25,617 research outputs found
Ranatra rafflesi Tran & Polhemus 2012
<i>Ranatra rafflesi</i> Tran & Polhemus, 2012 <p> <i>Ranatra rafflesi</i> Tran & Polhemus, 2012: 104 –106, Figs. 7, 8, 10, 12–15.</p> <p> <b>Material examined.</b> MALAYSIA: 2 males, 3 females, Sarawak, Sg. Stunggang before Bg. Kayan, coll. HH Tan, 2 Sep.1996, THH9693 (ZRC); 2 males, 1 female, Sarawak, Sg Mautala, coll. M. Kottelat <i>et al.</i>, May 1994, MK94-20 (ZRC.6.17663); 1 male, 3 females, Sarawak, Sibu, coll. M. Kottelat <i>et al.</i>, 15 May 1994, MK94-24 (ZRC); 1 male, Sarawak, Serian, Sg. Kubas, coll. HH Tan, 10 Jun. 1999, THH9938 (ZRC); 1 male, Sarawak, 38km towards Simunjan fr. Kuching – Sri Anam Rd, coll. HH Tan, 4 Sep. 1996, THH96100 (ZRC); 1 male, 1 female, Sarawak, 43km to Sri Aman, nr. Batang Ai turnoff, coll. HH Tan, 3 Sep. 1996, THH9698 (ZRC); 1 male, 1 female, Sarawak, Sibu, Sg. Teku, coll. HH Tan & R Kerte, 3 Mar. 1998, THH9810 (ZRC). INDONESIA: 5 males, 3 females, Sumatra, Jambi, Danau Rasau, coll. M Kottelat & HH Tan, 1–2 Jun. 1994, MK94-39 (ZRC).</p> <p> <b>Remarks.</b> First record for Borneo (Sarawak) and Sumatra.</p>Published as part of <i>Tran, A. D. & Poggi, R., 2019, The genus Ranatra Fabricius (Heteroptera: Nepidae) in Borneo, with a redescription of Ranatra spinifrons Montandon and the description of a new species, pp. 236-246 in Zootaxa 4555 (2)</i> on page 244, DOI: 10.11646/zootaxa.4555.2.4, <a href="http://zenodo.org/record/2624329">http://zenodo.org/record/2624329</a>
Rhyacobates anderseni Tran & Yang 2006
Rhyacobates anderseni Tran & Yang, 2006 Rhyacobates anderseni Tran & Yang, 2006: 14 –16, Figs. 7 –16, 27 (type locality: Vu Quang, Ha Tinh Prov., Vietnam). Material examined. For holotype and paratypes, see Tran & Yang (2006). Size. Males, length 6.0– 6.1 (allotype 6.0), width 1.83–1.85 (allotype 1.83) (apterous), length 6.1–6.2, width 1.85–1.88 (macropterous); females, length 6.8 –7.0 (holotype 6.8), width 2.52–2.57 (holotype 2.52) (apterous). Remarks. The followings are diagnostic characteristics of R. anderseni: the mesonotum of the male has a slender yellow stripe on the posterior half; the mesonotum of the female has a broader yellow median marking on the posterior three-fifths; the abdomen of the male is relatively short; in lateral view, abdominal segment 8 of the male has a concave ventral surface; the male proctiger has round angular projections on each side (see Tran & Yang 2006: Fig. 12); the male paramere is falciform, slightly broad, long, and not conspicuously setose (see Tran & Yang 2006: Figs. 13, 14); the metanotum of the female has a pointed median process on the posterior margin (see Tran & Yang 2006: Fig. 7); the abdomen of the female is short (length about 0.2 times body length), sternum 7 is long, almost enclosing the genital segments with its connexival margin raised slightly upwards, with a pair of long posterior projections pointing outwards and downwards (see Tran & Yang 2006: Figs. 8, 9), and its posterior margin is almost straight, bearing two short lateral processes (see Tran & Yang 2006: Fig. 10). For detailed comparisons of Rhyacobates anderseni with its congeners and other ptilomerine genera (Andersenius and Pleciobates), refer to Tran & Yang (2006: 16). Habitats. See Tran & Yang (2006: 16). Distribution. Vietnam: Ha Tinh. China: Yunnan.Published as part of Tran, A. D. & Nguyen, X. Q., 2016, Three new species of the water strider genus Rhyacobates Esaki, 1923 (Hemiptera: Gerridae) from Vietnam, pp. 501-516 in Zootaxa 4121 (5) on pages 512-513, DOI: 10.11646/zootaxa.4121.5.1, http://zenodo.org/record/27168
