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FIGURE 9 in Two new species of the marine isopod genus Cirolana Leach, 1818 (Crustacea Isopoda: Cirolanidae) from the coast of the western Gulf of Thailand
No full textFIGURE 9. Cirolana parawongat sp. nov., paratype male (5.0 mm, PSUZC-CR2087-02). A, left mandible; B, distal part of mandible; C, maxillule; D, maxilla;,E, maxilliped; F, maxilliped endite.Published as part of Rodcharoen, Eknarin & Bruce, Niel L., 2021, Two new species of the marine isopod genus Cirolana Leach, 1818 (Crustacea Isopoda: Cirolanidae) from the coast of the western Gulf of Thailand, pp. 469-486 in Zootaxa 4950 (3) on page 482, DOI: 10.11646/zootaxa.4950.3.3, http://zenodo.org/record/464999
FIGURE 9 in Two new species of the genus Kamaka (Crustacea: Amphipoda: Kamakidae) from Songkhla Lagoon, southern Thailand
No full textFIGURE 9. Kamaka songkhlaensis sp. nov. A–H, holotype, male, 2.5 mm (OMNH-Ar-7754); I–K, allotype, female, 2.4 mm (OMNH-Ar-7755); L, paratype, male, 2.2 mm (OMNH-Ar-7756); M, paratype, male, 1.7 mm (OMNH-Ar-7757). A, left epimeral plates 1–3 (lateral view); B, left pleopod 1 (anterior view); C, D, left pleopods 2, 3 (posterior views); E, left uropod 1 (dorsomedial view); F, right uropod 2 (dorsomedial view); G, left uropod 3 (ventral view); H, telson & right uropod 3 (ventrolateral view); I, left antenna 2 (lateral view); J, right gnathopod 1 (lateral view); K, left gnathopod 2 (lateral view); L, M, carpi–dactyli of left gnathopods 2 (lateral views). Scales: 0.05 mm.Published as part of Ariyama, Hiroyuki, Angsupanich, Saowapa & Rodcharoen, Eknarin, 2010, Two new species of the genus Kamaka (Crustacea: Amphipoda: Kamakidae) from Songkhla Lagoon, southern Thailand, pp. 55-68 in Zootaxa 2404 on page 65, DOI: 10.5281/zenodo.19416
Kamaka Dershavin 1923
No full textGenus Kamaka Dershavin, 1923 Kamaka Dershavin, 1923: 188 [type species: K. kuthae Dershavin, 1923 by monotypy]. Diagnosis. Urosomites 1 and 2 coalesced. Ocular lobes strongly produced forward, tip blunt. Antenna 1 without accessory flagellum; antenna 2 longer than antenna 1. Mandibular palp slender; palp of maxilla 1 2 - articulate; maxilliped with outer plate elongate, exceeding apex of palp article 2, palp with 4 articles, article 2 long, article 4 with a nail. Coxae relatively long, weakly overlapping; coxa 1 dilated, produced forward; coxae 5 and 6 lobed; coxa 7 smaller than anterior coxae. Gnathopod 1 of both sexes small, poorly subchelate, carpus longer than propodus; male gnathopod 2 enlarged, propodochelate, carpus short, unlobed, propodus dilated, with false chela or large process on posterior margin, dactylus long; female gnathopod 2 small, slightly larger than gnathopod 1, subchelate. Pereopods 3 and 4 similar, each with slender basis; pereopods 5–7 similar to each other, progressively elongate from 5 to 7, each with expanded basis. Oostegites of female present on pereopods 2–5. Pleonal epimeron 3 not bisinuate. Uropods 1 and 2 biramous, stout, peduncle of uropod 1 with ventrodistal process; uropod 3 small, uniramous, peduncle short, as long as ramus. Telson entire, short.Published as part of Ariyama, Hiroyuki, Angsupanich, Saowapa & Rodcharoen, Eknarin, 2010, Two new species of the genus Kamaka (Crustacea: Amphipoda: Kamakidae) from Songkhla Lagoon, southern Thailand, pp. 55-68 in Zootaxa 2404 on page 56, DOI: 10.5281/zenodo.19416
Cirolana Leach 1818
No full textKey to the species of Cirolana ‘pleonastica- group ’ in the South-East Asia region 1 Body dorsal surfaces without nodules or tubercles on posterior margin of pereonites and pleonites............................................................................................. C. tuberculata (Richardson, 1910) - Body dorsal surfaces with or without nodolsoes on posterior margin of pereonites, but with tubercles on pleonites........ 2 2 Body dorsal surfaces without nodules on posterior margin of pereonites 6 and 7................................... 3 - Body dorsal surfaces with nodules on posterior margin of pereonites 6 or 7....................................... 4 3 Head anteriorly produced and overriding the antennular bases; posterior of pleotelson (in male) with 2 tubercles and scattered nodules................................................................. C. trulla Sidabalok & Bruce, 2018 - Head anteriorly not produced and overriding the antennular bases; pleotelson with a median row of fine setae................................................................................. C. lembeh Sidabalok & Bruce, 2018 4 Uropod not sexually dimorphic, male uropodal exopod dorsal posterior surface without distolateral stiff setae........... 