1,361 research outputs found
Tranzscheliella helutae K. G. Savchenko 2023, sp. nov.
<i>Tranzscheliella helutae</i> K.G. Savchenko <i>sp. nov.</i> <p>MycoBank: 846947</p> <p>Etymology: After a Ukrainian mycologist Dr. Vasyl P. Heluta.</p> <p>Sori in the culms and surrounding the upper internodes and axes of abortive inflorescences, initially covered by the leaf sheath, later exposed. Peridium absent. Spore mass semi-agglutinated to powdery, dark brown. Spores globose to subglobose, 4.5–5.5 × 5–6.5 μm, dark olive-brown. Spores rarely with polar cups, medium brown, without appendages; spore wall c. 0.5–1 μm, smooth. In SEM unevenly, moderately verruculose.</p> <p> Typification: UKRAINE. Kherson Oblast: Hola Prystan District, Chernomorsky Biosphere Reserve, 46°45′15′′ N, 32°12′74′′ E, 7 m. asl, On <i>Agropyron cristatum,</i> 07 June 2009, K. G. Savchenko, BUT-F 304 (holotype).</p> <p> Other specimens examined: UKRAINE. Kherson Oblast: Hola Prystan District, Chernomorsky Biosphere Reserve, 46°31′35′′ N, 31°94′22′′ E, 1 m. asl, on <i>Agropyron cristatum</i>, 09 June 2009, K. G. Savchenko, BUT-F 305 (paratype).</p> <p> Note— <i>Tranzscheliella helutae</i> is a part of <i>T. hypodytes</i> species complex. <i>Tranzscheliella helutae</i> is the only species from this genus occurring on <i>Agropyron</i> (subfamily Pooideae, tribe Triticeae). <i>Agropyron</i> contains about seven species found in temperate, mostly arid regions of Eurasia (Dewey 1986). Several species of <i>Agropyron</i> were listed as hosts of <i>T. hypodytes</i> s.l. by Vánky (2012). <i>Tranzscheliella helutae</i> has slightly larger and darker spores than <i>T. hypodytes</i> s.str. (4–4.5 × 4.5–5.5 μm). The spores of <i>T. helutae</i> are unevenly, and moderately verruculose in SEM, which differ from the densely, minutely, uniformly verruculose spores of <i>T. hypodytes</i> s.str. In the phylogenetic analysis, specimens of <i>T. helutae</i> were resolved in a well-supported monophyletic clade (Fig. 1). In a concatenated alignment there are 25 base pair differences between <i>T. helutae</i> and the most closely related species, <i>T. yupeitaniae.</i></p>Published as part of <i>Savchenko, Kyryll & Mahmood, Huda, 2023, Tranzscheliella helutae sp. nov. on Agropyron cristatum from Ukraine, pp. 151-156 in Phytotaxa 592 (2)</i> on pages 154-155, DOI: 10.11646/phytotaxa.592.2.9, <a href="http://zenodo.org/record/7840526">http://zenodo.org/record/7840526</a>
Tipula (Pterelachisus) octomaculata Savchenko
Tipula (Pterelachisus) octomaculata Savchenko Figures 1 a, 2 a, 3 a, b Savchenko 1964: 44 (description), Fig. 19 (hypopygium lateral, 9 th tergite dorsal, inner gonostylus lateral); Theowald 1980: 454, Fig. 355 (a 9 th tergite dorsal, b inner gonostylus lateral, c outer gonostylus lateral, all redrawn after the original description). Material examined: Finland, Ks: Taivalkoski, Lehtorinnanoja 7289411: 3560846, 3.7.- 4.8.2006 Malaise trap, J. Salmela leg, 1 ♂ (PCJS, in alcohol). Other material studied: Tipula (Pterelachisus) angulata Loew USA: Minnesota, Clearwater Co # Itasca State Park, 4 July 1970 1500 ’, George W. Byers leg, 1 ♂ (ANS). Tipula (Pterelachisus) entomophthorae Alexander USA: Lake Tear 1310 m Essex Co. N.Y. 44.06.25-73.56.05 10 July 1980 McCabe & Teale leg, 1 ♂ (ANS). Tipula (Pterelachisus) trivittata Say Canada: Ontario, Ottawa 10 June 2001, F. Brodo leg, 1 ♂ (PCJS); Ont. Bells Corners 45 ° 17 ’N, 75 ° 48 ’W 2 July 1969 Fenja Brodo leg, 1 ♂ (PCJS). Redescription: Male. Head. Top of head brown, rostrum light brown. Hairs covering head brown to dark brown. Antenna 13 -segmented, scape, pedicel and 1 st flagellomere light brown, other flagellomeres dark brown. Palpus light brown. Thorax. General coloration of thorax brown (pruinosity not seen in specimens stored in alcohol, but holotype with grey pruinosity and three brownish prescutal stripes). Parts of post sutural scutum, katatergite, metakatepisternum and metaepimeron somewhat lighter. Stem of halter light, yellowish, knob infuscated. Wing 17 mm, patterned with light and dark areas. Dark clouds present at base of Rs, R 2 and R 3, tip of wing infuscated. Dark seams along CuA, CuA 2 and distal half of A 2. Discal cell and area around R 1 + 2 light. Coxae dark brown, trochanters light brown. Base of hind femur light brown, other parts of leg missing. Other legs missing. Abdomen. 