1,723,147 research outputs found
Tarjeta Postal de Joaquín Almeda a Pedro Dorado Montero
No full textTarjeta Postal de D. Joaquín Almeda, jurista catalán, a D. Pedro Dorado Montero, remitiéndole su trabajo "La costumbre como fuente de derecho"
The ancestry and the journal of Almeda Widger Hitchcock
No full textA history of the Widger family / Mrs. Caroline Gardner -- Notes on the Angel family / by Lucy Widger Alton -- Notes on the Atwell family / by Caroline Widger Gardner -- Early life of Almeda Widger (Hitchcock) / C.A. Gardner -- A copy of the journal and letters of Almeda Widger Hitchcock / copied by Caroline Widger Gardner.Unpublished bound typescript left to Almeda Townsend Goss by her mother Cora Hitchcock Townsend. Compiled and typed by Cora Hitchcock Townsend sometime after 1922 from notes hand-written by Caroline Widger Gardner between 1918 and 1922, letters hand-written by Almeda Widger Hitchcock in 1857, and journal entries hand-written by Almeda Widger Hitchcock 1856-1857, hand-copied by Caroline Widger Gardner in 1922, and mailed to Cora Hitchcock Townsend by Caroline Widger Gardner in 1922. Journal entries dated November 23, 1856-March 12, 1857; letters dated March 20, 1857-August 3, 1857
Dewey and Almeda Stewart Papers, MSS.2158
No full textAbstract: This collection consists mainly of correspondence between a young married couple during World War Two. It also includes various miscellaneous World War Two-era items and service-related materials.Scope and Content Note: Almost all of this collection is personal correspondence between Dewey and Almeda Stewart, a husband and wife separated by his service in the Navy during World War Two. The couple had only been married eight months when he departed for the Pacific Islands. Therefore, much of the content of the letters is of a very personal nature- detailing the longing and loneliness of a young couple separated by war and the excitement of planning a happy future together. Dewey was away for just under one year yet he wrote her almost every day, sometimes twice a day.Dewey makes several references that may be of special interest to historians. In a letter dated April 13, 1945, he expresses sadness over President Roosevelt's death saying that the country had experienced the "loss of a great man-"describing him as "intensely patriotic." In a letter dated August 7, 1945, a reference is made to a newsreel shown to servicemen about the "new bomb" that will bring the war "to a screeching halt." In a letter dated August 11, 1945, Dewey says that he and his fellow servicemen received "unofficial news" from the Navy that Japan had surrendered. On August 14, 1945, he describes a huge celebration held on the base. *August 15, 1945 is the "official" date historically recorded for Japan's surrender.*Many of the letters had portions cut out by the war censor, which was officially lifted on September 4, 1945. Dewey' subsequent letters are significantly more personal and provide more details about the Navy, what he is doing, and where he is.Dewey writes on various service letterhead that may be of special interest including: U.S. Naval Air Station; USO; U.S. Navy; U.S. Naval Receiving Station; Airmail; and American Red Cross.The collection also includes the last four letters Almeda wrote to Dewey before he came home. They discuss her happiness that the war is over and plans for their reunion. Additionally, there is one letter to Dewey Stewart from Roger H. Biddle, a Navy buddy who is stationed on the U.S.S. Moon near Okinawa. Letters from two of Almeda Stewart's brothers (both enlisted in the Army) are enclosed- the first is from Melvin Wagner, who is stationed in Japan; the second is from Raymond Wagner, who is stationed in the Panama Canal Zone.The collection contains several scenic "Pictorial Wonderland" postcards; telegraphs; pamphlets distributed by the U.S. Navy; several "Orders of Worship" for church services "at sea;" and various holiday cards from 1944-1945.Biographical/Historical Note: Dewey and Almeda Stewart were married on June 10, 1944 in Hernando, Mississippi. Dewey was an aviation metal smith, 1st class, in the U.S. Navy. He met Almeda while training at the U.S. Naval Air Station in Memphis, Tennessee. After marrying, the couple lived in Memphis until Dewey completed his training at the end of 1944. In February 1945, Dewey was sent to the Naval Receiving Station in Shoemaker, California. There he had two surgeries, one for a burst appendix and one to remove his tonsils. He shipped out on the U.S.S.
Audubon on June 1, 1945. The ship arrived at Pearl Harbor on June 7, 1945, staying in port two days for supplies before departing for the Pacific Islands. Almost a month later, on July 2, Dewey and the
Audubon docked at Samar Island in the Philippines. He worked in the mailroom and serviced ships and airplanes while serving on Samar. Dewey would remain here until the end of the war, obtaining transport home at the end of September 1945. He never saw any combat action during his brief Navy service during World War Two.While Dewey served in the Pacific, Almeda stayed with various relatives. First she returns to her parents' home in Dulaplaine, Arkansas. After a few months, she moved to Union Springs, Alabama, to live with Dewey's mother. She found a job working in a five and dime store and remained there for the duration of Dewey's service.Almeda Stewart had three brothers--Harrison Wagner, Melvin Wagner, and Raymond Wagner--all of whom served overseas during World War Two
Miconia veraguensis Gamba & Almeda 2014, spec. nov.