5 - Uropod sexually dimorphic, male uropodal exopod dorsal posterior surface with distolateral stiff setae................. 7 5 Antennula peduncle with 4 unfused articles; lateral margin of uropodal exopod having continuous row of plumose setae..................................................................................... C. khamensis sp. nov. - Antennula peduncle with articles 1 and 2 fused; lateral margin of uropodal exopod having discontinuous of plumose setae..................................................................................................... 6 6 Body dorsal with nodules on posterior of both pereonites 6 and 7; frontal lamina pentagonal, lateral margins anteriorly concave................................................................. C. merlion Sidabalok & Bruce, 2018 - Body dorsal with nodules on posterior of pereonite 7 only; frontal lamina pentagonal, lateral margins anteriorly straight....................................................................... C. phuketensis Rodcharoen et al., 2017 7 Pereonites 2–3 without transverse impressed line; pleonite 3 without median tubercle, pleonite 4 with median tubercle and 4 indistinct sublateral tubercules on each side (set as one row)................................. C. parawongat sp. nov. - Pereonites 2–3 with each a single transverse impressed line; pleonite 3 and 4 with 1 median tubercle (the largest) and 10 sublateral tubercles on each side (set as two row).................................. C. fasfes Sidabalok & Bruce, 2018Published as part of Rodcharoen, Eknarin & Bruce, Niel L., 2021, Two new species of the marine isopod genus Cirolana Leach, 1818 (Crustacea Isopoda: Cirolanidae) from the coast of the western Gulf of Thailand, pp. 469-486 in Zootaxa 4950 (3) on page 485, DOI: 10.11646/zootaxa.4950.3.3, http://zenodo.org/record/464999
Cirolana khamensis Rodcharoen & Bruce 2021, sp. nov.
No full textCirolana khamensis sp. nov. (Figs 2–7) Material examined. Holotype: ♂ (6.5 mm), Koh Kham, Songkhla Province, 6°58’18.84” N, 100°51’25.54”E, 9 July 2011, trapped at 8 m depth, coral reef, coll. E. Rodcharoen (PSUZC-CR2087-01). Paratypes: 9 ♂ (7.9, 8.2, 7.3, 7.7, 8.3, 7.1, 7.6, 8.2, 8.4 mm) [dissected]), 7 ♀ (7.4, 7.8, 8.5, 7.9, 8.3, 8.5, 7.8 mm [dissected]), same data as holotype, (PSUZC-CR2087-02). Description of male. Body dorsal surfaces punctuate, widest at pereonite 5 or pereonite 6, lateral margins subparallel. Rostral point absent. Eyes colour brown in alcohol; eyes separated by about 80% width of head. Pereonites 1–3 each with a single transverse impressed line, pereonites 4–6 each with 2 transverse impressed lines; pereonite 7 with 3 tranverse lines with posterior line forming nodulose ridge. Pereonite 1 and coxae 2–3 each with posteroventral angle rounded; coxae 5–7 with entire oblique carina. Pleon with pleonite 1 largely concealed by pereonite 7; pleonites 3–5 with tubercles; posterolateral angles of pleonite 2 rounded, extending posteriorly to anterior of pleonite 4; pleonite 3 with row of 11–13 tubercles, posterolateral margins extending to but not beyond posterior margin of pleonite 5, narrow rounded; pleonite 4 with prominent median tubercles and 3–4 sublateral tubercles on each side, posterolater margins rounded, clearly extending beyond posterior margin of pleonite 5; pleonite 5 with prominent median tubercles and 2–3 sublateral tubercles on each side, posterolateral angles overlapped by lateral margins of pleonite 4. Pleotelson 0.7 times as long as anterior width, dorsal surface with paired submedian logitudinal carina, each of which has 4 tubercles; lateral margins weakly concave; margins smooth, posterior margin evenly rounded, with 8 robust setae. Antennula peduncle articles 1 and 2 distinct articulated; article 2 0.5 times as long as article1; article 3 and 4 0.9 times as long as combined lengths of articles 1 and 2, article 3 1.8 times as long as wide; flagellum with 9 articles, extending to posterior margin of eye. Antenna peduncle article 4 1.8 times as long as wide, 2.6 times as long as article 3, posterodistal angle with 1 short simple seta; article 5 1 