1 st segment of abdomen dark brown, 2 nd segment yellowish, with narrow dark longitudinal stripe both dorsally and ventrally. Hairs of segments light. Male terminalia. 9 th tergite (Fig. 1 a) with deep median emargination, bearing two close spines in midpoint. Inner margin of emargination with blunt, weakly developed outgrowth. Gonocoxite characterized by relatively dense setosity (Figs. 3 a, b), aedeagal guides conspicuous, darkly sclerotized. Outer gonostylus (Fig. 2 a) humpbacked (subsigmoid), apically rounded, about three times longer than wide at mid point, bearing hairs subequal in length to width of outer gonostylus. Inner gonostylus (Fig. 2 a) with conspicuous, sclerotized outer basal lobe, with sub-apical hyaline seta. Sub-basal spine sclerotized, lying along dorsal margin of inner gonostylus. Lower beak of equal width along its entire length, with moderate, pointed outgrowth on dorsal side of apex. Dorsal crest dark brown, resembling hammer in lateral view. Female unknown. Discussion: The identity of the Finnish specimen is based on the original description by Savchenko (1964) and its subsequent translation by Theowald (1980). The holotype deposited at St. Petersburg was not studied by us directly, but we were able to examine photos of the holotype male. The similarity between the original description and the specimen from Finland is remarkable, so we do not hesitate to identify it as T. octomaculata. However, the only notable variation observed was that the sub-basal spine of the inner gonostylus is projected from the dorsal margin of the gonostylus in the holotype but lying along it in the Finnish specimen. According to the German translation of the original description (Theowald 1980), the outer gonostylus (“od”), is “Einen breiten lanzettförmigen od”. The outer gonostylus was not entirely figured by Savchenko (1964), the apex of it is hidden by the 9 th tergite, but Theowald has drawn the apex too, giving it an oval appereance, although Theowald did not see the specimen. Thus, we assume that the drawing by Theowald is misleading. In the Finnish specimen the outer gonostylus is humpbacked, widest at the midpoint, with the apex clearly narrower than the midpoint. Additionally, in the original description (Savchenko 1964, Fig. 19) and subsequent translation (Theowald 1980, Fig. 355 a) the 9 th tergite in dorsal view is somewhat confusing. Both the holotype and the specimen from Finland have somewhat wider, conical caudal lobes of the 9 th tergite; the lateral outline of the 9 th tergite in dorsal view is not as constricted as that figured by Savchenko (1964) and Theowald (1980); the outgrowth of the inner margin is blunt, not as sharp as that figured in Savchenko (1964) and Theowald (1980). FIGURE 1. Male 9 th tergite and inner gonostylus, dorsal view. a) Tipula octomaculata Savchenko, b) T. angulata Loew c) T. entomophthorae Alexander, d) T. trivittata Say. Savchenko (1964) states that T. octomaculata is similar regarding the wing pattern to species around T. irrorata Macquart, T. procliva Alexander and T. caerulea Mannheims & Nielsen. However, T. octomaculata is clearly distinguished from T. irrorata and related species by the structure of 9 th tergite. T. procliva has a quite similar inner gonostylus (see Alexander 1938, original description or Savchenko 1964, redrawn on p. 46), considering the dorsal crest and outer basal lobe, but the 9 th tergite of T. procliva has a strong spine in the midpoint of the emargination whereas in T. octomaculata there are two close spines. T. caerulea is a synonym of T. laetibasis Alexander (Oosterbroek 2006), a species quite different from T. octomaculata in all aspects of male terminalia (see, for example Savchenko 