No full text33. Miconia veraguensis Gamba & Almeda, spec. nov. (Fig. 26) Related to M. approximata by virtue of the densely fasciculate glomerules. Distinguished by the elliptic ovate berries which are larger than in its closest relatives; 6.59–7.34 × 4.31–5.3 mm. Type: PANAMA. Prov. Veraguas: Trail to Reserva Biológica Serranía de Tute and the summit of Cerro Tute about 0.7 km beyond the Escuela Agrícola Río Piedra just outside Santa Fe, 860–1300 m, 18 February 1996, Almeda et al. 7620 (holotype: CAS!; isotype MO!, NY!, PMA!). Little-branched shrub 1–1.5 m tall, bark green-brown. Upper internodes rounded-quadrate 1.09–1.91 cm long, cauline nodes slightly compressed becoming terete with age, nodal line present. Indumentum on branchlets, petioles, adaxial leaf surface, primary, secondary and tertiary veins adaxially and abaxially, bracts, bracteoles, pedicels, hypanthia, calyx lobes and calyx teeth densely covered with caducous white-translucent elongate slightly roughened trichomes 1–1.5 mm long, each trichome deflexed and somewhat flattened, intermixed with a dense understory of dendritic trichomes 0.2–0.5 mm long with moderately long thin-walled arms. Leaves of each pair slightly anisophyllous in size; subsessile to short-petiolate, the free rounded-quadrate petioles 0.42–0.95 cm long (on larger leaves) or 0.21–0.4 cm long (on smaller leaves), widely canaliculate adaxially, convexly 3-grooved abaxially, succulent, brownish; larger blades 12.5–20.5 × 6–9 cm, elliptic-obovate, the base acute or roundedcordate, shortly decurrent on the petiole, the margin crenulate to subentire, the apex bluntly apiculate; smaller blades 6.5–14 × 4.75–8.7 cm, elliptic-obovate to obovate, the base slightly rounded to attenuate, shortly decurrent on the petiole, the margin crenulate to subentire, the apex bluntly apiculate; chartaceous; adaxial surface of mature leaves, primary, secondary and tertiary veins glabrescent, the elongate roughened trichomes denser toward the base, the higher order veins glabrous; abaxial surface superficially glabrous, microscopically papillose with resinous unfurrowed or slightly furrowed glands to 0.1 mm in diameter, the indumentum on the secondary veins intermixed with a resinous understory of minute sessile to short-stalked glands 0.1 mm long with thin-walled short heads, these glands also present on the tertiary and higher order veins, sparsely intermixed with white furrowed sessile glands ca. 0.1 mm long; 5–(7-) plinerved, including the tenuous marginals, innermost pair of secondary veins diverging symmetrically from the primary vein 0.5–3 cm above the base, areolae 0.3–0.4 mm, adaxially the primary, secondary and tertiary veins deeply impressed, the higher order veins slightly so, abaxially the primary and secondary veins elevated and terete, somewhat succulent, the tertiary and higher order veins slightly raised to flat. Inflorescences a congested, axillary and fasciculate many-bracted glomerule 1.22–2.18 cm long, sessile, SYSTEMATICS OF THE OCTOPLEURA CLADE OF MICONIA Phytotaxa 179 (1) © 2014 Magnolia Press 139 unbranched, paired or appearing verticillate in the upper leaf axils and at defoliated nodes; bracts 5.43–7.04 × 2.75–3.95 mm, elliptic to elliptic-ovate, concave, the apex acute, greenish, glabrescent, persistent in fruit. Flowers not seen, probably 4-merous based on persistent calyx lobes in fruit, sessile. Hypanthia in fruit 6.4–6.85 × 1.5–2 mm, free portion of hypanthium 1.6–1.8 mm long, subcylindric to urceolate, bluntly 8-ribbed, green, the indumentum mostly consisting of dendritic trichomes to 0.3 mm long, intermixed with minute sessile glands and with white furrowed sessile glands, both ca. 0.1 mm long, ridged on the inner surface, glabrous, the torus adaxially glabrous, somewhat glossy. Calyx persistent in fruit, green to brown; tube 0.3–0.5 mm long, glabrous adaxially, with the same vestiture as the hypanthium abaxially; lobes 2–2.5 × 1.5–1.8 mm, triangular, slightly concave, the margin entire, the apex bluntly acute, the indumentum intermixed with the same two types of glands present on the hypanthium, spreading to reflexed in fruit; exterior calyx teeth to 1.8 mm long, linear-deltoid, thick, inserted half way up the lobes and projecting beyond them. Ovary (in fruit) 4-locular, completely inferior, 4.8–5 mm long, the apical collar 1 × 0.8–1 mm, conic, glandular-puberulent. Berries 6.59–7.34 × 4.31–5.3 mm when dry, globoseelliptic to globose-obovate, light green, ripening orange, the hypanthial indumentum persistent at maturity. Seeds 0.57–0.75 × 0.39–0.44 mm, pyramidal, yellow-brown; lateral symmetrical plane triangular, the highest point near the central part of the seed, with a foot-like projection at the micropylar end; antiraphal symmetrical plane suboblong; raphal zone suboblong, ca. 80% the length of the seed; multicellular sculpture rugose throughout the seed. Individual cells elongate and isodiametric, the latter found at the highest point of the seed, anticlinal boundaries channeled, undulate, with Ω- and U-type patterns; periclinal walls convex, low- to high-domed, microrelief striate. Additional specimens studied:— PANAMA. Veraguas: Along trail to summit of Cerro Tute about 1/ 2 mile above the Escuela Agrícola Alto Piedra near Santa Fe, 8.48222°N, - 1.09805°W, 900–1100 m, 29 January 1989, Almedaet al. 6480 (CAS, MO, NY, PMA). Illustration:— Fig. 26. Common names and documented uses:— None recorded. Habitat, distribution and ecology:— A local and uncommon species known only from cloud forests of Cerro Tute in the province of Veraguas, Panama (Fig. 16), at 860–1300 m. Phenology:— Collected in fruit in January and February. Etymology:— The specific epithet refers to the province of Veraguas in Panama, where this species appears to be endemic. Discussion:— This species has a distinctive white-translucent lanate vegetative indumentum, flowers that are congested in fasciculate glomerules with conspicuous bracts, and large bright orange mature berries. In its poorly developed inflorescences and rugose seeds, M. veraguensis is most similar, and also most closely related, to those species in the Approximata subclade that have sessile fasciculate glomerules. Miconia approximata which occurs nearly throughout Central America south to Ecuador has a thicker and darker vestiture, shorter, globose-oblate fruits at maturity (2–2.5 × 5–6 mm vs. 6.59–7.34 × 4.31–5.3 mm that is globose-elliptic to globose-obovate). In Veraguas province, M. approximata is only known from Isla de Coiba. Miconia veraguensis is also similar to M. chocoensis and M. quadridomius, two South American species that have a longer lanate indumentum (1.5–3 mm vs. 1–1.5 mm) and smaller berries. Although flowers of this species remain unknown, it is clearly distinct from its close relatives in the characters mentioned above. Conservation status:— This species would be considered Critically endangered CR Dbased on IUCN criteria. Because this species is rare and local in a protected area of the Cerro Tute in Veraguas, Panama, a status of Vulnerable VU is warranted. 140 Phytotaxa 179 (1) © 2014 Magnolia Press GAMBA & ALMEDA SYSTEMATICS OF THE OCTOPLEURA CLADE OF MICONIA Phytotaxa 179 (1) © 2014 Magnolia Press 141 142 Phytotaxa 179 (1) © 2014 Magnolia Press GAMBA & ALMEDA SYSTEMATICS OF THE OCTOPLEURA CLADE OF MICONIA Phytotaxa 179 (1) © 2014 Magnolia Press 143 144 Phytotaxa 179 (1) © 2014 Magnolia Press GAMBA & ALMEDA SYSTEMATICS OF THE OCTOPLEURA CLADE OF MICONIA Phytotaxa 179 (1) © 2014 Magnolia Press 145 146 Phytotaxa 179 (1) © 2014 Magnolia Press GAMBA & ALMEDA SYSTEMATICS OF THE OCTOPLEURA CLADE OF MICONIA Phytotaxa 179 (1) © 2014 Magnolia Press 147 148 Phytotaxa 179 (1) © 2014 Magnolia Press GAMBA & ALMEDA SYSTEMATICS OF THE OCTOPLEURA CLADE OF MICONIA Phytotaxa 179 (1) © 2014 Magnolia Press 149 Excluded species Clidemia radicans Pilger (1905: 179). Type: PERÚ. Amazonia: close to Leticia, Ule 6869 (holotype: MG; isotype: B-internet image!, photograph: F!). = Clidemia epiphytica var. trichocalyx (Blake) Wurdack (1964: 215–216). Ossaea ciliata (Triana) Cogniaux (1891a: 1067). Davya ciliata Grisebach (1860b: 265). Octopleura ciliata Triana (1871: 146). Type: In insula TRINITATIS, Crueger s.n. (holotype: BR-internet image!). = Miconia lateriflora Cogniaux (1909: 255). Ossaea involucrata (Grisebach) Triana (1871: 147). Type: CUBA. Prope Monte Verde, 1856–1857, Wright 194 (holotype: BR- 2 sheets-internet images!). = Calycogonium involucratum Grisebach (1860c: 184).Published as part of Gamba, Diana & Almeda, Frank, 2014, Systematics of the Octopleura Clade of Miconia (Melastomataceae: Miconieae) in Tropical America, pp. 1-174 in Phytotaxa 179 (1) on pages 139-150, DOI: 10.11646/phytotaxa.179.1.
