second largest male (MNHN-NA 12003) was collected on a gorgonian octocoral by the CALSUB expedition.</p> <p> <b>Systematic position</b>. <i>Spongiocaris neocaledonensis</i> closely follows the definition of the genus <i>Spongiocaris</i> given by Bruce & Baba (1973). The genus <i>Spongiocaris</i> needs to be revised (Saito & Komai 2008; Goy 2010) since some members assigned to other spongicolid genera actually need to be transferred to <i>Spongiocaris</i>. These species include <i>Spongicola japonica</i> Kubo, 1942, <i>Spongicola cubanica</i> Ortiz, Gómez & Lalana, 1994, and <i>Spongicoloides koehleri</i> (Caullery, 1896). A revision of the genera <i>Spongiocaris</i> Bruce & Baba, 1973 and <i>Spongicolides</i> Hansen, 1908 is in progress to affect these transfers. That said, <i>Spongiocaris neocaledonensis</i> is most similar to <i>Spongicola japonica</i> in rostral, carapacial and telsonal spination, while the new species’ eye pigmentation, morphology of the third pereiopods approaches the conditions seen in <i>Spongicoloides koehleri</i>. In overall appearance <i>S. neocaledonensis</i> resembles <i>S. yaldwyni</i> (Goy, 2010) and its branchial formula is the same as <i>S. yaldwyni</i>, except for the presence of an epipod on the third maxillipeds. The new species can be easily distinguished from the other seven species in the genus by the segmentation of the carpus of the last two pairs of pereiopods, the presence of the third maxilliped’s setiferous organ, and the dense fringe of setal brushes on the third pereiopod propodus.</p> <p> <b>Remarks</b>. <i>Spongiocaris neocaledonensis</i> is the only member of the genus with well developed setiferous organs on the third maxillipeds and first pereiopods; approaching the condition of many members of the family Stenopodidae. All other members of the genus <i>Spongiocaris</i> lack these structures, except <i>Spongicoloides koehleri</i> and <i>Spongicola japonica</i> which bear rudimentary setiferous organs on the first pereiopods. The main function of these setiferous organs in shallow water stenopodideans is for self-grooming (Bauer 1981, 1989). The function of these structures in a deep water hexactinellid sponge commensal is hard to explain. Perhaps in combination with the dense setal brushes on the third pereiopods, these setiferous organs are used to sweep silty deposits off the shrimp’s body as well as the internal sponge chamber in which the shrimp occupies.</p> <p> The bathymetric range of the new species is from 460 to 966 m, which is similar to that of <i>Spongicola japonica</i> at 300– 807 m. Collection of deep water hexactinellid sponges and careful examination of their internal chambers can yield commensals, as evidenced by the capture of four specimens of <i>Spongiocaris neocaledonensis</i> from two specimens of the hexactinellid sponge <i>Regardella okinoseana</i>. The unique holotype of <i>S. yaldwyni</i> was taken from <i>Regardella okiniseana</i> like the new species. Use of deepsea submersibles can uncover new commensal relationships that are normally lost to trawling or dredging sampling methods. The capture of a male <i>S. neocaledonensis</i> on a gorgorian octocoral was accomplished by the use of the deepsea submersible “Cyana” during the CALSUB expedition. This represents the first record of a stenopodidean shrimp in a commensal relationship with an alcyonarian. Continued future use of deepsea submersibles for invertebrate collections may reveal other new and interesting symbiotic associations such as this one.</p>Published as part of <i>Goy, Joseph W., 2015, Stenopodidean shrimps (Crustacea: Decapoda) from New Caledonian waters, pp. 301-344 in Zootaxa 4044 (3)</i> on pages 313-320, DOI: 10.11646/zootaxa.4044.3.1, <a href="http://zenodo.org/record/240460">http://zenodo.org/record/240460</a>
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