1964; Martinovsky 1974). There are not many species of Tipula (Pterelachisus) having a similar set of spines in the mid-point of the emargination of the 9 th tergite. Of the Holarctic species, we found only T. trivittata having two spines in the 9 th tergite (Fig. 1 d). T. trivittata, however, has a pointed, well developed outgrowth in the inner margin of the emargination: this outgrowth is weak and blunt in T. octomaculata. Further, the basal part of the inner gonostylus in T. trivittata has no sclerotized, pointed lobe or sub-basal spine. In addition, the lower beak resembles a spoon: it is rounded and very wide distally. The outer gonostylus is digitiform, very narrow. The male 9 th tergite of T. trivittata has been figured by Alexander (1919 a; 1966). Tipula angulata, a species quite close to T. trivittata, is distinguished from T. octomaculata by differences in the 9 th tergite (Fig. 1 b) (median emargination relatively small, only a weak protuberance in the mid-point, caudal margin truncate), outer gonostylus (digitiform and apex broadly rounded) and inner gonostylus (no spine in the outer basal lobe of the inner gonostylus; lower beak quite narrow compared to other species in lateral view). The male 9 th tergite of T. angulata has been figured by Alexander (1966). The closest relative of T. octomaculata is probably T. entomophthorae. The basic structure of the male hypopygia of the species is very similar, but in T. entomophthorae there is only a weak protuberance in the mid-point of the emargination of the 9 th tergite (Fig. 1 c); the inner margin of the emargination has a sub-basal, blunt outgrowth; the outer basal lobe of the inner gonostylus (Fig. 2 b) is stronger in T. octomaculata; the lower beak is relatively wide in the apical part (Fig. 2 b); and the flap in the margin of the dorsal crest is roughsurfaced, whereas it is even in T. octomaculata. The male 9 th tergite of T. entomophthorae has been figured by Alexander (1966). See also Alexander (1918) (original description) and Dietz (1921) (syn. T. similissima). Tipula (P.) penobscot Alexander, 1915 is probably a fourth Nearctic species close to T. octomaculata and related species. T. penobscot seem to have a strong outer basal lobe of the inner gonostylus (Alexander 1915, plate XVIII, Fig. 32) but have a caudal margin of 9 th tergite which is quite different than that of other species: there is no strong emargination, margin is concave, slightly notched, with a sharp outgrowth in the midpoint (Alexander 1915, plate XIX, Fig. 45). In addition, Tipula (P.) diflava Alexander, 1919 (Alexander, 1919 b) perhaps should be mentioned here. This arctic species (Brodo 1990) has a U-shaped median emargination of the 9 th tergite, with a small median outgrowth in the midpoint; lobes of the caudal margin of the 9 th tergite are oblique. The outer gonostylus is wide and rounded at the apex, the base is very narrow. The inner gonostylus has a pointed basal lobe and a sub-basal spine, but dorsal crest and beak without peculiar structures. The holotype male of T. octomaculata was collected from Russia, Archangelskaja Oblast, Obozerskaja by N. Krivosheina from a spruce forest in 1959 (Savchenko 1964; Theowald 1980). So far, the species has been known only from the type locality. The new discovery of the species in Finland is from the North boreal ecoregion (65 ° 40 ’N) (see above Material examined). The locality is an approximately 150 m wide strip of old-growth forest surrounding a forest brook dominated by Norway spruce (Picea abies), deciduous trees being very scarce. The ground layer vegetation is characterized by tall herbs (eg. Filipendula ulmaria) and ferns (Dryopteris carthusiana, Phecopteris connectilis). Decaying spruce trunks are abundant at the site. Tipulid species such as Tipula limitata Schummel, T. pseudoirrorata Goethgebuer, T. grisescens Zetterstedt, T. variicornis (Schummel) and T. nubeculosa Meigen were also found at the site. The related Nearctic species discussed above are apparently confined to woods (eg. Alexander 1941; Petersen et al. 2005), except T. diflava, which is an arctic species (Brodo 1990).Published as part of Salmela, Jukka & Autio, Olli, 2007, Redescription of Tipula octomaculata Savchenko, with notes on related Holarctic species (Diptera, Tipulidae), pp. 53-58 in Zootaxa 1527 on pages 54-57, DOI: 10.5281/zenodo.17759