times as long as article 4, 2.5 times as long as wide, anterodistal angle with cluster of 4 setae (2 simple and 2 palmate setae), anterodistal angle with cluster of 2 simple setae and 2 palmate setae; flagellum with 20 articles, extending to middle of pereonite 3. Frontal lamina pentagonal, 2.2 times as longer than greatest width, lateral margins straight, diverging slightly towards anterior, anterior margin acute. Left mandible molar process anterior margin with 15 flat teeth; with proximal cluster of long simple setae; left mandible spine row composed of 8 spines; palp articles 2 with 14 distolateral setae, palp article 3 with 17 biserrate RS. Left maxillula mesial lobe with 3 large and circumplumose RS; lateral lobe with 12 RS. Right maxilla lateral lobe with 5 long setae; middle lobe with 13 long setae; mesial lobe with 3 distal long setae and 12 proximal long setae. Right maxilliped palp article 2 mesial margin with 7 slender setae, lateral margin distally with 1 slender seta; article 3 mesial margin with 14 slender setae, lateral margin with 9 slender setae; article 4 mesial margin with 13 slender setae, lateral margin with 5 slender setae; article 5 distal margin 17 setae, lateral margin with 4 setae; endite with 5 long CPS, and 1 coupling setae (both left and right). Pereopod 1 basis 2.3 times as long as greatest width, infererior distal angle with cluster of 2 acute setae; ischium times 0.6 times as long as basis, inferior margin with 2 setae, superior distal margin with 1 RS; merus inferior margin with 6 molariform RS (set as in row of 4+2), superior distal angle with 4 setae (slender); carpus inferior distal margin with 2 setae (1 RS and 1 slender seta). Propodus 2.0 times as long as wide, inferior margin with 3 RS; dactylus 0.7 times as long as propodus. Pereopod 2 ischium inferior margin with 2 stout, bluntly rounded RS, superior distal margin with 3 RS (2 simple and 1 biserrate); merus inferior margin with 8 stout RS (set in row of 5 molariform and 3 acute), superior distal margin with 4 acute RS; carpus inferodistal angle with 3 RS (plus 1 slender seta); propodus 1.7 times as long as wide; dactylus 0.9 times as long as propodus. Pereopod 3 similar to pereopod 2. Pereopod 4 intermediate in form between pereopod 3 and pereopod 5. Pereopod 6 similar to pereopod 7. Pereopod 7 basis 2.8 times as long as greatest width, superior margin convex, inferior margin with 4 palmate setae; ischium 0.6 times as long as basis. inferior margin with 7 RS (set in groups of 3, 1 and 3), superior distal angle with 6 RS, inferior distal angle with 2 RS; merus 0.7 times as long as ischium, 1.3 times as long as wide, inferior margin with 4 RS (set in groups of 1 and 3), superior distal angle with 13 RS (2 simple and 11 biserrate), inferior distal angle with 6 RS; carpus times 0.9 times as long as ischium, 1.7 times as long as wide, inferior margin with 3 RS, superior distal angle with 13 RS (simple and biserrate), inferior distal angle with 10 RS (6 simple and 4 biserrate); propodus 1.0 time as long as ischium, 4.4 times as long as wide, inferior margin with 4 RS (set in group 2 and 2), superior distal angle with 3 RS, 1 slender seta and 1 palmate seta, inferior distal angle with 2 RS; dactylus 0.4 times as long as propodus. Penial openings two low tubercles, separated by 25% of sternal width. Pleopod 1 exopod 1.4 times as long as wide, lateral margin straight, distally broadly rounded, mesial margin strongly convex, with 30 PMS from distal one-third; endopod 2.2 times as long as wide, distally broadly rounded, lateral margin sinuate, with 10 PMS on distal margin only; peduncle 1.6 times as wide as long, mesial margin with 4 coupling hook. Pleopod 2 exopod with 38 PMS, endopod with 13 PMS; appendix masculina with parallel margins, 1.1 times as long as endopod, distally narrowly rounded. Pleopod 3 exopod with 40 PMS, endopod with 11 PMS. Pleopod 4 exopod with 38 PMS, endopod with 9 PMS. Pleopod 5 exopod with 34 PMS. Pleopods 2–5 peduncle distolateral margin with prominent acute RS. Uropod peduncle ventrolateral margin with 2 RS, lateral margin with 1 mesial short acute RS, posterior lobe about one-half as long as endopod; rami extending beyond pleotelson, marginal setae in single tier, apices broadly round. Endopod apically not bifid; lateral margin distally convex, without prominent excision, with 2 RS, mesial margin strongly convex, with 7 