Ranatra bilobata Tran & Nguyen 2016
Ranatra bilobata group Diagnosis. Body length: males 34–38, females 32–40; siphon long, ratio of siphon length: body length ca. 0.9–1.1; vertex in lateral view slightly higher than eye, evenly round, tubercle small or absent; process of second antennal segment very long, about equal to third segment; flexor side of fore femur with only one tooth at midlength, without carina, pre-apical teeth absent or only with tooth-like elevation (R. bilobata); metasternum with posterior margin deeply emarginated, usually with a pair of sublateral tumescences (except R. sulawesii); operculum of female not reaching apex of connexivum; male paramere with finger-like pre-apical process before hook, apical hook with pointed tip. Species included. Ranatra sulawesii Nieser & Chen, 1991, R. sterea Chen, Nieser & Ho, 2004, and R. bilobata Tran & Nguyen, 2016. Note. For comparison between these three species, see Tran & Nguyen (2016).Published as part of Tran, A. D. & Zettel, H., 2021, Taxonomic review of the Ranatra gracilis group sensu Lansbury, 1972 (Nepomorpha: Nepidae), with descriptions of four new species, pp. 45-70 in Raffles Bulletin of Zoology 69 on page 47, DOI: 10.26107/RBZ-2021-0005, http://zenodo.org/record/535167
Rhyacobates gongvo Tran & Yang 2006
Rhyacobates gongvo Tran & Yang, 2006 (Figs. 41, 42) Rhyacobates gongvo Tran & Yang, 2006: 16 –19, Figs. 17 –25, 28 (type locality: Sa Pa, Lao Cai Prov., Vietnam). Material examined. For holotype and paratypes, see Tran & Yang (2006). Others: VIETNAM: Lao Cai Prov.: 1 female (apterous), Sa Pa, Nam Sai, Seo Nam Sai stream 1, 22° 15.761 ’N 103 ° 55.909 ’E, 844 m asl., coll. Dinh N.H. et al., 24 October 2012, DNH 12.09 (ZMHU); 1 female (apterous), Sa Pa, Nam Sai, Seo Nam Sai stream 2, 22° 14.67 ’N 103 ° 59.541 ’E, 469 m asl., coll. Dinh N.H. et al., 24 October 2012, DNH 12.10 (ZNHU); 6 males, 5 females (apterous), 2 males (macropterous, de-alated), Sa Pa, Ban Ho, Ban Den, Nam Pu stream (feeder stream of Muong Hoa stream), 22 ° 15.709 ’N 103 ° 58.054 ’E, 416 m asl., coll. Tran A.D. et al., 29 May 2013, TAD 1316 (ZMHU); 1 male, 2 females (apterous), Sa Pa, Thanh Phu, Nam Cang stream, 22 ° 15.401 ’N 103 ° 58.866 ’E, 398 m asl., coll. Tran A.D. et al., 26 October 2013, TAD 1359 (ZMHU); 13 males, 4 females (apterous), Sa Pa, Ban Ho, Nam Pu stream (feeder stream of Muong Hoa stream), site 1, at lower section, 22 ° 15.778 ’N 103 ° 58.270 ’E, 404 m asl., coll. Tran A.D. et al., 26 October 2013, TAD 1361 (ZMHU); 1 female (apterous), Sa Pa, Cat Cat, Ho stream (feeder stream of Muong Hoa stream), 22 ° 19.546 ’N 103 ° 49.880 ’E, 1233 m asl., coll. Tran A.D. et al., 27 October 2013, TAD 1366 (ZMHU). Size. Males, length 6.2–6.5 (allotype 6.5), width 1.88–2.20 (apterous), length 6.4, width 1.97 (macropterous, de-alated); females, length 7.8–8.3 (holotype 8.3), width 2.52–2.67 (holotype 2.52) (apterous), length 7.5, width 2.44 (macropterous, de-alated). Remarks. Rhyacobates gongvo differs from other species of Rhyacobates by the following diagnostic characteristics: in the apterous morph, the mesonotum has a median yellow stripe on the posterior three quarters; the male proctiger has small angular projections on each side (see Tran & Yang 2006: Fig. 22); the male paramere is