Oleksandra Yakivna Savchenko. Lessons of cooperation and communication: a collection of materials of the All-Ukrainian pedagogical readings in memory of O.Ya. Savchenko "Alexandra Yakivna Savchenko. Lessons of cooperation and communication ", Kyiv, Nov. 4
До збірника увійшли матеріали учасників І Всеукраїнських педагогічних читань пам’яті О.Я. Савченко «Олександра Яківна Савченко. Уроки співпраці і спілкування» (Київ, 4 листопада 2021 року), в яких висвітлені події життя і віхи діяльності видатного педагога, дидакта початкової освіти, автора численних наукових праць, навчально-методичних посібників для широкого кола освітян, підручників для учнів початкових класів закладів загальної середньої освітиThe collection includes materials of participants of the I All-Ukrainian pedagogical readings in memory of O.Ya. Savchenko "Alexandra Yakivna Savchenko. Lessons of Cooperation and Communication ”(Kyiv, November 4, 2021), which highlights the life and milestones of an outstanding teacher, didactic teacher, author of numerous scientific papers, teaching aids for a wide range of educators, textbooks for primary school students. secondary education
Contribution of O.Ya. Savchenko to the development of historical and pedagogical science of Ukraine
У статті автором проаналізовано науковий доробок О. Я. Савченко, зокрема зосереджено увагу на ідеях історико-педагогічного характеру, висловлених ученою у своїх працях.In the article the author analyzes the scientific achievements of O.Ya. Savchenko, in particular the attention is focused on the ideas of historical and pedagogical nature expressed by the scholar in her writings
Thaumastoptera (Thaumastoptera) intermixta Savchenko 1974
Thaumastoptera (Thaumastoptera) intermixta Savchenko, 1974 (Figs. 2, 7, 8) Thaumastoptera intermixta Savchenko 1974: 34 (description), Fig. 34 (male terminalia). Diagnosis. Small species, yet slightly larger than other species treated here. Body colouration in general pale yellow. Wing without any darker marks. Venation with upper branch of medial fork about 8 times as long as its petiole. Femora and tibiae without any darker rings. Male terminalia with aedeagus exceedingly long, by about one third its length longer than gonocoxite, tapered into subacute tip (Figs. 7, 8). Wing length 5.2–5.6 mm. Material examined: 2 males, 1 female. Russia: Krasnodar Territory, Sochi / Khosta (43°31'24"N 39°52'47"E), 15.v.2011, male (D. Gavryushin leg.) (JSO). Georgia: “Autonomous Soviet Socialist Republic of Adjara: …, …, 16.vi.1978, E. Savchenko leg.” [handwritten label in Cyrilic, illegible in part], male, female (JSO). Redescription. Male: Head. Suffused with dark greyish brown pruinosity on frons and vertex. Rostrum and palpus brown. Antenna 16-segmented, moderately long, reaching to about base of wing, brown throughout. Flagellomeres spherical proximally, passing into ovoid and narrower distally, with verticils moderately dense, longest ones about twice as long as their respective flagellomeres. Thorax. Generally yellow, mostly shiny dorsally, including pronotum, scutellum, and mediotergite. Pleuron likewise shiny yellow. Wing membrane tinged yellowish, veins yellow. No dark spots or darkly tinged sections of veins on wing. Venation generally usual for Thaumastoptera s. str. Sc1 ending about opposite slightly more than one third length of Rs; upper branch of medial fork (M1+2) differing from other species in being very long, about 8 times as long as its petiole (M1+2+3), or longer (Fig. 2). Halter yellow throughout. Legs pale yellow, without any darker rings. Abdomen. Yellow. Male terminalia (Figs. 7, 8) yellow. Segment 9 (basal ring) simple, more or less straight at dorsal posterior margin. Proctiger