1,721,044 research outputs found
Cranial morphology of Jonkeria truculenta (Therapsida, Dinocephalia) and a taxonomic reassessment of the family Titanosuchidae
No full textTitanosuchidae are a group of herbivorous, long-snouted dinocephalians with definitive records known only from the middle Permian Tapinocephalus Assemblage Zone of the Beaufort Group (Karoo Supergroup) of South Africa. Here, the taxonomy of this family is revised; of the nine species currently recognized, only two are found to be valid: Titanosuchus ferox and Jonkeria truculenta, which can be distinguished on the basis of appendicular proportions. Jonkeria boonstrai, J. haughtoni, J. ingens, J. parva, J. rossouwi, and J. vanderbyli are synonymized with Jonkeria truculenta, and J. koupensis is considered a nomen dubium (Titanosuchidae indet.). Several new cranial features are described for J. truculenta, such as pachyostosis of the prefrontals, postorbitals and parietals, and an ontogenetic series for the species is presented.Fil: Jirah, Sifelani. University of the Witwatersrand; SudáfricaFil: Rubidge, Bruce. University of the Witwatersrand; SudáfricaFil: Abdala, Nestor Fernando. Consejo Nacional de Investigaciones Científicas y Técnicas. Centro Científico Tecnológico - Tucumán. Unidad Ejecutora Lillo; Argentin
Tooth replacement in the non-mammalian cynodont Cynosaurus suppostus (Therapsida) from the late Permian of South Africa
No full textFive specimens of the late Permian cynodont Cynosaurus suppostus, inferred to represent an ontogenetic growth series, were imaged using X-ray microtomography in order to determine the pattern of tooth replacement for the species. Due to the small sample size, and damage to the rostrum in several specimens, replacement patterns of the incisors could not be accurately determined. Evidence supporting the presence of two maxillary canine loci in a juvenile Cynosaurus is presented. Replacement of the mandibular canines ceased at a basal skull length of ∼88 mm, whereas replacement of the maxillary canines ceased at a basal skull length of ∼90 mm. The final canine generation to erupt in both the maxillae and mandible had open-rooted apices. An alternating pattern of replacement is described for the maxillary and mandibular postcanine series. Postcanine replacement generations in the maxilla are seemingly separated by two tooth loci, whereas in the mandible replacement activity occurs at alternating loci. Cessation of replacement in the first maxillary postcanine locus (PC1) is synchronized with the eruption of the replacement tooth in the third locus (PC3). This cessation of replacement at PC1 results in a distal migration of the entire maxillary postcanine series. This pattern provides possible evidence for a finite number of replacement generations occurring at the active PC1. There appear to have been two replacement cycles per postcanine locus in Cynosaurus, which is a lower number of total replacements per postcanine locus than previously estimated for the Early Triassic basal cynodont Thrinaxodon liorhinus.Fil: Norton, Luke A.. Evolutionary Studies Institute; SudáfricaFil: Abdala, Nestor Fernando. Consejo Nacional de Investigaciones Científicas y Técnicas. Centro Científico Tecnológico - Tucumán. Unidad Ejecutora Lillo; ArgentinaFil: Rubidge, Bruce. Evolutionary Studies Institute; SudáfricaFil: Botha, Jennifer. National Museum; Sudáfric
Postcranial morphology of the South African middle Permian pareiasaurs from the Karoo Basin of South Africa
No full textPareiasaurs were relatively abundant and globally distributed herbivorous parareptiles of the middle to late Permian. The basal-most pareiasaurs, the Bradysauria, are restricted to the middle Permian of South Africa and went extinct at the end of the Guadalupian (Capitanian) at the top of the Tapinocephalus Assemblage Zone. Currently, three genera are recognized in this group: Bradysaurus, Embrithosaurus and Nochelesaurus, but their postcrania are poorly known, and consequently poorly understood. In this paper, our third contribution designed to improve understanding of the Bradysauria, we present a detailed comparative postcranial description and updated diagnoses for Bradysaurus baini, Embrithosaurus schwarzi and Nochelesaurus alexanderi. Bradysaurus baini has one postcranialautapomorphy: anterior dorsal osteoderms smooth and strongly convex, with an incipient central boss, and very light ornamentation. Three pelvic autopomorphies of Embrithosaurus schwarzi are confirmed: anterior portion of the iliac blades flat and vertical (not evertedor upturned); iIiac blades diverge anteriorly, oriented at 45?60° off the sagittal plane; and a very thick pelvic symphysis. For Nochelesaurus alexanderi we remove all three of the postcranial autapomorphies previously proposed. To the diagnoses of each species, we have added several new distinguishing postcranial features, within the local group of middle Permian pareiasaurs. The results reinforce our previous cranial studies concluding that three valid species of pareiasaurs are represented in the South African middle Permian.Fil: Van den Brandt, Marc Johan. Evolutionary Studies Institute; SudáfricaFil: Benoit, Julien. Evolutionary Studies Institute; SudáfricaFil: Abdala, Nestor Fernando. Consejo Nacional de Investigaciones Científicas y Técnicas. Centro Científico Tecnológico - Tucumán. Unidad Ejecutora Lillo; ArgentinaFil: Rubidge, Bruce. Evolutionary Studies Institute; Sudáfric
A new large cynodont from the Late Permian (Lopingian) of the South African Karoo Basin and its phylogenetic significance
Get PDFThe Karoo Basin of South Africa has the best global record of Lopingian (Late Permian) non-mammaliaform cynodonts, currently represented by five species. We describe Vetusodon elikhulu gen. et sp. nov., documented by four specimens from the Daptocephalus Assemblage Zone. With a basal skull length of ~18 cm, it is the largest Lopingian cynodont and is also larger than Induan representatives of the group. Vetusodon elikhulu has a cranial morphology that departs notably from that previously documented for Permo-Triassic cynodonts. It features a short and extremely wide snout, resembling that of the contemporaneous therocephalian Moschorhinus, and has large incisors and canines that contrast with the small unicusped postcanines, suggesting a more important role of the anterior dentition for feeding. The dentary is extremely long and robust, with the posterior margin located closer to the craniomandibular joint than in other Lopingian and Induan cynodonts (e.g. Thrinaxodon). The secondary palate morphology of V. elikhulu is unique, being short and incomplete and with the posterior portion of the maxilla partly covering the vomer. A phylogenetic analysis suggests that V. elikhulu is the sister taxon of Eucynodontia and thus the most derived of the Lopingian to Induan cynodonts yet discovered.Fil: Abdala, Nestor Fernando. Consejo Nacional de Investigaciones Científicas y Técnicas. Centro Científico Tecnológico - Tucumán. Unidad Ejecutora Lillo; ArgentinaFil: Gaetano, Leandro Carlos. Universidad de Buenos Aires. Facultad de Ciencias Exactas y Naturales. Departamento de Geología; ArgentinaFil: Smith, Roger M. H.. Evolutionary Studies Institute; SudáfricaFil: Rubidge, Bruce. Evolutionary Studies Institute; Sudáfric
Cranial morphology and phylogenetic relationships of the Middle Permian pareiasaur Embrithosaurus schwarzi from the Karoo Basin of South Africa
No full textPareiasaurs were globally distributed, abundant, herbivorous parareptiles of the Middle to Late Permian, with the basal-most members found in the Middle Permian of South Africa. These basal taxa were particularly abundant and went extinct at the end of the Gaudalupian (Capitanian) at the top of the Tapinocephalus Assemblage Zone. Currently four taxa are recognized in this group: Bradysaurus seeleyi, B. baini, Nochelesaurus alexanderi and Embrithosaurus schwarzi, but they are all poorly understood. We here present the first detailed cranial description and updated diagnosis for Embrithosaurus schwarzi. No cranial autapomorphies were identified. However, Embrithosaurus schwarzi is a distinct taxon in this group, based on its unique dentition and using a combination of cranial features. It has nine marginal cusps on all maxillary and mandibular teeth, and wider maxillary teeth than in the co-occurring taxa, due to the marginal cusps being arranged more regularly around the crown, and the apex of the crown lacking the long, central, three-cusped trident. Our updated phylogenetic analysis recovers the four Middle Permian South African taxa as a monophyletic group for the first time, which we call Bradysauria, comprising a clade including Embrithosaurus, Bradysaurus baini and a polytomy including Nochelesaurus and Bradysaurus seeleyi.Fil: Van den Brandt, Marc Johan. University of the Witwatersrand; SudáfricaFil: Abdala, Nestor Fernando. University of the Witwatersrand; Sudáfrica. Consejo Nacional de Investigaciones Científicas y Técnicas. Centro Científico Tecnológico - Tucumán. Unidad Ejecutora Lillo; ArgentinaFil: Rubidge, Bruce Sidney. University of the Witwatersrand; Sudáfrica. DST National Research Foundation. Centre for Excellence in Palaeosciences ; Sudáfric
