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Stenodactylina pseudoventrosa Devillez & Charbonnier 2021, n. comb.
Stenodactylina pseudoventrosa (Beurlen, 1928) n. comb. (Fig. 17A, B) Eryma pseudoventrosa Beurlen, 1928: 158; 1933: 90. — Schweitzer et al. 2010: 24. Eryma modestiformis (pars.) – Förster 1966: 118. TYPE MATERIAL. — Holotype GPIT Ar/294/3. TYPE LOCALITY. — Hülben, Baden-Würrtemberg, Germany. TYPE AGE. — Kimmeridgian. DESCRIPTION Carapace Sub-cylindrical carapace; spiny rostrum; fusiform intercalated plate; narrow post-orbital area; deep and wide cervical groove, strongly inclined dorsally, inflected at carapace mid-height, joined to dorsal margin and to antennal groove; deep and wide antennal groove; short, wide gastro-orbital groove, deep and oblique, originating as a median inflexion of cervical groove; postcervical and branchiocardiac grooves subparallel, slightly curved, very close; deep postcervical groove, not joined to dorsal margin and interrupted in hepatic region; deep branchiocardiac groove, strongly inclined, not joined to dorsal margin. Ornamentation Carapace covered by small tubercles preceded by crescentshaped depressions; cephalic region with an oblique orbital row of tubercles. DISCUSSION Known by a single cast of carapace, this species was originally assigned to Eryma (Beurlen 1928), and then regarded as a synonym of Eryma modestiforme (see Förster 1966). Careful examination of the holotype clearly shows the absence of junction between postcervical and branchiocardiac grooves and the interruption of the postcervical in the hepatic region. This groove pattern is diagnostic of Stenodactylina. So, the new combination Stenodactylina pseudoventrosa, n. comb. is here proposed. The strong proximity of the postcervical and branchiocardiac grooves allows the distinction of S. pseudoventrosa, n. comb. from S. burgundiaca and S. delphinensis. Moreover, the ventral extremity of the postcervical groove of S. pseudoventrosa, n. comb. is lower than in S. burgundiaca, S. guisei, S. shotoverigiganti n. sp., S. triglypta, and S. walkerae. Contrary to S. australis, S. burgundiaca, S. delphinensis, S. deslongchampsi, S. granulifera, S. guisei, and S. triglypta, an orbital row is present in S. pseudoventrosa. There are also depressions and no coarse elements in its ornamentation contrary to S. australis, S. deslongchampsi, S. lagardettei, S. shotoverigiganti n. sp., S. triglypta, and S. walkerae.Published as part of Devillez, Julien & Charbonnier, Sylvain, 2021, Review of the Late Jurassic erymoid lobsters (Crustacea: Decapoda), pp. 25-73 in Geodiversitas 43 (2) on page 61, DOI: 10.5252/geodiversitas2021v43a2, http://zenodo.org/record/448647
Pustulina Quenstedt 1857