generally semioval, in part covered with spinulae, broadly rounded at posterior margin. Gonocoxite moderately long, slightly arched. Gonostylus rather long, produced into slightly curved, setose beak, directed medially and subacute at apex, and with another, broadly obtuse projection on outer margin of gonostylus before apex; latter projection with a few spinoid setulae. Group of longer setae at inner margin, at about half-length of gonostylus. Aedeagus exceedingly long, by about one third its length longer than gonocoxite, generally parallel-sided, rather broad proximally, abruptly narrowed to about half its breadth at its half-length, apex of aedeagus tapered into subacute tip. Paramere very short, just as lobe at base of aedeagus. Female. Resembling male in general appearance. Distribution. Russia: North Caucasus; Georgia. Distribution in Uzbekistan and Tajikistan given by Oosterbroek (2017) probably is not based on published primary data (V. Lantsov, pers. comm.). Discussion. Externally, T. (T.) intermixta differs from the other species treated here by its slightly larger size and its completely yellow body colouration, without any dark marks on the wings and any apical rings on the femora and tibiae. Male terminalia are conspicuous in having a very long aedeagus (Figs. 7, 8).Published as part of Starý, Jaroslav & Oboňa, Jozef, 2018, Palaearctic species of Thaumastoptera (s. str.) Mik (Diptera: Limoniidae), pp. 227-234 in Zootaxa 4394 (2) on pages 230-231, DOI: 10.11646/zootaxa.4394.2.5, http://zenodo.org/record/119767
O. Ya. Savchenko on the activities of the All-Ukrainian Association of Vasyl Sukhomlynskyi on the development of Sukhomlynskyi studies in educational and scientific space
Автором розглянуто низку статей голови Всеукраїнської асоціації Василя Сухомлинського академіка Олександри Яківни Савченко (1991 – 2020), присвячених розвитку сухомлиністики в освітньо-науковому просторі України та за її межами.The author considers some of articles by the head of the All-Ukrainian Association of Vasyl Sukhomlynskyi, Academician Oleksandra Yakivna Savchenko (1991 - 2020), devoted to the development of studies of Sukhomlynskyi studies in the educational and scientific space of Ukraine and abroad
Sustainable infrastructure technologies: impact and policy
This dissertation presents three studies that examine localized spatial effects of sustainable infrastructure investments in several contexts. The first chapter estimates public local economic benefits and costs of contaminated land cleanup and redevelopment through New York’s Brownfield Cleanup Program. Unlike prior literature, I value site redevelopment separately from cleanup, assess whether green redevelopment adds value, and examine time to capitalization. Using a property level fixed effects model and controlling for time-varying shocks, I find that the program has added 4% to nearby property values, producing a $1.26 billion property tax gain that outweighs its public costs. Most benefits stem from program participation and site cleanup. Furthermore, site entry and site cleanup do not immediately capitalize into nearby property values and are valued by the housing market with a lag of approximately three years.
The second chapter estimates the value of contaminated land cleanup through Illinois’ Site Remediation Program in the presence of foreclosures and demonstrates the potential consequences of not controlling for time-varying unobservables in hedonic regressions. Failure to control for incidence of foreclosures, which can act as a time-varying correlated unobservable, can lead to overestimation of the property effects of contaminated site participation in SRP by as much as 46% in absolute terms. Mean differencing of time-varying neighborhood amenities substantially reduces this bias. Given that foreclosures may be correlated with a myriad of neighborhood amenities, these findings provide cautionary evidence suggesting that studies employing hedonic analysis should control for time-varying market conditions.