Fig. 1. – Dichaetanthera schatzii H. Ranariv. & Almeda. A in A new Dichaetanthera (Melastomataceae: Melastomateae) from Masoala National Park in Madagascar
No full textFig. 1. – Dichaetanthera schatzii H. Ranariv. & Almeda. A. Habit; B. Enlargement of inflorescence node; C. Representative leaf (abaxial surface); D. Detail of indumentum on elevated primary vein; E. Petal (adaxial surface); F–G. Antepetalous (small) stamens (profile view); H–I. Antesepalous (large) stamens (profile view); J. Hypanthium and calyx lobes enveloping capsule (profile view); K. Ovary, style and stigma (profile view). [Schatz et al. 2806, CAS] [Drawings: A. Chou]Published as part of Ranarivelo, Heritiana & Almeda, Frank, 2019, A new Dichaetanthera (Melastomataceae: Melastomateae) from Masoala National Park in Madagascar, pp. 131-136 in Candollea 74 (2) on page 133, DOI: 10.15553/c2019v742a2, http://zenodo.org/record/572470
Miconia amplipedunculata Almeda & O. Ortiz 2022, sp. nov.
No full textMiconia amplipedunculata Almeda & O. Ortíz, sp. nov. (Figs. 1 & 2) Diagnosis:— Miconia amplipedunculata can be recognized by its shortly petiolate lanceolate to oblanceolate leaf blades that are alternately 3-plinerved, brown amorphous stellulate-lepidote indumentum on the primary and secondary veins of abaxial foliar surfaces and inflorescence, long-pedunculate (to 12.5 cm) inflorescence, persistent bracts and bracteoles, unlobed calyx that is truncate and flangelike, prominent subulate calyx teeth that exceed the calyx in length, 5-merous flowers, distally constricted anther filaments, and isometric stamens that lack appendages. Type:— PANAMA. Colón: Distrito Donoso, Valle Grande, antiguas oficinas MPSA, Quebrada arriaba nacia Patio Limosa, 08º4934.1’ N, 80º40’15.9’ W, 279 m, 4 November 2015, fl., I. Vergara-Pérez 1050 & Rojas (holotype PMA [127052]!; isotype MO [6718180]!). Shrub to 2 m tall, cauline nodes of uppermost branchlets somewhat swollen, the internodes rounded, glabrous and vaguely ridged when dry. Leaves of a pair equal to slightly unequal in size, concolored when dry, petioles 0.4–1 cm long, canaliculated adaxially; blades 16–25 × 4–6 cm, subcoriaceous when dry, lanceolate to oblanceolate, apex acuminate, base rounded to obtuse, margin entire, 3-plinerved with an elevated midvein and one pair of secondaries mostly alternately diverging from the midvein 2.5–8 cm above the blade base, the tertiary veins conspicuous, elevated and spaced 5–8 mm apart at the wider portions of the abaxial surface of the blade, adaxially glabrous at maturity, abaxially essentially glabrous or glabrate on the actual surface but moderately to copiously beset with a brown stellulate-lepidote indumentum on primary and secondary veins. Inflorescence terminal, thyrsoid, 15 cm long with distal branching, the elongate typically widely spreading peduncle 12.5 cm long, copiously stelluate-lepidote; bracts of the rachis nodes 1.5–3 × 0.5 mm, sessile, persistent, lanceolate to elliptic, glabrous adaxially, abaxially moderately and inconspicuously brown amorphous lepidote; bracteoles 0.75–1 × 0.5 mm, sessile, persistent, ovate-lanceolate, adaxially glabrous and abaxially with an indumentum like the bracts. Flowers 5-merous, perfect, pedicels 1–1.5 mm long, hypanthia campanulate, 1.5 × 1.5 mm (excluding tube and flangelike calyx), copiously to moderately brown stelluate-lepidote, the calyx unlobed, reduced to a truncate flangelike rim ca. 0.25 mm long; exterior calyx teeth 5, erect, bluntly subulate, 0.5 mm long and exceeding the truncate calyx; torus minutely glandular-puberulent adaxially. Petals 3.5–4 × 0.75–1 mm, pink to pink-magenta, glabrous, elliptic-oblong, apically obtuse with a minute apical mucro. Stamens 10, isometric and isomorphic, forming a ring around the style at anthesis, filaments 1.5 mm long, white, glabrous, complanate, constricted distally just below the anther thecae; anthers 1 mm long and ca. 0.25 mm wide, pale yellow, linear oblong, the apex with a ± truncate or somewhat dorsally inclined terminal pore; connective slightly thickened but unappendaged. Ovary (at anthesis) ca. 3/5 inferior, 5-locular, ± globose, apex conspicuously fluted, densely white-papillate. Style straight or somewhat declinate, glabrous, 3.5–4 mm long, white; stigma capitellate. Mature berries and seeds not seen. Distribution, habitat, and phenology:— Miconia amplipedunculata is presently known only from low elevations (below 300 m) in the Donoso district of coastal Colón province on the Caribbean slope of Panama (Fig. 3). At its only known locality, where it is evidently uncommon, M. amplipedunculata grows in or at the margins of mature secondary rainforest. The type and only known collection was flowering in early November. Conservation:— The Caribbean slope rainforest habitat where Miconia amplipedunculata grows is not part of a protected area. Because the type was the only collection available for this study we were unable to calculate area of occupancy and extent of occurrence for a recommended conservation assessment using IUCN guidelines and criteria (IUCN 2019). The type locality appears to be within the Concesión del Proyecto Mina de Cobre, a large-scale open-pit copper mine development in Panama. The concession consists of four zones amounting to 13,600 hectares (https://www.first-quantum.com). We have no information on the exact occurrence of M. amplipedunculata within this concession. Until the distribution of this species is better known we regard its conservation status as Data Deficient (DD). Etymology:— The epithet for this species, amplipedunculata, is derived from the Latin word “amplus” (ample, large, abundant, or great). It highlights the unusually long inflorescence peduncle of this species which readily distinguishes it from all allied species. Affinities:— Miconia amplipedunculata is morphologically most similar to a group of species that includes M. iteophylla Almeda (1989: 214), M. jefensis Almeda (2000: 43), M. ligulata Almeda 1989: 216), and M. peltata Almeda (1989: 217). Like M. amplipedunculata, most of these species are restricted to Panama; only M. ligulata has a range that extends from Nicaragua to Colombia and Venezuela (Almeda 1989, 2009; Almeda et al. 2016). All of these species share a brown amorphous stellulate-lepidote indumentum, oblong