RS. Exopod not extending to end of endopod, 1.5 times as long as greatest width, apically shallowly and equally sub-bifid; lateral margin weakly convex, with 5 RS; mesial margin convex, with 4 RS. Female. Similar to male but on average a little larger (see ‘Size’). Variation. Pleotelson (n=17 [10♂ and 7♀]) with 7–8 RS, with 8 RS (4+4) most frequent (76%). Uropod endopod mesial margin with 6–8 RS, with 7 (76%) and 6 (18%) most frequent and 8 (6%) occurring only once, lateral margin with 2 RS (100%); exopod mesial margin with 4 RS (100%), lateral margin with 5 RS (100%). Size. Adult males (n=10) 6.5–8.3 mm (mean 7.7 mm); females (n=7) 7.4–8.5 mm (mean 8.0 mm). Remarks. Cirolana khamensis sp. nov. is characterized by pereonites 1–3 each with a single transverse impressed line; pereonites 4–6 with each with two transverse impressed lines; pereonite 7 with three transverse lines with the posterior line forming a nodulose ridge; the antennula peduncular articles 1 and 2 are distinct and articulated; pleotelson dorsal surface with two sub-median longitudinal carinae, each of which has four prominent tubercles, posterior margin evenly rounded, with eight robust setae; pereopod 1 merus inferior margin with six molariform RS; lateral margin of uropodal exopod having continuous row of PMS; and penes in the form of two low tubercles separated by 25% of sternal width. We place Cirolana khamensis into the Cirolana ‘pleonastica -group’ as the species shows all the characters for that group, with the exception of the uropodal exopod lateral margin, which has setae along the distal two-thirds of its length, though not the entire length. Nonetheless, the morphology and setation of the uropodal exopod is closer to that of the Cirolana ‘pleonastica -group’ than to the other clades, and furthermore C. khamensis lacks the defining characters of the other groups (see Sidabalok 2019). In their diagnosis of the ‘pleonastica -group’, Sidabalok & Bruce (2018a) state the uropod exopod lateral margin character is usually present. Similar South-East Asian species in the Cirolana ‘pleonastica -group’ are Cirolana phuketensis Rodcharoen et al., 2017 (Thailand), C. merlion Sidabalok & Bruce, 2018 (Singapore), C. fasfes Sidabalok & Bruce 2018 (Indonesia) (Sidabalok & Bruce 2018a). However, the new species can be distinguished from those species by lateral margin of uropodal exopod having a more continuous row of PMS (vs uropodal exopod lateral margin with widelyspaced and acute robust setae). Moreover, C. khamensis sp. nov. can be separated from the allopatric species C. bruscai Bruce & Olesen, 2002 by the head not being produced and overriding the antennular bases; and the inferior margins of pereopods 6 and 7 are smooth (vs head weakly produced and overriding the antennular bases; inferior margins of pereopods 6 and 7 serrate in C. bruscai). Etymology. The epithet is taken from the type locality.Published as part of Rodcharoen, Eknarin & Bruce, Niel L., 2021, Two new species of the marine isopod genus Cirolana Leach, 1818 (Crustacea Isopoda: Cirolanidae) from the coast of the western Gulf of Thailand, pp. 469-486 in Zootaxa 4950 (3) on pages 471-478, DOI: 10.11646/zootaxa.4950.3.3, http://zenodo.org/record/464999
Kamaka appendiculata Ariyama, Angsupanich & Rodcharoen, 2010, sp. nov.
No full textKamaka appendiculata sp. nov. (Figs 2–5) Kamaka sp. 2 Angsupanich et al., 2005: 375. Material examined. Holotype: male, 2.1 mm (OMNH-Ar- 7750), Outer Songkhla Lagoon (Fig. 1 A; 07°08' 47 ''N, 100 ° 33 ' 15 ''E), 1.2 m deep, brown soft clay, 30 psu in salinity, 12 Oct. 2008, coll. S. Angsupanich. Allotype: female, 1.7 mm (OMNH-Ar- 7751), same data as holotype. Other paratypes: 2 males, 1.9, 1.7 mm (OMNH-Ar-7752, 7753); 1 male and 2 females, 2.1, 2.0, 1.8 mm (PSUZC-CR0224 to 0226); 7 males and 4 females, 1.8 –2.0, 1.7–2.3 mm (PSUZC-CR0227, 0228; not dissected), same data as the holotype. Description. Male [based on holotype, 2.1 mm (OMNH-Ar- 7750), and paratype 1.9 mm (OMNH-Ar- 7752) for pereopod 5]. Body (Fig. 2) slender, smooth. Eyes medium-sized. Antenna 1 (Fig. 3 A) with ratio of lengths of peduncular articles 1–3 1: 0.9: 0.7, peduncular article 1 with 1 penicillate and 1 simple setae at ventrodistal corner; flagellum with 5 medium-sized articles and 1 tiny distal article, articles 2–5 each with aesthetasc at ventrodistal corner. Antenna 2 (Fig. 3 B, B 1) with article 2 