The third chapter studies electricity consumption patterns generated by newly commissioned renewable energy plants in Indonesia to determine whether these facilities provide electricity to the regions in which they are located. This paper uses a novel application of satellite nighttime light data used as a proxy for energy consumption combined with data on geocoded locations of 15 renewable energy plants and Village Potential Statistics survey for Indonesia. I estimate a difference-in-difference model to compare the intensity of nighttime lights in nearby areas around incoming plants to that in geographically matched locations before and after each plant becomes operational. Results show that a 1% increase in capacity of a newly commissioned renewable energy plant leads to a 0.09% increase in nighttime lights relative to the matched control group. This corresponds to a 0.15% increase in residential electricity consumption and implies that local populations benefit, although to a small degree, from the energy these plants produce. Results further show that the immediate increase in nighttime lights associated with plant commissioning grows over time, but at a decreasing rate.Submission published under a 24 month embargo labeled 'Closed Access', the embargo will last until 2019-12-01The student, Olesya Savchenko, accepted the attached license on 2017-09-12 at 11:31.The student, Olesya Savchenko, submitted this Dissertation for approval on 2017-09-12 at 11:38.This Dissertation was approved for publication on 2017-09-13 at 14:02.DSpace SAF Submission Ingestion Package generated from Vireo submission #11642 on 2018-03-13 at 10:32:49Made available in DSpace on 2018-03-13T17:29:07Z (GMT). No. of bitstreams: 2
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Holophryxus quadratahumerale
HOLOPHRYXUS QUADRATAHUMERALE (SCHULTZ, 1978) Synonym: Isophryxus quadratahumerale Schultz, 1978. Original description: S c h u l t z (1 9 7 8— a s I. quadratahumerale): 79–82, fig. 9 (immature and adult ♀♀ and ♂).Published as part of Huys, Rony, Savchenko, Alexandra S. & Kosobokova, Ksenia N., 2023, Discovery of a new species and host record of Holophryxus Richardson, 1905 (Isopoda: Dajidae) from the central Arctic: a model of enhanced descriptive standards for epicaridean isopods, pp. 592-649 in Zoological Journal of the Linnean Society 198 (2) on page 639, DOI: 10.1093/zoolinnean/zlac105, http://zenodo.org/record/801100
A new species of the Y-larva genus <i>Hansenocaris</i> Itô, 1985 (Crustacea: Thecostraca: Facetotecta) from the Azores, with notes on its morphology and biogeography
A new facetotectan species, described herein as Hansenocaris spiridonovi Kolbasov, Savchenko et Høeg sp.n. and based on its cypridiform stage, was found in the plankton off Azores Islands. We employed scanning electron microscopy to document the fine-scale external morphology of this species. We discuss the findings of facetotectan larvae showing cosmopolitan distribution of these crustaceans. The presence of Y-cyprid exuviae in plankton, findings of y-larvae in the deep sea plankton, occurrence in the oceanic plankton, as well as their permanent presence in the tropical waters may evidence on a pelagic host.</p
Tipula (Vestiplex) coronifera Savchenko
Tipula (Vestiplex) coronifera Savchenko (Figs 25, 50–62; Map 3) Tipula (Vestiplex) coronifera Savchenko, 1960: 210; 1964: 201; Oosterbroek & Theowald, 1992: 154. Type material examined. HOLOTYPE ♂: MONGOLIA, Khangai Mts. (south-east), Lamyn-Gegen, 16 VII 1926 (Kirichenko), Nr 750 (ZIN). PARATYPE: ♀, Nr 770, topotypic (ZIN). Additional material examined. MONGOLIA. Uvs Aimag: 1 ♀, Khovd Soum, Shiver / Ireg Gol, 25 km NE Khovd, N49.43119, E91.19093, 1964 m, 5–6.vii.2010, MAIST, MAIS2010070504. Hovsgol Aimag: 1 ♂, Ulaan-Uul Soum, unnamed trib. of Beltes Gol 34 km NE Bayanzurh, 2113 m, N50.42980, E99.21690, 28.vi.2006, J. Gelhaus #1054-A, SRP06062803. Ovorhangay Aimag: 1 ♀, Zuunbayan-Ulaan Soum, Millenium Road A 0301 marker 380.6 km ~ 50 km NE Arvayheer, N46.60919, E103.10400, 1820 m, 5.vii.2004, SRPT, SRP04070502. Elevation range in Mongolia. Adults were collected at altitudes ranging from 1960 m to 2100 m. Period of activity. Adults are active from the end of June through to early July. Known distribution. Mongolia (Map 3). Redescription. Male. Body length 20.4 mm, wing length 20.8 mm. General body color brownish yellow. Head. Brownish yellow, vertex and occiput brown gray pruinose with brown median line. Rostrum brownish yellow, nasus distinct. Antenna 13-segmented, if bent backward