petals, unappendaged anther connectives, filaments that are constricted distally, a torus that is puberulent adaxially, persistent bracts and bracteoles, and 5-locular ovaries. Miconia peltata can be distinguished from all other species in this complex by its peltate, 5–7-plinerved leaves that are broadly rounded to subcordate at the base, an inflorescence that is divaricately branched at the initiating node, five welldefined triangular calyx lobes, and higher elevation (850–1000 m). Our new species is probably closest to M. jefensis which has wider (7–14 cm) leaf blades that are 5–7-nerved (vs. blades 4–6 cm and 3-plinerved) and well-defined rounded-triangular calyx lobes (vs. an unlobed truncate calyx), and grows at higher elevations (700–1000 m vs. 279 m). Miconia iteophylla also has 3-plinerved leaves with the inner pair of secondary veins diverging from the median nerve in opposite or subalternate fashion like M. amplipedunculata but its leaves are narrowly elliptic and smaller (4.5–9 × 0.6–1.7 cm), its inflorescence is a terminal panicle that branches 0.6–3 cm above the node initiating the inflorescence, its calyx consists of five depressed triangular undulations, its petals are white, and it grows at somewhat higher elevations (200–600 m). Miconia amplipedunculata is only superficially similar to M. ligulata, the most widespread species among this group of congeners; the latter grows in habitats spanning the elevational range of all closely related species in this group (0–1100 m). Miconia ligulata has leaf blades of comparable size to M. amplipedunculata but they are 5-plinerved, attenuate to long-acuminate apically with a base that is gradually tapering and decurrent on the petiole, its inflorescence branches at the initiating node, its calyx consists of five depressed triangular lobes, and its petals are white. All of the close relatives of M. amplipedunculata were included by Kriebel (2016) in his enlarged concept of the genus Conostegia D. Don (1823: 284, 316). A cladogram of a molecular phylogenetic hypothesis in Kriebel’s monograph (Fig. 1) shows three of the four sampled species (M. jefensis, M. ligulata, and M. peltata) grouping with Clidemia trichosantha Almeda (1984: 274) and Miconia brenesii Standley (1938: 816) in his sect. Geniculatae. We here interpret all of these species to be part of a greatly expanded Miconia (Michelangeli et al. 2022).Published as part of Almeda, Frank & Ortíz, Orlando O., 2022, Miconia amplipedunculata (Melastomataceae: Miconieae), a new species from the Caribbean lowlands of Panama, pp. 294-300 in Phytotaxa 575 (3) on pages 295-298, DOI: 10.11646/phytotaxa.575.3.7, http://zenodo.org/record/743133
Miconia neocoronata Gamba & Almeda 2014, nom. nov.
No full text21. Miconia neocoronata Gamba & Almeda, nom. nov. Basionym: Clidemia coronata Gleason (1939a: 114–115). Type: COSTA RICA. Bords du Río Tuis, July 1893, Pittier 8080 (holotype: US-internet image!; isotypes: BR-3 sheets-internet images!). Nec Miconia coronata (Bonpland) de Candolle (1828: 187). Small shrub, rarely epiphytic, (0.5–)1–2(–2.75) m tall, loosely and irregularly branched. Upper internodes [1.75–4.25 cm long] and cauline nodes terete, nodal line absent. Indumentum on branchlets, petioles, primary, secondary, tertiary leaf veins abaxially, inflorescence axes, bracts, bracteoles, and pedicels (when present) densely to copiously composed of brownish sessile or thinly stipitate dendritic trichomes 0.2–0. 5(–1.9) mm long with short axes and few-moderate number of terete arms, rarely sparsely intermixed with caducous elongate smooth trichomes 1–1.5 mm long. Leaves of each pair somewhat anisophyllous in size; the petiole 0.5–3.9 cm long, canaliculate adaxially and shallowly grooved abaxially; larger blades 9–16 × 5–9.5 cm, smaller blades 3.5–10 × 2.1–5.5 cm, ovate to elliptic-ovate, the base rounded to obtuse, the margin ciliate and repand-entire, the apex shortacuminate, chartaceous; mature leaves with adaxial surface, primary, secondary, tertiary and higher order veins glabrous; abaxial surface essentially glabrous except for a few dendritic trichomes and resinous glands on the venules, the dendritic trichomes on the secondary and tertiary veins sparsely intermixed with caducous resinous slightly furrowed more or less stalked glands ca. 0.08 mm long, the higher order veins moderately beset with the same type of resinous glands; 5- or 7-plinerved, including the tenuous marginals, innermost pair of secondary veins diverging slightly to notably asymmetrically from the primary vein 0.25–0.35 cm above the base, areolae 0.5–0.75 mm, adaxially the primary and secondary veins slightly impressed to flat, the tertiary and higher order veins flat, abaxially the primary and secondary veins elevated and terete, the tertiary and higher order veins slightly elevated. Inflorescences a pseudolateral group of few-flowered modified dichasia 1.5–4 cm long, sessile, commonly with three or more paracladia arising from the base, borne on the upper foliar nodes, the rachis together with bracts and bracteoles pale magenta; bracts 0.65–0.75 × ca. 0.25 mm, spatulate-oblong, aristate at the apex, decurrent at the base, sometimes with a swollen glabrous structure at the base abaxially, both surfaces glabrescent with age, persistent in flower and tardily deciduous in fruit; bracteoles 0.65–0.95 × 0.21–0.49 mm, sessile, spatulate-oblong or triangular, the apex aristate-ciliate, the base decurrent, with one visible central vein, both surfaces glabrescent with age, persistent in fruit. Flowers 5-merous, sessile or on pedicels <0.4 mm long. Hypanthia at anthesis 3–3.2 × 0.95–1.25 mm, free portion of hypanthium 0.8–1 mm long, urceolate and constricted distally into a cylindric neck, bluntly 10-ribbed, bright pink, copiously and caducously resinous with slightly furrowed more or less stalked glands ca. 0.05 mm long, sparsely intermixed with the general and caducous dendritic trichomes, ridged on the inner surface, glabrous, the torus densely to moderately ciliolate adaxially, rarely glabrous. Calyx open in bud and deciduous on fruiting hypanthia; tube 0.15–0.3 mm long, adaxially sparsely ciliolate to glabrescent, abaxially SYSTEMATICS OF THE OCTOPLEURA CLADE OF MICONIA Phytotaxa 179 (1) © 2014 Magnolia Press 95 sparsely resinous glandular with the same