Genus Pustulina Quenstedt, 1857 (Fig. 1 H-I) Pustulina Quenstedt, 1857: 807. — Glaessner 1969: 481. — Feldmann et al. 2015: 3. — Devillez et al. 2016: 531, fig. 1K- L. — Devillez et al. 2017: 792. Phlyctisoma Bell, 1863: 34. — Zittel 1885: 695. — Glaessner 1929: 314. — Secrétan 1964: 74. — Förster 1966: 135. — Glaessner 1969: 626. TYPE SPECIES. — Pustulina suevica Quenstedt, 1857, by monotypy. EMENDED DIAGNOSIS BY Devillez et al. (2016). — Fusiform intercalated plate; inflated hepatic, cardiac and branchial regions; deep cervical groove, joined to dorsal margin and to antennal groove; deep, long gastro-orbital groove, originating as a slight median inflexion of the cervical groove, with two divergent, curved branches, delimiting two gastro-orbital lobes; strongly inclined postcervical groove, inflected before joining hepatic groove, not joined to dorsal margin; short and shallow branchiocardiac groove, joined to dorsal margin and not joined to postcervical groove; concave hepatic groove, joined to cervical groove; shallow cardiac groove, straight, inclined forward, rising from postcervical groove, joined to dorsal margin; cephalic region with strongly tuberculate antennal row and distal antennal spine; carapace with tuberculate ornamentation; chelate P1-P3; P1 with strongly tuberculate ornamentation; short P1 propodus with fingers barely longer; P1 dactylus longer than P1 index.Published as part of Devillez, Julien & Charbonnier, Sylvain, 2021, Review of the Late Jurassic erymoid lobsters (Crustacea: Decapoda), pp. 25-73 in Geodiversitas 43 (2) on page 52, DOI: 10.5252/geodiversitas2021v43a2, http://zenodo.org/record/448647
Stenodactylina Beurlen 1928
Genus Stenodactylina Beurlen, 1928 (Fig. 1 J-L) Stenodactylina Beurlen, 1928: 175. — Glaessner 1969: 456. — Schweigert 2013: 411. — Devillez et al. 2016: 522, figs 1D-F. Erymastacus Beurlen, 1928: 171 (pars.). — Secrétan 1964: 71. — Glaessner 1969: 456 (pars.). — Hyžný et al. 2015: 375. TYPE SPECIES. — Stenodactylina liasina Beurlen, 1928, by monotypy. DIAGNOSIS BY Devillez & Charbonnier (2019). — Fusiform intercalated plate; narrow post-orbital area; deep, very wide cervical groove, joined to dorsal margin and to antennal groove; short gastro-orbital groove originating as a slight median inflexion of cervical groove; postcervical and branchiocardiac grooves nearly parallel; narrow postcervical groove, not joined to branchiocardiac groove and interrupted in hepatic region; branchiocardiac groove strongly inclined, joined to hepatic groove; hepatic groove concavo-convex, joined to cervical groove; inferior groove convex posteriorly, joined to hepatic groove; chelate P1; P1 propodus rectangular or trapezoidal, with inner margin more compressed than outer margin, with a wide dactylar bulge; P1 with extremely long and slender fingers, equal in length; P1 chela (form I; Fig. 1K) with strong, rectangular or trapezoidal propodus, bearing straight or sinuous fingers, strongly narrowing immediately after their basis; outer margin convex at the basis of the index; P1 chela (form II; Fig. 1L) with trapezoidal propodus, outer margin straight or convex, straight fingers, narrowing gradually to their distal extremity.Published as part of Devillez, Julien & Charbonnier, Sylvain, 2021, Review of the Late Jurassic erymoid lobsters (Crustacea: Decapoda), pp. 25-73 in Geodiversitas 43 (2) on page 58, DOI: 10.5252/geodiversitas2021v43a2, http://zenodo.org/record/448647
Stenodactylina shotoverigiganti Devillez & Charbonnier 2021, n. sp.