relatively long and slender, not setose (see Tran & Yang 2006: Figs. 24, 25); the abdomen of the female is elongate and straight (length about 0.4 times body length), the posterior part of sternum 7 is slightly depressed dorsoventrally (see Tran & Yang 2006: Fig. 17); sternum 7 of the female does not totally enclose the genital segments, the posterior margin is straight and without a process, and the connexival projections are long, straight, and flat (see Tran & Yang 2006: Figs. 18–20). Rhyacobates gongvo is relatively similar to R. malaisei Andersen & Chen, 1995, but can be separated from the latter by the diagnosis above (for a comparison between these two species, see Tran & Yang 2006: 18–19). Habitats. See Fig. 40; also see Tran & Yang (2006: 18). Distribution. Vietnam: Lao Cai.Published as part of Tran, A. D. & Nguyen, X. Q., 2016, Three new species of the water strider genus Rhyacobates Esaki, 1923 (Hemiptera: Gerridae) from Vietnam, pp. 501-516 in Zootaxa 4121 (5) on page 513, DOI: 10.11646/zootaxa.4121.5.1, http://zenodo.org/record/27168
Phuc Tran: Author, Teacher, Tattoo Artist
Profile of Phuc Tran. Tran taught classical languages in middle and high schools in Maine and New York for years. He also owns a tattoo shop in Portland and has written a highly acclaimed novel titled Sigh, Gone about his family\u27s move to Pennsylvania from Vietnam in 1975. In this piece, Tran briefly discusses community, language, and how it feels being an Asian American in Maine
TRAN Quyet Thang
학위논문(박사)--아주대학교 일반대학원 :전자공학과,2017. 2CHAPTER 1: INTRODUCTION 1
1.1 Introduction to Light Modulating Devices 1
1.2 Introduction to Graphene 1
1.3 Overview of the Numerical Methods 3
1.4 Organization of Dissertation 7
CHAPTER 2: TUNABLE WIDE-ANGLE TUNNELING IN GRAPHENE-ASSISTED FRUSTRATED TOTAL INTERNAL REFLECTION 9
2.1 Introduction 9
2.2 Frustrated Total Internal Reflection (FTIR) Effect 10
2.3 Controllable Wide Angle Tunneling Effect with ENZ Effect in FTIR Configuration 11
2.4 Waveguide-type optical modulator based on a GA-FTIR structure 20
2.5 Chapter Summary 26
CHAPTER 3: LOW LOSS ELECTRICALLY CONTROLLABLE VERTICALLY COUPLED DIRECTIONAL COUPLER 28
3.1 Introduction 28
3.2 Vertically Coupled Directional Coupler 29
3.3 Chapter Summary 40
CHAPTER 4: OPTICAL PHASE MODULATOR BASED ON GRAPHENE EMBEDDED ALL PASS FILTER 42
4.1 Introduction 42
4.2 Phase Modulating All Pass Filter with Graphene 42
4.3 Chapter Summary 48
CHAPTER 5: COUPLED MODE THEORY OF PERFECT GRAPHENE ABSORBERS IN DUAL-MODE/SINGLE-MODE COUPLED RESONATORS SYSTEM 50
5.1 Introduction 50
5.2 Dual Mode - Single Mode Coupled Resonators System 51
5.3 Temporal Coupled Mode Theory of the "Triple-mode absorber" 52
5.4 Numerical verification of the CMT 59
5.5 Chapter Summary 66
CHAPTER 6: CONCLUSION AND FUTURE WORK 68
6.1 Conclusion 68
6.2 Future Work 68
REFERENCES 70
APPENDIX: AUTHOR'S PUBLICATIONS LIST 79DoctoralOne of the most important component of integrated photonics is a class of devices known as light modulation devices, which allow us to modulate and manipulate the flow of light, similar to the role transistor played in electronics. Only recently introduced, but graphene have shown incredible promises as a "miracle" material in electronics, with properties ranging from zero band gap, very high electrical mobility, ultra broadband optical responses, and the ability to drastically modify its optical properties through chemical or electrical doping.