type of glands as the hypanthium; lobes 0.15–0.45 × 0.65–0.85 mm, rounded-triangular, the margin entire to vaguely undulate, the apex subacute, sparsely and caducously papillose on both surfaces; exterior calyx teeth ca. 0.5 mm long, subulate, inserted at the base of the calyx lobes, equaling or occasionally exceeding the lobes in length, caducously resinous-glandular on both surfaces. Petals 0.75–1.15 × 0.65–0.85 mm, obovate and somewhat asymmetrical, the margin entire to sinuate, the apex rounded-obtuse, rather asymmetric, cream to white, densely papillose on both surfaces, reflexed at anthesis. Stamens 10; filaments 0.95–1.25 × 0.25 mm, whitish, glabrous; anther thecae ca. 2 × 0.33–0.34 mm, linear-oblong and subulate, truncateacuminate at the apex, opening by one dorsally inclined pore 0.11–0.13 mm in diameter, pale yellow to yellow at anthesis; connective yellow, its prolongation and appendage 0.35–0.45 mm long, the appendage oblong-spatulate to somewhat deltoid, obtuse to truncate at the apex, minutely and caducously glandular, the glands sessile and rounded. Ovary 5-locular, completely inferior, 2–2.2 mm long at anthesis, the apical collar absent, the apex 0.13–0.2 mm in diameter, slightly depressed, caducously glandular-puberulent; style ca. 5 mm long, parallel-sided (i.e. terete), white, glabrous; stigma truncate to expanded truncate at anthesis. Berries 3.1–4.1 × 3–3.9 mm when dry, globose, initially bright pink but ripening bright purple-black, the hypanthium indumentum subpersistent at maturity. Seeds 0.3–0.45 × 0.16–0.18 mm, ovoid, angled, light-brown; lateral and antiraphal symmetrical planes ovate, the highest point near the central part of the seed; raphal zone suboblong, ca. 20% larger than the corpus of the seed, extending along its entire length, ventrally and longitudinally expanded, dark-brown; individual cells elongate, anticlinal boundaries channeled, undulate, with Ω- and U-type patterns; periclinal walls convex, lowdomed to nearly flat, microrelief punctate. Additional specimens studied:— COLOMBIA. Chocó: (Bahía Solano), Ciudad Mutis, Quebrada Seca, at waterfall near by, 29 m, 6.2°N, - 77.4°W, 6 February 2012, Almeda et al. 10470 (CAS, CHOCO, COL). COSTA RICA. Cartago: In forest in mountains above the Río Pacuare, near Platanillo, 800 m, 3 May 1956, Williams 19517 (EAP, US); Pendiente muy empinada que va desde los edificios del Instituto Interamericano de Ciencias Agrícolas de Turrialba hacia el Río Reventazón, 600 m, 19 September 1964, Jiménez 2380 (CR, NY); (Turrialba), Forested slope leading down to the Río Reventazón behind main building of CATIE, 9°53.5’N, 83°39’W, 560–600 m, 30 July 1985, Grayum & Hammel 5755 (CAS, MO); (Turrialba), Across the Río Reventazón from Interamerican Institute in Turrialba, 610 m, 21 July 1947, DeWolf 369 (US); Bosque húmedo y empinado entre el Inst, de Turrialba y el Río Reventazón., 600 m, 12 July 1965, Jiménez 3276 (CR, F, NY, US). San José: (Tarrazu), Estribaciones del Cerro Diamante, 9°32’30"N, 84°1’20"W, 500–600 m, 23 June 1998, Estrada et al. 1636 (CAS, CR); (Pérez Zeledón), Basin of El General, 9.1°N, - 83.29°W, 675–900 m, May 1940, Skutch 4948 (MO, US). Illustration:— None found. Common names and documented uses:— None recorded. Habitat, distribution and ecology:— This is a rare species known from few collections in the low-elevation forested slopes of primary rain and riparian forests, commonly near rivers, in Costa Rica at 560–900 m. Although M. neocoronata has long been considered to be endemic in Costa Rica, it was recently collected in Colombia, from the tropical wet forest in the department of Chocó at 29 m (Almeda et al. 10470, CAS!, CHOCO, COL). This occurrence suggests that the geographic and elevational range of this species is more extensive than expected. It probably occurs in suitable habitats in intervening areas of Panama (Fig. 15). Phenology:— Collected in flower and fruit in February, from May through July, and in September. Etymology:— The specific epithet comes from the Greek word neo (new), in reference to this new combination, and from the Latin word coron (a crown), that probably refers to the minute pubescence in the ovary apex. Gleason did not explain the designation of the name coronata in the protologue. Discussion:— Miconia neocoronata is distinct from its closest relatives by elimination of characters, particularly indumentum details. It differs from M. atropurpurea, its sister species, in hypanthial indumentum; in M. atropurpurea it consists of elongate smooth pinkish trichomes, and in M. neocoronata by the slightly furrowed more or less stalked resinous glands. Vegetatively and in floral characters M. neocoronata is very similar to M. quinquenervia. See Almeda (2004) for a thorough discussion of differences. Almeda (2004) commented on the slight difference in their connective appendages; although they are similar in shape and both are glandular, in M. neocoronata the edge is entire (vs. gland-edged). These two species are also close to M. reitziana, which shares the rusty-asperous vegetative indumentum and the resinous-glandular hypanthium, but lacks the decurrent leaf bases present in M. quinquenervia and its venation is prevailingly 7-plinerved (vs. 9-plinerved). Miconia quinquenervia is also distinct in its inflorescence architecture (dichasial dithyrsoid vs. groups of modified dichasia). The rusty- 96 Phytotaxa 179 (1) © 2014 Magnolia Press GAMBA & ALMEDA asperous trichomes in M. reitziana are almost concealed by the typically dense pink or red indumentum, rarely present throughout and completely absent on the foliar surfaces except for the margins in M. neocoronata. Miconia neocoronata has unique petal morphology, being obovate and somewhat asymmetrical (vs. more oblong in M. quinquenervia and M. reitziana), and its hypanthium is constricted above the ovary into a neck (vs. subcylindric to campanulate in M. reitziana and urceolate to campanulate in M. quinquenervia). Conservation status:— Endangered EN B2ab(iii). This species is rare and not known from any protected areas in CostaRica.Published as part of Gamba, Diana & Almeda, Frank, 2014, Systematics of the Octopleura Clade of Miconia (Melastomataceae: Miconieae) in Tropical America, pp. 1-174 in Phytotaxa 179 (1) on pages 95-97, DOI: 10.11646/phytotaxa.179.1.