<i>Stenodactylina shotoverigiganti</i> n. sp. <p>(Fig. 16 D-G)</p> <p>urn:lsid:zoobank.org:act: E2B19293-97D9-41F9-AF43-4B03CA6F7E2F</p> <p> <i>Eryma</i> cf. <i>bedelta</i> – Woods 1930: pl. 21, fig. 2.</p> <p>TYPE MATERIAL. — Holotype NHMUK 24559.</p> <p> ETYMOLOGY. — The specific epithet comes from the Latin “ <i>gigantes</i> ”, meaning giant, associated to the name of the type locality, Shotover. It refers to the giant of Shotover, a geoglyph, mentioned during the 17th century by Jean Aubrey, that has disappeared now.</p> <p>TYPE LOCALITY. — Shotover, Oxfordshire, United Kingdom.</p> <p>TYPE AGE. — Oxfordian.</p> <p>DESCRIPTION</p> <p> <i>Carapace</i></p> <p>Sub-cylindrical carapace; fusiform intercalated plate; elongated cardiac region; deep and wide cervical groove, becoming wider ventrally, slightly sinuous at its median inflexion, joined to dorsal margin and to antennal groove; deep and very wide antennal groove; short, shallow gastro-orbital groove, oblique, originating as a slight median inflexion of cervical groove; subparallel postcervical and branchiocardiac grooves, slightly curved; deep and wide postcervical groove, not joined to dorsal margin and interrupted in hepatic region; deep and wide branchiocardiac groove, becoming wider towards its junction to the hepatic groove, joined to dorsal margin, joined to the posterior extremity of the hepatic groove; wide hepatic groove, concavo-convex, joined to the cervical groove; slightly inflated ω area, ventrally delimited by a shallow depression extending between cervical and hepatic grooves; flat χ area; deep and wide inferior groove, joined to hepatic groove.</p> <i>Ornamentation</i> <p>Carapace covered by rounded tubercles; intercalated plate with small tubercles irregularly spaced; gastric region with an oblique row of tubercles ended by an orbital spine.</p> <p>DISCUSSION</p> <p> <i>Stenodactylina shotoverigiganti</i> n. sp. is known by a unique carapace previously mentionned and figured byWoods (1930) as <i>Eryma</i> cf. <i>bedelta</i>. Careful examination of the specimen shows the absence of junction between postcervical and branchiocardiac grooves, and the interruption of the postcervical groove in hepatic region while the branchiocardiac groove is joined to the hepatic groove. This groove pattern is diagnostic of <i>Stenodactylina</i>. Moreover, some morphological features allow the distinction of this specimen from all other representatives of the genus for which the carapace is known. Indeed, the postcervical groove is shorter than that of <i>S. australis</i>, <i>S. delphinensis</i> (Moret, 1946), <i>S. deslongchampsi</i> (Van Straelen, 1925), <i>S. granulifera</i>, <i>S. lagardettei</i>, <i>S. pseudoventrosa</i>, and <i>S. walkerae</i> (Feldmann & Haggart, 2007). It is also slightly curved contrary to <i>S. burgundiaca</i> (Crônier & Courville, 2004), <i>S. delphinensis</i>, <i>S. granulif-era</i>, <i>S. triglypta</i>, and <i>S. walkerae</i>. Only the ω area is inflated while it is flat in some species (<i>S. australis</i>, <i>S. delphinensis</i>, <i>S. deslongchampsi</i>) or both ω and χ areas are inflated in some others (<i>S. burgundiaca</i>, <i>S. granulifera</i>, <i>S. guisei</i> (Wright, 1881), <i>S. lagardettei</i>). The ornamentation of the specimen is thin and only made of tubercles contrary to <i>S. australis</i>, <i>S. burgundiaca</i>, <i>S. delphinensis</i>, <i>S. granulifera</i>, <i>S. guisei</i>, <i>S. lagardettei</i>, and <i>S. pseudoventrosa</i>. Finally, there is an orbital row, which is absent in <i>S. australis</i>, <i>S. burgundiaca</i>, <i>S. delphinensis</i>, <i>S. deslongchampsi</i>, <i>S. guisei</i>, and <i>S. triglypta</i>. Considering all these elements, we consider this specimen as a representative of a new species of erymid lobsters: <i>Stenodactylina shotoverigiganti</i> n. sp.