In this dissertation, the author presented several unique nanostructures to exploit the aforementioned graphene characteristics to create light modulation devices with superior performance characteristics. Novel effects including wide angle extraordinary reflection causes by epsilon near zero effect and wide angle extraordinary transmission causes by coupling of plasmonic supermodes, phase modulation with near unity amplitude transmission with all pass filter, and graphene perfect absorber with a coupled system of dual mode/single mode resonator was thoroughly investigated, theoretically and numerically. The effects was also presented in practical nanostructures better suited for applications, which are also numerically investigated with various numerical methods
Mon Tran
Photograph used for a story in the Daily Oklahoman newspaper. Caption: "Mon Tran of Oklahoma City has r eceived a FEL-PRO Automotive Scholarship.
Monologue, Dialogue, and Tran Vietnam
A manuscript comprised of materials completed by O. W. Wolters before his death.O. W. Wolters was a twentieth-century historian of early Southeast Asia
who began his academic career with the study of early commercial
relations in the Malay world and the maritime empire of Srivijaya, which
dominated the Straits of Malacca and neighboring seas for several
centuries. During the last twenty-five years of his life, he became
interested in the Tran dynasty of Vietnam (1225-1400). From 1976 to 1996,
he published twelve articles about the Tran dynasty. When he died in 2000,
he left a nearly-completed manuscript of a book-length work about that
dynasty, which is herewith made available to the world of readers.
What makes this manuscript particularly interesting is how the author
shaped a work of historical research into what he liked to call a novel.
He became convinced that there was a certain way of thinking and speaking
that was distinctive to educated people in the Tran period, and he
believed that the best way to present this was through conversational
dialogue. He further presents the Tran way of thinking as a critical
perspective on the regimes that followed.
This manuscript also contains self-reflexive meditations on what the
author was endeavoring to achieve and his critique of his success in doing
so. He offers readers a rare glimpse into the craftsmanship of the most
creative and adventurous scholar of early Southeast Asia in his
generation
Phallostethus cuulong Shibukawa, Tran & Tran, 2012, new species
Phallostethus cuulong, new species New Vietnamese name: Cá bụng đầu (Figures 1–4) Holotype. ZRC 53233, male, 24.2 mm SL, a branch of Hau River (a distributary of Mekong), Cu Lao Dung, Soc Trang Province, Vietnam (9 ° 30.8 ’ N, 106 ° 13.7 ’ E), 0–0.5 m depth, 31 July 2009, collected by K. Shibukawa. Paratypes. Total eight specimens (five males and three females), 20.0– 24.5 mm SL: CTU-P 2327, 1 specimen (female), 23.7 mm SL, Duyen Hai, Tra Vinh Province, Vietnam (9 ° 40.9 ’ N, 106 ° 30.7 ’ E), 0.3–0.8 m depths, 5 April 2009, collected by K. Shibukawa, V.V. Tran and L.X. Tran; CTU-P 2494, 1 specimen (male), 22.5 mm SL, My Thanh River (a distributary of Mekong), Vinh Chau, Soc Trang Province, Vietnam (9 ° 22.7 ’ N, 106 ° 0.7 ’ E), 0.5–1.2 m depths, 1 August 2009, collected by H.P. Ha, V.V. Tran and L.X. Tran; CTU-P 5020, 1 specimen (male, cleared and stained), 23.5 mm SL, Cho Lach, Ben Tre, Vietnam (10 ° 10.5 ’ N, 106 °. 8.9 ’ E), 0.5 m depth, 3 February 2010, collected by L.X. Tran; NSMT-P 106664, 1 specimen (male), 20.0 mm SL, collected with CTU-P 2494; NSMT-P 106665, 1 specimen (female), 22.0 mm SL, collected with CTU-P 2494; USNM 404477, 1 specimen (female), 23.8 mm SL, collected with CTU-P 2494; USNM 404478, 1 specimen (male), 20.3 mm SL, Cau Ke, Tra Vinh Province, Vietnam (9 ° 57.1 ’ N, 106 ° 1.8 ’ E), 0.5–3.5 m depths, 28 May 2010, collected by L.X. Tran; USNM 404479, 1 specimen (male, cleared and stained), 24.5 mm SL, collected with CTU-P 5020. Diagnosis. Phallostethus cuulong is distinguished from congeners in having following characters: seven serrae on the second ctenactinium in adult males (vs. five and eight in P. dunckeri and P. l e h i, respectively); 11–13 pectoral-fin rays (vs. 9–10 and 12 in P. dunckeri and P. l e h i, respectively); 