FIGURE 1. Miconia glandulipetala Ocampo & Almeda. A. Branches with infructescences. B in A new species of Miconia (Melastomataceae: Miconieae) from the eastern slope of the Peruvian Andes
No full textFIGURE 1. Miconia glandulipetala Ocampo & Almeda. A. Branches with infructescences. B. Flower at anthesis. C. Longitudinal section of a flower (petals and stamens removed). D. Petal showing a subapical glandular hair. E. Stamens in lateral (left) and dorsal (right) views. F. Transversal section of an ovary. G. Mature berry.Published as part of Ocampo, Gilberto & Almeda, Frank, 2014, A new species of Miconia (Melastomataceae: Miconieae) from the eastern slope of the Peruvian Andes, pp. 166-172 in Phytotaxa 163 (3) on page 168, DOI: 10.11646/phytotaxa.163.3.3, http://zenodo.org/record/513237
Microlicia daneui R. B. Pacifico & Almeda 2022, sp. nov.
No full textMicrolicia daneui R.B.Pacifico & Almeda, sp. nov. (Figs. 16–17). Type:— BRAZIL. Bahia: Mucugê, arredores do Pico da Batávia, Parque Nacional da Chapada Diamantina, 13°18’30.3”S, 41°17’31.2”W, 1603 m, 24 June 2022, fl., fr., R. Pacifico 691, V.E. Bressan & L. Daneu (holotype: HUEM!, isotypes: CAS!, HUEFS!, RB!). Diagnosis:—Differs from Microlicia noblickii (Wurdack 1995: 821) A.B.Martins & Almeda in Almeda & Martins (2001: 4) by its leaves on petioles 4–12 mm long (vs. sessile), longer hypanthia 6–8 mm long (vs. 5–5.2 mm long), calyx lobes oblong to triangular 7–9 mm long (vs. deltoid to triangular, 3.5–4 mm long), petals 19–25 mm long (vs. 14– 16 mm long) that are entirely magenta (vs. pink with a yellow base), antepetalous pedoconnectives with appendages 1–1.3 mm long (vs. ca. 0.5 mm long) and ovaries 3–5-locular (vs. 5-locular). Erect shrubs 0.5–1.5 m tall, much-branched. Upper cauline internodes 4–7 mm long, light green (when fresh) or reddish, becoming pale brown (when dry) and defoliated with age, obscurely quadrangular to terete, not sulcate, unwinged, densely glandular-punctate and sparsely to densely covered with inconspicuous gland-tipped trichomes 0.1–0.3 mm long, Leaves decussate, ascending, concealing uppermost internodes (when dry), chartaceous to coriaceous, concolored and light green (younger leaves) to vivid green (older leaves when fresh), both leaf surfaces becoming brownish to blackened (when dry); petioles 0.4–1.2 mm long, light green flushed with red (when fresh), rectangular; blades 9–19 × 4–12 mm, narrowly to widely elliptic or slightly obovate, apex rounded or acute, base attenuate, margins entire and glabrous, flat, adaxial surface glandular-punctate to glabrescent, abaxial surface glandular-punctate and covered with inconspicuous gland-tipped trichomes 0.1–0.3 mm long mainly on the veins, eventually becoming glabrescent, 3-nerved from the base (basal acrodromous), venation prominent on the abaxial surface and impressed on the adaxial surface, secondary venation not evident. Flowers 5-merous on short pedicels 0.5–1 mm long, apical, solitary, ebracteolate. Hypanthia (at anthesis) 6–8 mm long, 5–7 mm wide at the torus, light green flushed with red (when fresh) becoming brownish (when dry), campanulate, surpassing the capsule in length at maturity, densely glandular-punctate and sparsely covered with inconspicuous gland-tipped trichomes 0.1–0.3 mm long. Calyx tubes 0.2–0.3 mm long. Calyx lobes 7–8 mm long, 4–5 mm wide at the base, light green flushed with red (when fresh) becoming brownish (when dry), patent at anthesis, oblong to triangular, apex rounded to truncate, margins entire and glabrous to inconspicuously ciliate with gland-tipped trichomes 0.1–0.3 mm long, externally with an indumentum like that of the hypanthia. Petals 19–25 × 14–16 mm, obovate, magenta, the base attenuate, apex acuminate, both surfaces glabrous, margins entire and glabrous. Stamens 10, dimorphic, antesepalous anthers ± horizontal (at anthesis), subparallel and clustered, antepetalous anthers ± erect and clustered near the floral axis; antesepalous (larger) stamens with filaments 7–8 mm long, magenta, glabrous, thecae (excluding rostra) 2.9–3.1 × 0.9–1 mm, yellow becoming brownish after pollination, oblong, slightly corrugate (polysporangiate), rostra 0.7–0.8 mm long, white, the ventrally inclined pores ca. 0.4–0.5 mm wide, nearly circular, pedoconnectives 8–9 mm long, magenta with a yellow apex, appendages 1.4–1.7 mm long, apex emarginate, yellow; antepetalous (smaller) stamens with filaments 6–7 mm long, magenta, glabrous, thecae (excluding rostra) 2.5–2.8 × 0.9–1 mm, yellow becoming brownish after pollination, oblong, slightly corrugated (polysporangiate), rostra 0.6–0.7 mm long, white, the ventrally inclined pores ca. 0.4–0.5 mm wide, nearly circular, pedoconnectives 3–3.5 mm long, magenta with a yellow apex, appendages 1–1.3 mm long, apex emarginate, yellow. Ovary (at anthesis) ca. 4 × 3 mm, superior, subglobose, glabrous, 3–5-locular, adnate to the hypanthium basally for ½ of its length; style 11–12 mm long, magenta, glabrous, sigmoid, stigma punctiform. Fruit at maturity a subglobose loculicidal capsule ca. 5–6 × 5–6 mm, pale brown, glabrous, 3–5-valvate, enveloping hypanthia rupturing and flaking away with age, dehiscent from the apex to the base (basipetal), columellas deciduous. Seeds ca. 1 × 0.5 mm, yellow, oblong, testa foveolate, raphal zone nearly circular, ca. 20–30% the length of the seed. Additional specimens examined:— BRAZIL. Bahia: Mucugê [“Ibicoara”], Serra da Batávia, 13°18’26”S, 41°17’31”W, 1578 m, 23 June 2012, fl., fr., H .A. Ogasawara et al. 222 (ALCB!, HUFU-online image!). Mucugê. Arredores do Pico da Batávia, Parque Nacional da Chapada Diamantina, 13°18’30.3”S, 41°17’31.2”W, 1603 m, 24 June 2022, fl., fr., R . Pacifico et al. 690 (CAS!, HUEM!, HUEFS!, RB!). Distribution, habitat and phenology:— Apparently endemic to the campo rupestre of Serra da Batávia in Mucugê, Bahia, Brazil (Fig. 12, Appendix 2). It was collected at elevations of 1578–1603 m, in areas exposed to full sun, flowering and fruiting in June. Etymology:— The epithet honors our friend and colleague Lukas Halla Daneu (b. 1983–). During the last decade, Lukas has been a valued field assistant on several botanical expeditions to the interior of