</p>Published as part of <i>Devillez, Julien & Charbonnier, Sylvain, 2021, Review of the Late Jurassic erymoid lobsters (Crustacea: Decapoda), pp. 25-73 in Geodiversitas 43 (2)</i> on page 60, DOI: 10.5252/geodiversitas2021v43a2, <a href="http://zenodo.org/record/4486479">http://zenodo.org/record/4486479</a>
Stenodactylina strambergensis
Stenodactylina strambergensis (Bachmayer, 1959) (Fig. 17 G-I) Erymastacus strambergensis Bachmayer, 1959: 940, pl. 2, fig. 2. Phlyctisoma strambergensis – Förster 1966: 144, pl. 18, fig. 8. Eryma strambergensis – Schweitzer et al. 2010: 24. Stenodactylina strambergensis – Devillez et al. 2016: 524. TYPE MATERIAL. — Holotype NMW 344/1959. TYPE LOCALITY. — Stramberg, Moravian-Silesian, Czech Republic. TYPE AGE. — Tithonian. DESCRIPTION Thoracic appendages Chelate P1; P1 propodus sub-rectangular, slightly globose dorso-ventrally; inner margin strongly compressed; rounded outer margin with a slight deviation at the basis of the index; slender index; wide and inflated dactylar bulge. Ornamentation P1 propodus covered by rounded, coarse and widely spaced tubercles; inner margin with a row of spines. DISCUSSION This species is known by some P1 propodi. Initially included within Erymastacus, then moved into Phlyctisoma (see Förster 1966) and Eryma (see Schweitzer et al. 2010), and finally to Stenodactylina by Devillez et al. (2016). This assignment is supported by the stronger compression of the inner margin than that of outer margin, the presence of a row of coarse tubercles on inner margin (present in most of the species of the genus), the strongly inflated dactylar bulge, and the fine basis of the index. Because only the P1 propodus of this species is known, the comparisons with other species of Stenodactylina are limited. Stenodactylina strambergensis has the particularity to have a relatively short propodus. Moreover, the ornamentation of S. strambergensis is homogeneous – only made of coarse tubercles – contrary to other species, except S. armata and S. triglypta.Published as part of Devillez, Julien & Charbonnier, Sylvain, 2021, Review of the Late Jurassic erymoid lobsters (Crustacea: Decapoda), pp. 25-73 in Geodiversitas 43 (2) on page 62, DOI: 10.5252/geodiversitas2021v43a2, http://zenodo.org/record/448647
FIG. 1 in The decapod crustacean fauna from the Late Jurassic of Cricqueboeuf, Normandy (France)
FIG. 1. — Geographic and geological settings: A, location of the Cricqueboeuf outcrop in Normandy;B, stratigraphic log of the upper Oxfordian- lower Kimmeridgian succession based on Guyader (1968) and modified after Devillez et al. (2018). The red arrow indicates the interval of beds yielding crustacean-bearing nodules.Published as part of <i>Charbonnier, Sylvain, Garassino, Alessandro, Gendry, Damien, Devillez, Julien & Picot, Laurent, 2023, The decapod crustacean fauna from the Late Jurassic of Cricqueboeuf, Normandy (France), pp. 573-588 in Geodiversitas 45 (19)</i> on page 575, DOI: 10.5252/geodiversitas2023v45a19, <a href="http://zenodo.org/record/10066067">http://zenodo.org/record/10066067</a>
FIG. 5 in Review of the Late Jurassic erymoid lobsters (Crustacea: Decapoda)