Reassessment of the Phylogeny of Basal Therapsida Using Micro-Tomography and Bayesian Phylogeny
No full textA thesis submitted in fulfilment of the requirements for the degree of Doctor of Philosophy (Evolutionary Studies Institute) to the Faculty of Science, School of Geosciences, University of the Witwatersrand, Johannesburg, 2023.This PhD research is focussed on the evolution and cranial anatomy of the earliest therapsids. The Therapsida is a paraphyletic group of land tetrapods that lived mostly during the Permian and Triassic Periods. They emerged quite suddenly in the fossil record during the middle Permian, already showing diversification into five distinct groups: Biarmosuchia, Dinocephalia, Anomodontia, Gorgonopsia and Therocephalia. A sixth group, the Cynodontia, appeared during the late Permian. Of these, the anomodonts, therocephalians, and cynodonts survived the devastating Permo-Triassic mass extinction and continued to be the dominant tetrapods during the Early and Middle Triassic Periods. Ultimately, the cynodonts would evolve into the earliest mammals. Chapter 1 introduces the scientific questions tackled in this thesis. Chapter 2 provides the necessary explanations about the material and methods used for this work. The use of CT-scanning technology allows for the detailed examination of cranial internal morphology, with the goal of incorporating inner cranial characters into a phylogenetic framework. Chapters 3 and 6 focus on the fossilisation gap at the early to middle Permian transition, which has led to debates over the timing of the early diversification of the Therapsida. In Chapter 3, the geologic position and anatomy of one of the oldest known therapsids, Raranimus dashankouensis is reviewed. The chapter concludes with a reassessment of the basal phylogenetic position of Raranimus and suggests a Roadian origin for the taxon. Chapter 6 presents a comprehensive analysis of cranial morphological characters based on the descriptions and findings from Chapters 3, 4, and 5. Using both traditional maximum parsimony and clock-based Bayesian inference of phylogeny for the first time on the Therapsida, the results suggest that therapsids originated around 280.5 million years ago and rapidly diversified into several distinct clades during the Kungurian and Roadian Epochs. The phylogenetic relationships of several taxa are reevaluated and the findings suggest that Biseridens is more closely related to the Dinocephalia and Biarmosuchia, rather than the Anomodontia. Furthermore, the results suggest that Sinophoneus may represent the basal-most dinocephalian, and that Therocephalia might be paraphyletic. Therapsids were characterised by a wide range of cranial morphologies, with the Biarmosuchia and Anomod ontia exhibiting distinct cranial features. Both groups are considered basal among therapsids. Chapter 4 of this PhD research focuses on the postnatal ontogenetic development of cranial ornamentation in Biarmosuchia and possible impact on phylogeny. The results suggests that cranial bosses and ridges are ontogenetic features and the parietal bone originates from multiple centres of ossification. Chapter 5 is a comprehensive cranial description of several basal anomodont specimens. The Chapter concludes with a review of the taxonomy of the genus Eodicynodon and proposes the creation of a new taxonomical unit for specimen NMQR 2913.National Research Foundation (NRF).GENUS.University of the Witwatersrand, Johannesburg.MMM202
The Taxonomy and Phylogeny of Varanopidae from the Middle Permian of outh Africa