bearing ovoid lamellate appendix mediodistally, ratio of lengths of peduncular articles 3–5 1: 2.2: 1.7; flagellum with 4 medium-sized articles and 1 tiny distal article, tip of article 4 with several setae. Upper lip (Fig. 3 C) with a hollow on ventral margin, bearing short thin setae. Mandible (Fig. 3 D, D 1, E, E 1) with each incisor bearing 4 cusps; left lacinia mobilis with 5 cusps, right 3 cusps; palp article length ratio 1: 1.6: 1.7, articles 1–3 with 0, 1 and 11 setae, respectively. Lower lip (Fig. 3 F) with mandibular process short, central part of outer lobe with a roundish cavity-like structure, apical parts of inner and outer lobes covered with thin setae. Maxilla 1 (Fig. 3 G, G 1) with outer plate bearing 10 robust setae apically; tip of palp article 2 with 4 robust and 2 thin setae. Maxilla 2 (Fig. 3 H) with both plates setose apically. Maxilliped (Fig. 3 I) with inner plate bearing several thick plumose setae distally; outer plate with 8 medial robust setae. Gnathopod 1 (Fig. 4 A) with coxal plate roundish-trapezoidal, posteroventral corner rounded, bearing 9 fine setae and 1 robust seta on anterior/ventral and posterior margins, respectively; basis with 3 long setae posteriorly; carpus with many thick setae and several normal setae on posterior margin and medial surface, respectively; propodus setose anteriorly and posteriorly; dactylus with denticle on posterior margin. Gnathopod 2 (Fig. 4 B) with coxal plate roundish, bearing 2 ventral setae and 1 posterior robust seta (lost); basis elongate, posterodistal corner with short seta; merus short, posterodistal corner rounded; carpus triangular; propodus enlarged, with long acute process on posteroproximal corner, posterior margin of main part with many setae, posterior margin of process with several setae; dactylus elongate, strongly curved posteriorly, anterior and posterior margins each with several setae. Pereopods 3 and 4 (Fig. 4 C, C 1, D, D 1) with coxal plates roundish, each bearing 4 ventral setae and 1 posterior robust seta; basis of pereopod 3 with 2 long setae; meri with several setae at anterodistal corners; dactyli short. Pereopod 5 (Fig. 4 E) with coxal plate wide, anterior lobe with several setae on ventral margin, posterior lobe with short plumose seta at posteroventral corner; basis slightly expanded, anterior margin with 1 plumose and 3 simple setae, anterodistal corner with 3 simple setae, posterior margin with 1 plumose and 2 simple setae; merus with thick seta at posterodistal corner; carpus with 1 + 3 robust setae; propodus with 1 + 2 robust setae and 1 apical plumose seta. Pereopod 6 (Fig. 4 F) with coxal plate narrow, anterior margin with long robust seta, posteroventral corner with short plumose seta; basis slightly curved posteriorly, anterior margin with 5 plumose and 3 simple setae, anterodistal corner with 2 simple setae, posterior margin with 2 plumose and 3 simple setae; merus with anterodistal robust seta and posterodistal long seta; carpus with 3 posterodistal robust setae; propodus with 1 + 1 + 1 robust setae, distal end with 1 long and 2 short simple setae and 1 plumose seta; dactylus curved. Pleon 2 with 2 plumose setae ventrally (Fig. 5 A). Pleopod 3 shortest (Fig. 5 B–D); peduncles of pleopods each with several plumose setae and 2 coupling hooks; outer rami shorter than inner, outer rami of pleopods 1–3 with 7, 6 and 6 articles and inner rami with 5, 4 and 4 articles, respectively. Uropod 1 (Fig. 5 E) with peduncle bearing 3 robust setae on dorsal surface; outer ramus shorter than inner, outer ramus with 3 marginal and 4 terminal robust setae, inner ramus with 2 marginal and 4 terminal robust setae. Uropod 2 (Fig. 5 F) with peduncle bearing robust seta each on dorsal and medial surface; outer ramus shorter than inner, outer ramus with 3 marginal and 4 terminal robust setae, inner ramus with 2 marginal and 4 terminal robust setae. Uropod 3 (Fig. 5 G) with peduncle bearing 1 fine seta and 1 distal robust seta on medial margin; ramus short, distal end obtuse, with 2 thick and 3 normal setae. Telson (Fig. 5 H) short, not coalesced with urosomite 3; dorsodistal margin with 2 and 1 short setae, lateral margins each with 2 penicillate setae. Female [based on allotype, 1.7 mm (OMNH-Ar- 7751)]. Generally similar to male except for antenna 2, gnathopods and oostegites. Antenna 2 (Fig. 5 I) with more slender peduncle lacking lamellate