FIG. 5. — Additionnal synonyms of Eryma ventrosum (Meyer, 1835): A, B, holotype BSPG AS VIII 114 of Eryma radiatum Oppel, 1861 (Oxfordian, Balingen, Germany): carapace (A), P1 chela (B); C, holotype IRSNB of Eryma cumonti Van Straelen, 1921 (Callovian, Pougues-les-Eaux, France); D, holotype MNHN.F.B13231 of Eryma babeaui Étallon, 1861 (Kimmeridgian, Boulogne-sur-mer, France); E, syntype MNHN.F.A29782 of Bolina thirriae Étallon, 1859 (Kimmeridgian, Gray, France); F, syntype MNHN.F.A29727 of Eryma affinis Ferry, 1865 (Bathonian, Fuissé, France); G, original figure of Garassino & Krobicki (2002: fig. 7) of the holotype of Galicia marianae (Oxfordian, Rudno, Poland); H, syntype MNHN.F.A29783 of Bolina girodi Étallon, 1859 (Saint-Claude, France); I, lectotype MJSN Col.Del.475 of Eryma greppini (Oppel, 1861) (Vellerat, Switzerland); J, paralectotype MJSN Col.Del.1 of E. greppini (Vellerat, Switzerland); K, dorsal view of the syntype MNHN.F.A29783 of B. girodi; L, dorsal view of the lectotype MJSN Col.Del.475 of E. greppini; M, specimen MFN 2236 P1383/2 MB.A.1537 from the Late Jurassic of Tanzania. Abbreviation: dd, dorsal domes. Photographs: A-C, M, J. Devillez; D-F, P. Loubry; H, K, L. Cazes; I-J, L, D. Becker. Scale bars: 1 cm.Published as part of Devillez, Julien & Charbonnier, Sylvain, 2021, Review of the Late Jurassic erymoid lobsters (Crustacea: Decapoda), pp. 25-73 in Geodiversitas 43 (2) on page 34, DOI: 10.5252/geodiversitas2021v43a2, http://zenodo.org/record/448647
FIG. 10 in A synthesis of the evolutionary history of erymoid lobsters (Crustacea, Decapoda, Erymoidea)
FIG. 10. — Erymoid fauna from the Kimmeridgian-Tithonian "Plattenkalk" of Bavaria (Germany): A, specimen MFN 2236 P1383/3 MB.A.2891 of Eryma modestiforme (Schlotheim, 1822) from Solnhofen; B, holotype SMNS 3682 of Eryma major Oppel, 1861, from Nusplingen; C, holotype SMNS 24227 of Eryma westphali Schweigert, Dietl & RÖper, 2000, from Nusplingen; D, holotype BSPG AS VII 186 of Eryma veltheimii (Münster, 1839), from Kehlheim; E, specimen SMNS 64681 of Eryma punctatum Oppel, 1861, from Nusplingen; F, specimen SMNS 64521 of Palaeastacus fuciformis (Schlotheim, 1822), from Zandt; G, holotype BSPG 1993 XXVIII 200 of Palaeastacus rothgaengerae Schweigert & RÖper, 2001, from Brunn; H, holotype SMNS 70507 of Stenodactylina geigerae Schweigert & Härer, 2020, from Marxheim; I, holotype SMNS 64872 of Stenodactylina devillezi Schweigert & Härer, 2020, from Nusplingen; J, specimen SMNS 64319 of Pustulina suevica Quenstedt, 1857, from Nusplingen; K, specimen BSPG AS I 619 of Pustulina minuta (Schlotheim, 1822), from Solnhofen. Scale bars: 1 cm. Photographs: J. Devillez (A-F, J-K), G. Schweigert (G-I).Published as part of Devillez, Julien & Charbonnier, Sylvain, 2022, A synthesis of the evolutionary history of erymoid lobsters (Crustacea, Decapoda, Erymoidea), pp. 265-289 in Geodiversitas 44 (9) on page 278, DOI: 10.5252/geodiversitas2022v44a9, http://zenodo.org/record/633564
Enoploclytia Mesozoique, M'Coy 1849
Genus Enoploclytia M’Coy, 1849 (Fig. 1 M-N) Enoploclytia M’Coy, 1849: 330; 1854: 137. — Zittel 1885: 694. — Fritsch & Kafka 1887: 27. — Van Straelen 1925: 278. — Beurlen 1928: 164. — Rathbun 1926: 128. — Secrétan 1964: 81. — Förster 1966: 146. — Taylor 1979: 25. — Aguirre-Urreta 1989: 514. — Feldmann et al. 2015: 3. — Devillez et al. 2016: 530, fig. 1I-J; 2017: 786, fig. 4A-B; 2018: 144, fig. 2A-B. Enoploclytia Enoploclytia – Mertin 1941: 160. — Glaessner 1969: 455. TYPE SPECIES. — Astacus leachii Mantell, 1822, by original designation. DIAGNOSIS BY Devillez et al. (2018). — Fusiform intercalated plate; wide, deep cervical groove, joined to