reaching base of wing. Scape and pedicel yellow, first flagellomere yellow, darkened brown mesally. Flagellomeres 2–6 brownish, succeeding flagellomeres yellowish, apical flagellomere brownish, small, reduced, distinctly shorter than preceding flagellomere. Each flagellomere, except first, with darkened basal enlargement and moderately incised. Verticils shorter than respective segments. Palpus brown. Thorax. Brownish gray. Pronotum brownish yellow with brown median line. Prescutum and presutural scutum brown with 4 darker longitudinal stripes bordered by brown, intermediate stripes completely separated from each other. Interspace between median and lateral stripes yellowish with pale setae arising from dark sockets. Postsutural scutum brown. Scutal lobe with 2 indistinct spots bordered by brown. Scutellum brownish, postnotum brownishyellow, with brown median line. Pleura brown, sparsely dusted with gray. Coxae yellow, sparsely dusted with grey. Trochanters, femora and tibiae yellow. Tarsal segments indistinctly passing into brownish. Distal part of femora and tibiae narrowly darkened. Tarsal claws without tooth. Wing indistinctly patterned. Halter with pale stem and brown knob. Abdomen. Yellow; tergite 1 laterally darkened and sparsely pruinose. Dorsal median stripe indistinct, interrupted at caudal margins of segments. Lateral stripe brownish and distinct. Lateral margins of tergites pale. Sternite 1 sparsely dusted with gray. Hypopygium. Brown, at base slightly broader than abdomen. Tergite 9 in shape of large concave crescent sclerotised saucer (Fig. 50). Main body of tergal saucer brownish yellow with blackened rim. Posterior margin of tergal saucer broadly emarginated, with small denticles or teeth. Median V-shaped notch inconspicuous. Lateral angle of tergal saucer with broad obtuse tooth. Anterior and lateral portions of tergal saucer raised into sclerotised border. Border anteriorly inconspicuously toothed, laterally produced into acute tooth directed caudad and situated under lateral angle of tergal saucer so tergite 9 with 2 teeth in lateral view. Gonocoxite unarmed (Fig. 51). Outer gonostylus flattened, nearly oval (Fig. 52). Inner gonostylus in shape of claw-shaped plate, with 2 mid-dorsal teeth, anterior tooth twice as large as posterior. Beak extended into blackened, obtuse rostrum (Fig. 53). Gonocoxal fragment large with lateral and medial sclerites well-developed (Fig. 54). Medial sclerites fused, anterior apodeme long and broad, posterior part flat and arched. Lateral sclerite large and bilobed. Aedeagal guide in shape of long, slender tube (Fig. 55). Sperm pump with central vesicle small and flattened (Fig. 56). Compressor apodeme with median incision. Posterior immovable apodeme much longer than compressor apodeme, large and extended laterally into broad plate. Anterior immovable apodeme in shape of rounded plate. Aedeagus shaped as very long tube similar to T. (V.) subcentralis. Distal part ventrally membranous, shovel-shaped (Fig. 57). Female. Body length 26.8–27.0 mm, wing length 22.0– 26.6 mm. Antenna short, flagellomeres except first with very small incisions. Mesopleura brown, gray pruinose. Distal part of femora darkened. Ovipositor (Figs 58–62). Tergite 10 shiny, brown. Cercus brownish yellow, slender, about as long as tergite 10, with tip narrowed, outer margin almost smooth with small serration (Fig. 58). Hypovalva in shape of pale, elongated slender filament, with short trichia at tip (Fig. 59). Median incision between hypovalvae deeper than posterior margin of sternite 8. Lateral incision relatively deep and narrow (about twice as deep as maximal width). Lateral angle of sternite 8 rounded. Sternite 9 with anterior part slightly broadened, posterior part rounded (Fig. 60). Furca anteriorly narrowed, posteriorly broad (Fig. 61). Bursa copulatrix with spermathecal ducts sclerotised at base, in shape of lightly curved, slender dark brown process. Wall of bursa copulatrix on connection site with spermathecal ducts partially sclerotised, sclerotisation extended short distance along wall (Fig. 62). Cul-de-sac of bursa copulatrix curved. Spermatheca spherical, greatly broadened at base (Fig. 25). Remarks. An endemic species for Mongolia.Published as part of Starkevich, Pavel, Podenas, Sigitas & Gelhaus, Jon K., 2020, Taxonomic review of Tipula (Vestiplex Bezzi) crane flies (Diptera: Tipulidae) in Mongolia, pp. 1-88 in Zootaxa 4837 (1) on pages 32-35, DOI: 10.11646/zootaxa.4837.1.1, http://zenodo.org/record/445472
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