appendix. Gnathopod 1 (Fig. 5 J) with coxal plate smaller than that of male. Gnathopod 2 (Fig. 5 K) with coxal plate roundish-square, bearing 5 ventral setae and 1 posterior robust seta; basis bearing 4 setae on anterior margin, posterior margin with long seta, posterodistal corner with 3 setae; merus relatively long, distal end truncate; carpus roundish triangular, anterior margin and medial surface weakly setose; propodus setose posteriorly, palm oblique, defined by small robust seta; dactylus slender; oostegite on gnathopod 2 small, with single long seta. Pereopod 7 (Fig. 4 G) with coxal plate weakly lobate, anterior margin with robust seta, posteroventral corner with short plumose seta; basis almost linear, anterodistal corner with simple seta, posterior margin with plumose seta; merus with seta on anterodistal corner; carpus with 2 robust setae distally; propodus with 2 anterodistal robust setae, posterodistal end with 3 long simple setae and 1 plumose seta; dactylus curved. Shape change in male antenna 2 and gnathopod 2 with growth. Paratype, 1.7 mm (OMNH-Ar- 7753): lamellate appendix on peduncular article of antenna 2 small (Fig. 5 L); propodus proper, posterior process and dactylus of gnathopod 2 shorter than those of the holotype (Fig. 5 M). Etymology. Referring to the appendix on peduncular article 2 of male antenna 2. Remarks. The ovoid lamellate appendix on peduncular article 2 of male antenna 2 is a unique character that all other Kamaka species do not possess, although the shape of male gnathopod 2 of this new species resembles that of Kamaka biwae and K. morinoi. Distribution. Outer Songkhla Lagoon (present study) and Middle Songkhla Lagoon (Angsupanich et al., 2005).Published as part of Ariyama, Hiroyuki, Angsupanich, Saowapa & Rodcharoen, Eknarin, 2010, Two new species of the genus Kamaka (Crustacea: Amphipoda: Kamakidae) from Songkhla Lagoon, southern Thailand, pp. 55-68 in Zootaxa 2404 on pages 57-62, DOI: 10.5281/zenodo.19416
Kamaka songkhlaensis Ariyama, Angsupanich & Rodcharoen, 2010, sp. nov.
No full text<i>Kamaka songkhlaensis</i> sp. nov. <p>(Figs 6–9)</p> <p> <i>Kamaka</i> cf. <i>taditadi</i> Angsupanich <i>et al.</i>, 2005: 375. — Ruensirikul <i>et al.</i>, 2007: 1233, fig. 5.</p> <p> Not <i>Kamaka taditadi</i> Thomas & Barnard, 1991: 311, figs 1–4. — Krapp-Schickel & Myers, 2006: 1091, figs 5–7.</p> <p> <b>Material examined.</b> Holotype: male, 2.5 mm (OMNH-Ar-7754), Outer Songkhla Lagoon (Fig. 1 B; 07°11'04''N, 100°25'03''E), 0.35 m deep, fine sand, 21 psu in salinity, 10 Oct. 2008, coll. S. Angsupanich. Allotype: female, 2.4 mm (OMNH-Ar-7755), same data as holotype. Other paratypes: 2 males, 2.2, 1.7 mm (OMNH-Ar-7756, 7757); 2 males and 1 female, 1.9, 1.8, 2.2 mm (PSUZC-CR0229 to 0231); 1 male and 1 female, 1.9, 2.2 mm (PSUZC-CR0232, 0233; not dissected), same data as the holotype.</p> <p> <b>Description.</b> Male [based on holotype, 2.5 mm (OMNH-Ar-7754)]. Body (Fig. 6) slender, smooth. Eyes medium-sized. Antenna 1 (Fig. 7 A) with ratio of lengths of peduncular articles 1–3 1:1.2:0.9, peduncular article 1 with 2 penicillate setae at ventrodistal corner; flagellum with 9 medium-sized articles, articles 7–9 each with aesthetasc at ventrodistal corner. Antenna 2 (Fig. 7 B, B1) with distinct swelled article 4, ratio of lengths of peduncular articles 3–5 1:2.6:1.7; flagellum with 1 long, 4 medium-sized and 1 tiny distal article, article 1 excavate posteriorly, with serrate posterior margins, tip of article 5 with several setae.</p> <p>Upper lip (Fig. 7 C) bearing short thin setae. Mandible (Fig. 7 D, D1, E, E1) with left and right incisors bearing 4 and 5 cusps, respectively; left lacinia mobilis with 4 cusps, right 2 cusps; tips of accessory blades blunt (acute in paratypes); palp with article 3 broadened distally, article length ratio 1:1.4:1.2, articles 1–3 with 1, 6 and ca. 13 setae, respectively. Lower lip (Fig. 7 F) with mandibular process short, central part of outer lobe with a roundish cavity-like structure, apical parts of inner and outer lobes covered with thin setae. Maxilla 1 (Fig. 7 G, G1) with outer plate bearing 10 robust setae apically; tip of palp article 2 with 4 robust, 4 thin and many feeble setae. Maxilla 2 (Fig. 7 H) with both plates wide, setose apically. Maxilliped (Fig. 7 I) with inner plate bearing several thick plumose setae distally; outer plate with 8 medial robust setae.