dorsal margin and to antennal groove; long, wide gastro-orbital groove originating as a median inflexion of cervical groove, delimiting two gastroorbital lobes; sinuous postcervical groove, joined to dorsal margin and to hepatic groove, with ventral extension at carapace mid-height; short branchio-cardiac groove, interrupted in upper part of carapace, joined to dorsal margin, not joined to postcervical groove; concavo-convex hepatic groove, joined to cervical groove; prominent ω and χ bulges; inferior groove convex posteriorly, joined to hepatic groove; carapace with heterogeneous coarse ornamentation; massive globose P1 propodus, rounded in transversal section; long and thin P1 fingers (straight in dorsal view); occlusal margins armed with sharp and slender tooth; P1 merus with strong, prominent distal process at extern side of its ventral extremity. DISCUSSION Recently, Devillez et al. (2018) reported a specimen identified as Enoploclytia sp. from the Oxfordian of France (Fig. 18). Despite its poor preservation, this fossil exhibits the typical carapace groove pattern of the genus: elongated gastro-orbital groove, with two divergent distal branches, a sinuous postcervical groove joined to the posterior extremity of the hepatic groove and not joined to the branchiocardiac groove, which is short and interrupted in the branchial region (Fig. 18C). This specimen is the oldest occurrence of Enoploclytia, and the only known in the Jurassic. Indeed, Enoploclytia is more typical of the Cretaceous.Published as part of Devillez, Julien & Charbonnier, Sylvain, 2021, Review of the Late Jurassic erymoid lobsters (Crustacea: Decapoda), pp. 25-73 in Geodiversitas 43 (2) on page 64, DOI: 10.5252/geodiversitas2021v43a2, http://zenodo.org/record/448647
Palaeastacus Bell 1850
Genus <i>Palaeastacus</i> Bell, 1850 <p>(Fig. 1 F-G)</p> <p> <i>Palaeastacus</i> Bell, 1850: 344. <i>—</i> Zittel 1885: 695. <i>—</i> Beurlen 1928: 180. <i>—</i> Förster 1966: 126. <i>—</i> Glaessner 1969: 626. <i>—</i> Aguirre-Urreta & Ramos 1981: 606. <i>—</i> Aguirre-Urreta 1989: 509. <i>—</i> Schweitzer & Feldmann 2001: 174. <i>—</i> Feldmann <i>et al.</i> 2015: 3. <i>—</i> Hyžný <i>et al.</i> 2015: 375. <i>—</i> Devillez <i>et al.</i> 2016: 525, fig. 1G-H. <i>—</i> Devillez <i>et al.</i> 2017: 782.</p> <p> <i>Enoploclytia Palaeastacus</i> – Mertin 1941: 161. <i>—</i> Glaessner 1969: 455.</p> <p> TYPE SPECIES. <i>—</i> <i>Astacus sussexiensis</i> Mantell, 1824, by subsequent designation of Glaessner (1929).</p> <p>DIAGNOSIS BY Devillez & Charbonnier (2019). — Fusiform intercalated plate; deep cervical groove, joined to dorsal margin and to antennal groove; short gastro-orbital groove, originating as a slight median inflexion of the cervical groove; postcervical and branchiocardiac grooves subparallel, joined to hepatic groove; hepatic groove concavo-convex, joined to cervical groove; inferior groove convex posteriorly, joined to hepatic groove; chelate P1; P1 propodus short, thick, slightly globose, with a narrow dactylar bulge; P1 fingers usually wide, slightly longer than propodus, progressively narrowing to their distal extremity, occlusal margin curved at the basis of the index.</p>Published as part of <i>Devillez, Julien & Charbonnier, Sylvain, 2021, Review of the Late Jurassic erymoid lobsters (Crustacea: Decapoda), pp. 25-73 in Geodiversitas 43 (2)</i> on page 49, DOI: 10.5252/geodiversitas2021v43a2, <a href="http://zenodo.org/record/4486479">http://zenodo.org/record/4486479</a>
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