11–14 + 25-26 = 37–40 vertebrae (vs. 13 + 27 = 40 and 12 + 28 = 40 in P. dunckeri and P. l e h i); approximately 5–19 teeth on paradentary (vs. 15–20 and 28 or more in P. dunckeri and P. le h i, respectively). All six examined males are dextral (vs. one and two known males of P. dunckeri are sinistral and dextral respectively, and all four know males of P. l e h i are sinistral). Description. Counts of the holotype are asterisked, and the frequency of each count is given in parentheses following the relevant count. Dorsal-fin rays 7 (2), 8 * (6) or 9 (1); anal-fin rays 24 * (4), 25 (1), 26 (3) or 27 (1); pectoral-fin rays 11 (2), 12 * (7) or 13 * (9); scales in lateral series 34 (5), 35 * (10) or 36 (3); predorsal scales 1 + 25 (1), 2 + 25 (1), 2 + 26 (4) or 2 + 27 * (3); transverse scales 6 (3), 7 * (12) or 8 * (3); circumpeduncular scales 12 * (8) or 13 (1); paradentary teeth approximately 5 (1), 6 (3), 7 (3), 8 (3), 10 (1), 13 * (1), 14 (2), 15 * (1), 18 (2) or 19 (1). The following measurements are % of SL: head length 22.1–24.1; snout length 7.0– 8.5; eye diameter 6.7–7.3; interorbital width 3.3–5.2; length of jaw 8.0– 9.4; predorsal length 78.0– 82.6; preanal length 46.4–48.7; maximum body depth 15.0– 18.7; body depth at anal-fin origin 12.4 –15.0; body width 9.4 –14.0; caudal-peduncle length 18.6–20.7; caudal-peduncle depth 5.6–7.6; length of dorsal-fin base 9.1–10.5; length of anal-fin base 32.3–36.7; pectoral-fin length 14.6–19.1; caudal-fin length 20.1–22.4. Head depressed anteriorly, with flat or barely concave interorbital space. Dorsal surface of head with membranous dome when alive or freshly collected (which can be seen in the cleared and stained specimens, e.g., Fig. 4), but shrunken and not apparent in alcohol specimen. Snout rather pointed. Eyes lateral on head, large, diameter slightly less than snout length. Mouth terminal or subterminal; jaws small, barely extend to a level of anterior margin of eye; upper jaws highly protractile. Body compressed, moderately deep. Anus and urogenital openings anterior, ventral to pectoral-fin base. A slightly frayed, fleshy membranous mid-ventral keel between urogenital opening and anal-fin origin. In males, a distinct mid-ventral groove, deepened and widened anteriorly, supports the priapium and mid-ventral keel. Pectoral fin falcate, the uppermost branched ray longest in most specimens; pectoral-fin rays branched, except for the uppermost 1 (uppermost nubby ossicle not included here; see “Materials and Methods” above) and lowermost 1–2 rays unbranched. Pelvic fin absent in males, present but rudimentary in females (Fig. 3). First dorsal fin absent; origin of second dorsal fin at, or slightly before, a level of posterior end of anal-fin base; anterior 2–3 rays simple, whereas the other rays branched. Anal fin with a long base, commencing well before midlength; anterior 2–4 rays simple, whereas the other rays branched. Caudal fin emarginate, symmetrical dorsoventrally. Male bilaterally asymmetric, dextral; namely, seminal papilla offset to right side of body (= aproctal side), and anus offset to left side of body (= proctal side); a long rod-like toxactinium curved from left to right; a large fleshy pad, the pulvinulus, covers articulation point of toxactinium and aproctal axial bone (Fig. 3). Scales on body cycloid, moderately large and deciduous; scales on abdomen largest; body entirely scaled, except for pectoral-fin base, mid-ventral groove before anal-fin origin, and mid-predorsal narrow naked space slightly behind occipital region; head and fins naked, except for posterior part of occipital region and basal part of caudal fin with some scales. Teeth on premaxilla and dentary unicuspid, slightly curved inward. Paradentary slender (as in Phallostethus dunckeri illustrated by Parenti, 1984: 4, fig. 1), with 5–19 minute teeth laterally; teeth on paradentary form a uniserial row or, in some larger specimens, biserial rows; teeth on inner row, if present, much smaller than those on outer row. Cephalic sensory canals reduced, comprising: two short infraorbital canals (each with terminal pores only) anteriorly and anteroventrally to eye; preopercular canal (with 6–7 pores). Main external bones in males including a long, curved toxactinium