No full textA dissertation submitted in fulfilment of the requirements of Master of Science, to the Faculty of Science, School of Geosciences, University of the Witwatersrand, Johannesburg, 2024.Varanopidae are the only pelycosaur-grade tetrapods from the Abrahamskraal Formation of the Beaufort Group. Although four varanopid species are described from the middle Permian of South Africa (SA), their validity has been questionable. Moreover, Ford and Benson recently proposed that varanopids belong to the Diapsida rather than Synapsida. Given this taxonomic and phylogenetic turmoil, this study i) describes two newly discovered and well-preserved varanopid specimens from the middle Permian of SA using CT scanning and ii) re-assesses the validity of the four species from the Karoo Basin in the light of new data from these specimens. The phylogeny of the group is addressed by updating two pre-existing cladistics matrices. The two new specimens exhibit a mosaic of features that overlap the diagnoses of several species of SA varanopids. In addition, some diagnostic traits of the four SA varanopid genera and species are found to be variable across ontogeny. For instance, dorsal osteoderms and the extent of ornamentation on the angular bone co-vary with size. It is therefore proposed that all varanopid specimens currently described from the middle Permian of South Africa belong to a single valid species, Heleosaurus scholtzi, with differences between specimens being accounted for by ontogenetic changes. The phylogenetic analysis supports Ford and Benson’s hypothesis but contra the prediction of this hypothesis, no differences in cranial fenestration during ontogeny are observed. Morphological characters preserved in the manus of the newly discovered specimen BP/1/8499 suggest that Heleosaurus was arboreal.National Research Fund (NRF)MMM202
Going Beyond Counting First Authors in Author Co-citation Analysis
Get PDFThe present study examines one of the fundamental aspects of author co-citation analysis (ACA) - the way co-citation
counts are defined. Co-citation counting provides the data on which all subsequent statistical analyses and mappings
are based, and we compare ACA results based on two different types of co-citation counting - the traditional type that
only counts the first one among a cited work's authors on the one hand and a non-traditional type that takes into
account the first 5 authors of a cited work on the other hand. Results indicate that the picture produced through this non-traditional author co-citation counting contains more coherent author groups and is therefore considerably clearer. However, this picture represents fewer specialties in the research field being studied than that produced through the traditional first-author co-citation counting when the same number of top-ranked authors is selected and analyzed. Reasons for these effects are discussed
Tooth replacement patterns in the Early Triassic epicynodont Galesaurus planiceps (Therapsida, Cynodontia)
Get PDFSixteen specimens of the Early Triassic cynodont Galesaurus planiceps (including eight that were scanned using micro-computed tomography) representing different ontogenetic stages were assembled to study the dental replacement in the species. The growth series shows that the incisors and postcanines continue to develop and replace, even in the largest (presumably oldest) specimen. In contrast, replacement of the canines ceased with the attainment of skeletal maturity, at a basal skull length of ~90 mm, suggesting that Galesaurus had a finite number of canine replacement cycles. Additionally, the functional canine root morphology of these larger specimens showed a tendency to be open-rooted, a condition not previously reported in Mesozoic theriodonts. An alternating pattern of tooth replacement was documented in the maxillary and mandibular postcanine series. Both postcanine series increased in tooth number as the skull lengthened, with the mandibular postcanine series containing more teeth than the maxillary series. In the maxilla, the first postcanine is consistently the smallest tooth, showing a proportional reduction in size as skull length increased. The longer retention of a tooth in this first locus is a key difference between Galesaurus and Thrinaxodon, in which the mesial-most postcanines are lost after replacement. This difference has contributed to the lengthening of the postcanine series in Galesaurus, as teeth continued to be added to the distal end of the tooth row through ontogeny. Overall, there are considerable differences between Galesaurus and Thrinaxodon relating to the replacement and development of their teeth.Fil: Norton, Luke A.. University of the Witwatersrand; SudáfricaFil: Abdala, Nestor Fernando. University of the Witwatersrand; Sudáfrica. Consejo Nacional de Investigaciones Científicas y Técnicas. Centro Científico Tecnológico - Tucumán. Unidad Ejecutora Lillo; ArgentinaFil: Rubidge, Bruce S.. University of the Witwatersrand; SudáfricaFil: Botha, Jennifer. Free State University; Sudáfrica. Karoo Palaeontology; Sudáfric
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