Bahia that have resulted in the collection of many new species and a significant increase in the floristic sampling for little-explored regions. Notes:— Microlicia daneui seems to be related to Microlicia noblickii (see diagnosis; Fig. 5E) and Microlicia mucugensis Wurdack (1988: 294) Almeda & A.B.Martins (2001: 4). These two putative relatives have sessile leaves and were initially described in Lavoisiera Candolle (1828: 102) by Wurdack (1988, 1995). Names in Microlicia for both species were later provided by Almeda & Martins (2001). Microlicia mucugensis shares with M. daneui the glandular-punctate indumentum on branches, leaves and hypanthia, somewhat elliptic leaf blades that are rigid and become blackened when dry, magenta petals and dimorphic yellow stamens. Microlicia daneui differs by the laxly imbricated leaves (vs. not imbricated) on petioles 0.4–1.2 mm long (vs. sessile), modally longer blades 9–19 mm long (vs. 5–10 mm long), longer hypanthia 6–8 mm long (vs. 4.5– 4.8 mm long), calyx lobes 7–8 mm long (vs. 3.5–4.2 mm long), petals 19–25 mm long (vs. 12–14 mm long), stamens with filaments 6–8 mm long (vs. 4.3–5.5 mm long), and polysporangiate anthers (vs. tetrasporangiate). Microlicia daneui is probably parapatric with the compared species. Both M. noblickii (Fig. 5E) and M. mucugensis have never been reported for Serra da Batávia (Appendix 1). Suggested conservation status:— Data Deficient (DD; see Appendix 2).Published as part of Pacifico, Ricardo & Almeda, Frank, 2022, New species of Marcetia and Microlicia (Melastomataceae) endemic to the campo rupestre of Chapada Diamantina, Bahia, Brazil, pp. 39-69 in Phytotaxa 573 (1) on pages 55-57, DOI: 10.11646/phytotaxa.573.1.3, http://zenodo.org/record/732945
Miconia quadridomius Gamba & Almeda 2014, nom. nov.
No full text24. Miconia quadridomius Gamba & Almeda, nom. nov. Basionym: Clidemia cuatrecasasii Wurdack (1981: 248–249). Type: COLOMBIA. Dept. Valle: Puerto Merizalde, costa del pacífico, Río Naya, 5–20 m, 20–23 February 1943, Cuatrecasas 13957 (holotype: US!). Nec Miconia cuatrecasae Markgraf in Cuatrecasas (1933: 27). Openly branched shrub or small tree (1.5–) 2–6 m tall. Upper internodes rounded-quadrate, (2.1–) 3.2–4.9 cm long, cauline nodes terete, nodal line absent. Indumentum on branchlets, petioles, surface of young leaves, primary and secondary veins adaxially, primary and secondary leaf veins abaxially, bracts apically, pedicels, hypanthia, calyx lobes abaxially, and exterior calyx teeth densely to copiously composed of ferruginous elongate slightly or moderately roughened trichomes 2–3 mm long, each trichome deflexed and somewhat flattened, densely intermixed with an understory of clavate dendritic trichomes 0.1–0.3 mm long with short to moderately long thinwalled (flattened) arms. Leaves of each pair slightly to commonly anisophyllous in size; the semiterete short petioles 0.8–1.5 mm long, superficially canaliculate adaxially; larger blades 18–27 × 6.7–10.2 cm, smaller blades 7–16 × 2.5–6.5 cm, elliptic to elliptic-obovate, the base typically rounded, frequently becoming attenuate but ending in a rounded-cordate base, the margin serrulate to crenulate, the apex acuminate to caudate-acuminate, chartaceous; mature leaves adaxially with both the dendritic and elongate trichomes on the surface, primary and secondary veins becoming sparse to caducous with age, the tertiary and higher order veins glabrescent; abaxial surface glabrous, the tertiary and higher order veins sparsely and caducously covered with the general dendritic trichomes; 5-(7-)plinerved, including the tenuous marginals, innermost pair of secondary veins diverging from the primary vein 0.3–2 cm above the base, forming a deeply tufted cavity beset with the general indumentum (acarodomatia?), areolae 0.3–0.4 mm, reticulation visible on both surfaces, adaxially the primary and secondary veins slightly impressed, the tertiary and higher order veins flat, abaxially the primary and secondary veins elevated and terete, the tertiary and higher order veins slightly elevated. Inflorescences a congested axillary fasciculate glomerule 1–1.5 cm long, sessile, unbranched, typically paired and seemingly cauliflorous on defoliated nodes; bracts 1.5–7 × 0.4–0.6 mm, subulate, thin, erect, with inconspicuous parallel venation, glabrous on the main surfaces but the general elongate roughened indumentum present at the apex, each bract seemingly branched, persistent to tardily deciduous in fruit. Flowers 4-merous on thick pedicels 0.4–1 mm long. Hypanthia at anthesis 2.7–3 × 0.9–1 mm, free portion of hypanthium 1.3–1.7 mm long, tubular to suburceolate, bluntly 8-ribbed, ridged on the inner surface, moderately scaly, the torus adaxially copiously beset with short-stalked glands with thinwalled short heads. Calyx open in bud and persistent in fruit; tube 0.1–0.3 mm long, adaxially with the same type of glands as the torus, abaxially with the same vestiture as the hypanthium; lobes 1.5–2 × 0.8–1.1 mm, ovate-oblong, 108 Phytotaxa 179 (1) © 2014 Magnolia Press GAMBA & ALMEDA the margin entire, the apex retuse to obtuse, the adaxial surface glabrous or minutely resinous-puberulent, reflexed at anthesis; exterior calyx teeth 1–1.5 mm long, subulate, inserted at the base of the calyx lobes and barely spreading beyond them. Petals 1.5–3.5 × 0.6–1 mm, oblong to linear-oblong, the margin entire, the apex roundedobtuse, white, glabrous on both surfaces, reflexed at anthesis. Stamens 8; filaments 1.4–1.5 × ca. 0.2 mm, white, glabrous; anther thecae 1.3–1.5 × 0.23–0.36 mm, linear-oblong and slightly clavate, somewhat emarginate at the apex, opening by one dorsally inclined pore 0.1 mm; connective darker than the thecae, its prolongation and appendage 0.2–0.3(–0.5) mm long, the appendage lanceolate, bluntly acute at the apex, copiously gland-edged, the glands rounded and conspicuously stalked to 0.2 mm long. Ovary 4-locular, 3/4 to completely inferior, 1.3–1.45 mm long at anthesis, the apical collar 0.3–0.4 × 0.55–0.65 mm, conic, copiously glandular-ciliate; style 5–7 mm long, parallel-sided (i.e. terete), white, glabrous; stigma truncate to capitellate when dry. Berries 2–2.5 × 2 mm when