</p> <p>Gnathopod 1 (Fig. 8 A) with coxal plate large, posteroventral corner right-angled, bearing 15 fine setae and 1 robust seta on anterior/ventral and posterior margins, respectively; basis with long seta posteriorly; carpus with many thick setae and normal setae on posterior margin and medial surface, respectively; propodus setose anteriorly and posteriorly; dactylus with denticle on posterior margin. Gnathopod 2 (Fig. 8 B) with coxal plate roundish, bearing 3 ventral setae and 1 posterior robust seta; basis elongate, posterodistal corner with short seta; merus relatively short, posterodistal corner rounded; carpus roundish triangular; propodus broad, with acute process on posteroproximal corner, distal margin rounded, with many setae, posterior margin of process with several setae; dactylus elongate, strongly curved posteriorly, anterior and posterior margins each with a few setae.</p> <p>Pereopods 3 and 4 (Fig. 8 C, C1, D) with coxal plates roundish square, each bearing 4 ventral setae and 1 posterior robust seta; basis of pereopod 3 with long seta; meri with several setae at anterodistal corners; dactyli short. Pereopod 5 (Fig. 8 E) with coxal plate wide, anterior lobe with a few setae ventrally, posterior lobe with short plumose seta at posteroventral corner; basis slightly expanded, anterior margin with 5 simple setae, anterodistal corner with 3 simple setae, posterior margin with 5 plumose and 1 simple setae; merus with thick seta at posterodistal corner; carpus with 1+3 robust setae; propodus with 2 robust and 1 plumose setae distally. Pereopod 6 (Fig. 8 F) about 1.6 times as long as pereopod 5; coxal plate narrow, anterior margin with long robust seta, posteroventral corner with short plumose seta; basis almost linear, anterior margin with 6 plumose and 2 simple setae, anterodistal corner with 1 plumose and 3 simple setae, posterior margin with 5 plumose and 2 simple setae; merus with anterodistal robust seta and posterodistal long seta; carpus with 1+1+4 robust setae; propodus with 1+2 robust setae, distal end with 3 long and 2 short simple setae and 1 plumose seta; dactylus curved. Pereopod 7 (Fig. 8 G) about 1.2 times as long as pereopod 6; coxal plate not lobate, posteroventral corner with short plumose seta; basis almost linear, anterior margin with 5 plumose and 1 simple setae, anterodistal corner with 3 simple setae, posterior margin with 6 plumose and 2 robust setae; merus with 1+1 long robust setae posteriorly; carpus with 1+1+2 robust setae; propodus with 1+2 anterior and 1 posterior robust setae, distal end with 4 long and 2 short simple setae and 1 plumose seta; dactylus curved.</p> <p>Pleon 1 with 1 plumose and 1 simple setae ventrally, pleon 2 with 2 ventral and 1 posteroventral plumose setae (Fig. 9 A). Pleopod 3 shortest (Fig. 9 B–D); peduncles of pleopods each with a few plumose setae and 2 coupling hooks; outer rami shorter than inner, outer rami of pleopods 2, 3 with 8 and 7 articles and inner rami of pleopods 1–3 with 7, 7 and 6 articles, respectively. Uropod 1 (Fig. 9 E) with peduncle bearing 3 robust setae on dorsal surface; outer ramus shorter than inner, both rami each with 1 marginal and 4 terminal robust setae. Uropod 2 (Fig. 9 F) with peduncle bearing 3 dorsal robust setae; outer ramus shorter than inner, both rami each with 1 marginal and 4 terminal robust setae. Uropod 3 (Fig. 9 G, H) with peduncle bearing distal robust seta on medial margin; ramus short, distal end obtuse, with 2 thick and 2–3 normal setae. Telson (Fig. 9 H) short, not coalesced with urosomite 3; dorsodistal margin with a pair of simple and penicillate setae.</p> <p>Female [based on allotype, 2.4 mm (OMNH-Ar-7755)]. Generally similar to male except for antenna 2, gnathopods and oostegites. Antenna 2 (Fig. 9 I) with peduncular article 4 slender, article 1 of flagellum without excavation. Gnathopod 1 (Fig. 9 J) with coxal plate smaller than that of male, posteroventral corner rounded. Gnathopod 2 (Fig. 9 K) with coxal plate roundish-square, bearing several ventral setae and 1 posterior robust seta; basis bearing 4 setae on anterior margin, posterior margin with 2 long setae, posterodistal corner with 3 setae; merus relatively long, distal end truncate; carpus roundish triangular, anterior margin and posterodistal corner setose; propodus setose anteriorly and posteriorly, palm oblique, defined by small robust seta; dactylus relatively stout; oostegite on gnathopod 2 small, with single long seta.