dry, globose and slightly oblate, bright orange when ripe, the hypanthium indumentum persistent in fruit. Seeds 0.68–0.79 × 0.5–0.65 mm, pyramidal, brown; lateral symmetrical plane triangular, the highest point near the central part of the seed; antiraphal symmetrical plane suboblong; raphal zone circular to suboblong, ca. 40% the length of the seed; multicellular sculpture rugose throughout the seed; individual cells elongate, anticlinal boundaries channeled, irregularly curved; periclinal walls convex, low-domed to nearly flat, microrelief striate. Additional specimens studied:— COLOMBIA. Chocó: Vía de Morro de Mico al “Mirador” por el camino hacia “Jurubidá” en dirección Sy posteriormente hacia “Copete de Pava” en dirección N, 6°5’N, 77°10’W, 0–100 m, 15 May 1990, Barbosa 6598 (MO, US); P.N. de Utría, Serranía ubicada al NE de la Ensenada de Utría por la trocha llamada del M-19, 6°20’N, 77°20’W, 0–100 m, 12 June 1990, García-Cossio & Agualimpia 500 (CHOCO, MO); Hoya del Río San Juan, Quebrada Taparal, afluente del Río San Juan, 4°12’N, 77°8’W, 5–10 m, 28 March 1979, Forero et al. 4247 (COL, MO). Valle: (Buenaventura), Along road between Buenaventura and Málaga vicinity of Bajo Calima, km 3.5.2 from main Cali-Buenaventura Hwy, at Gallinero, 4°0’N, 77°3’W, 100 m, 15 July 1993, Croat & Bay 75749 (MO); (Buenaventura), Bajo Calima, Ca, 15 km Nof Buenaventura, Cartón de Colombia concession, Dindo area, 3°59’N, 77°2’W, 50 m, 26 March 1986, Gentry et al. 53646 (MO, US); (Buenaventura), Bajo Calima, ca. 10 km due Nof Buenaventura, Cartón de Colombia concession, 3°56’N, 77°8’W, ca. 50 m, 5 December 1981, Gentry 35354 (CAS, MO); (Buenaventura), Bajo Calima, ca. 15 km Nof Buenventura, Cartón de Colombia concession, 3°56’N, 77°8’W, ca. 50 m, 18 February 1983, Gentry & Juncosa 40492 (MO); (Buenaventura), Bajo Calima, Granja Agroforestal, 40 m, 29 March 1984, Devia 491 (MO, TULV); (Buenaventura), Bajo Calima, Estación Agroforestal, Secretaría de Agricultura y fomento, Parte NE del campamento, 40–60 m, 5 August 1979, Cabrera 5181 (CUVC, MO); (Buenaventura), San Isidro, Bosque INDERENA-CONIF, 40 m, 5 March 1989, Devia & Prado 2641 (CAS, TULV); (Cordoba), Dagua Valley, 80–100 m, 6 May 1922, Killip 5119 (US); La Trojita, Río Calima (región del Chocó), 5–50 m, 19 February 1944, Cuatrecasas 16623 (US). ECUADOR. Esmeraldas: Road Lita-Alto Tambo-San Lorenzo, km 6.9 from Lita, 0°52’24.6"N, 78°29’33.2"W, 720 m, 30 September 2001, Cotton et al. 1794 (CAS, QCA). PANAMA. Comarca de San Blás: Headwaters of Río Nergala along continental divide, 350 m, 11 January 1985, de Nevers & Herrera 4515 (CAS, MO); Llano-Cartí Road, km 16, trail to creek on Atlantic drainage, 250–350 m, 2 February 1989, Almeda et al. 6522 (CAS, MO, NY, PMA); Nusagandi, Along continental divide on El Llano-Carti road, Headwaters of Atlantic draining creeks, 9°19’N, 78°15’W, 320 m, 12 August 1984, de Nevers & Pérez 3694 (CAS). Panamá: Along El Llano Carti-Tupile road, 12 mi above Pan-Am Hwy, 200–500 m, 26 March 1973, Liesner 1135 (CAS, NY). Illustration:— None found. Common names and documented uses:— None recorded. Habitat, distribution and ecology:— Local and uncommon in understories of rain forests, typically in deep shade and/or along streams in Panama, Colombia and Ecuador (Fig. 16), at 0–720 m. It is most common in the Bajo Calima region (part of the Chocó) in the department of Valle in Colombia. It was recently reported from Panama (Almeda 2009). Specimens from Ecuador that were previously attributed to this species are M. chocoensis. Phenology:— Collected in flower from January through March and from May through August; in fruit from February through March, June through September, and in December. Etymology:— The specific epithet honors José Cuatrecasas (1903–1996), Spanish botanist and prolific collector of Colombian plants. Discussion:— This distinctive species has an indumentum of elongate-roughened trichomes on vegetative and hypanthial parts, conspicuous 5-plinerved leaves and a poorly developed-sessile inflorescence. Almeda (2009) SYSTEMATICS OF THE OCTOPLEURA CLADE OF MICONIA Phytotaxa 179 (1) © 2014 Magnolia Press 109 noted that the Panamanian populations differ from those in Colombia by the presence of shorter bracts to 2 mm long (vs. 6–7 mm long) calyx lobes adaxially glabrous (vs. resinous-puberulent); we agree with these observations. In all other characters, the material from both countries is similar in foliar shape, indumentum details, and staminal and seed morphology. In M. quadridomius the leaves are plinerved, but there is a space formed in between the innermost pair of secondary veins and the primary vein before the point of divergence. This space may serve to house insects. Althought some ants were found in this leaf area among different specimens from the Bajo Calima region (Colombia), the morphology of this structure is not a usual ant-domatium. It is more similar to an acarodomatium; further natural history studies are required in order to elucidate the function of this structure. This species is similar to M. chocoensis, but differs in the foliar plinervation and in lacking white furrowed glands on the leaves abaxially. In M. quadridomius the vegetative and hypanthial indumentum is ferrugineous (vs. white), and consists of prominently roughened flattened-deflexed trichomes (vs. slightly roughened). Miconia quadridomius is also similar to M. approximata, which has vegetative pubescence that is shorter (0.3–0.7 mm long vs. 2–3 mm long), and the abaxial tertiary and higher order foliar veins densely resinous-glandular (vs. furfuraceous). In the protologue, Wurdack (1981) provides a detailed enumeration of the differences between these species. Conservation status:— Endangered EN B2ab(iii). Protected only in Colombia in the Ensenada de Utría National Park (Chocó); the threats include destruction of its natural habitat and the fact that it is not protected in other parts of its range.Published as part of Gamba, Diana & Almeda, Frank, 2014, Systematics of the Octopleura Clade of Miconia (Melastomataceae: Miconieae) in Tropical America, pp. 1-174 in Phytotaxa 179 (1) on pages 108-110, DOI: 10.11646/phytotaxa.179.1.
- …