</p> <p>Shape change in male gnathopod 2 with growth. Paratype, 2.2 mm (OMNH-Ar-7756): propodus narrower than that of the holotype, distal margin with small spine (Fig. 9 L). Paratype 1.7 mm (OMNH-Ar-7757): propodus narrower than that of OMNH-Ar-7756, posterior process short, distal margin with blunt projection posteriorly, dactylus stout (Fig. 9 M).</p> <p> <b>Etymology.</b> Referring to the type locality.</p> <p> <b>Remarks.</b> This new species is characterized by the swelled peduncular article 4 of antenna 2 and the wide propodus of gnathopod 2 in adult male, and shares these characters with <i>Kamaka poppi</i> and <i>K. taditadi</i>. However, <i>K. poppi</i> has the following characters (Bamber, 2003): (1) peduncular article 4 of male antenna 2 with distal spine (absent in <i>K. songkhlaensis</i>), (2) merus of male gnathopod 2 long (relatively short in <i>K. songkhlaensis</i>), (3) distal margin of propodus of male gnathopod 2 almost straight (convex in <i>K. songkhlaensis</i>), (4) carpi of male pereopods 3, 4 short (relatively long in <i>K. songkhlaensis</i>), (5) inner rami of uropods 1, 2 without marginal spine (present in <i>K. songkhlaensis</i>). On the other hand, <i>K. taditadi</i> has the following characters (Thomas & Barnard, 1991; Krapp-Schickel & Myers, 2006): (1) posteroventral corner of male coxa 1 round (right-angled in <i>K. songkhlaensis</i>), (2) merus and carpus of male gnathopod 2 long (relatively short in <i>K. songkhlaensis</i>), (3) anterior margin of propodus of male gnathopod 2 with blunt triangular process (absent in <i>K. songkhlaensis</i>). In addition, the new species inhabits brackish environment, whereas <i>K. taditadi</i> is a marine species.</p> <p> <b>Distribution.</b> Outer Songkhla Lagoon (present study), Middle Songkhla Lagoon (Angsupanich <i>et al.</i>, 2005) and Upper Songkhla Lagoon (Ruensirikul <i>et al.</i>, 2007).</p>Published as part of <i>Ariyama, Hiroyuki, Angsupanich, Saowapa & Rodcharoen, Eknarin, 2010, Two new species of the genus Kamaka (Crustacea: Amphipoda: Kamakidae) from Songkhla Lagoon, southern Thailand, pp. 55-68 in Zootaxa 2404</i> on pages 62-67, DOI: <a href="http://zenodo.org/record/194167">10.5281/zenodo.194167</a>
Going Beyond Counting First Authors in Author Co-citation Analysis
Get PDFThe present study examines one of the fundamental aspects of author co-citation analysis (ACA) - the way co-citation
counts are defined. Co-citation counting provides the data on which all subsequent statistical analyses and mappings
are based, and we compare ACA results based on two different types of co-citation counting - the traditional type that
only counts the first one among a cited work's authors on the one hand and a non-traditional type that takes into
account the first 5 authors of a cited work on the other hand. Results indicate that the picture produced through this non-traditional author co-citation counting contains more coherent author groups and is therefore considerably clearer. However, this picture represents fewer specialties in the research field being studied than that produced through the traditional first-author co-citation counting when the same number of top-ranked authors is selected and analyzed. Reasons for these effects are discussed
Variations on the Author
Get PDF“Variations on the Author” discusses two of Eduardo Coutinho’s recent films (Um Dia na Vida, from 2010, and Últimas Conversas, posthumously released in 2015) and their contribution to the general question of documentary authorship. The director’s filmography is characterized by a consistent yet self-effacing form of authorial self-inscription: Coutinho often features as an interviewer that rather than express opinions propels discourses; an interviewer that is good at listening. This mode of self-inscription characterizes him as an author who is not expressive but who is nonetheless markedly present on the screen. In Um Dia na Vida, however, Coutinho is completely absent form the image, while Últimas Conversas, on the contrary, includes a confessional prologue that moves the director from the margins to the center of his films. This article examines the ways in which these works stand out in the filmography of a director who offers new insights into the notion of cinematic authorship
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