and a short stout ctenactinium with seven serrae dorsally (not including a hook-like distal tip); two smallest males examined (CTU-P 2493 and USNM 404478, 20.0– 20.3 mm SL) bearing 5–6 serrae on ctenactinium, assumued to represent the immature condition (and not included in the diagnosis, above). First pleural rib attached to fifth vertebra in males, fourth in females; first pleural rib in female much shorter than in male. Branchiostegal rays 4. Color when alive or freshly collected. Body subtranslucent in life, but whitish immediately after death (Fig. 1); a bright white blotch over brain when alive (assumed to fade just after death); iris silvery; minute melanophores scattered on snout, cheek and jaws; a melanophore at angle of lower jaw; a large reddish yellow blotch, slightly smaller than eye, at mid-lateral caudal fin base; male priapium with several large and small melanophores, particularly on the aproctal side (= right side in the new species) just anterior to the base of second ctenactinium; a series of minute black dots along mid-lateral septum of body musculature at least on caudal part of body; inner side of pectoral fin with many melanophores at least dorsally (the area with melanophores much more broader in females than in males); a series of mid-ventral black dots from anal-fin origin to caudal-fin base; other fins transparent. Color in alcohol. Head and body pale straw-colored; a series of irregular-sized melanophores (several of them dash-like) along midlateral septum of body musculature (at least on caudal part); paired patches of melanophores on anterior part of snout dorsally; many melanophores scattered on head above neurocranium, those posterior distinctly larger than those anterior; a melanophore at angle of lower jaw; a patch of minute gular melanophores; two patches of melanophores at throat and just behind urogenital opening in females; male priapium with several large and small melanophores, particularly on the aproctal side (= right side in this species) just anterior to the base of second ctenactinium; a mid-ventral series of black dots from anal- to caudal-fin bases, each along anal-fin base on interspace between fin rays (continuous and forming a irregular blackish gray line in some specimens); inner side of pectoral fin with many melanophores dorsally (the area with melanophores much broader in females than in males); caudal fin covered by numerous minute melanophores; other fins transparent. Distribution, habitat and the other notes. Phallostethus cuulong is known from nine specimens, six males and three females, collected from shallow waters (<1.2m depth) around banks of slow-flowing turbid canals and rivers with soft muddy bottoms in Soc Trang and Tra Vinh Provinces, Vietnam. The first author (KS) observed that a fish, latter designated as one of the paratypes of Phallostethus cuulong (NSMT-P 106665), swam slowly at the water surface around a bank of the slow-flowing tidal canal with dense semi-aquatic vegetation. A bright white blotch on dorsal surface of head was clearly confirmed in the field, but less vivid than that of the sympatric aplocheilid, Aplocheilus panchax (well-known for its reflective “pineal” spot on the top of the head). When the fish was disturbed, it quickly swam a short distance away from the original position; it was subsequently scooped up carefully using a hand net by KS. The species was usually solitary, and collected by hand nets or seine nets. Like the other phallostethids in the Vietnamese Mekong, this species has never been seen in the fish markets. As far as we aware, all fishes of the family Phallostethidae have no vernacular names in the Vietnamese Mekong (except for the new species herein named), since they are usually overlooked. Etymology. The specific name, cuulong, is the Vietnamese name of the Mekong delta (Cưu Long), where the type series of the new species was collected. The name, here applied as a noun in apposition, means “nine dragons,” in reference to nine distributaries of the Mekong basin in Vietnam. Remarks. Following the key to genera of phallostethid fishes by Parenti (1989), the new species is clearly assigned to Phallostethus by having the combination of, e.g., shield-like pulvinulus, large seminal papilla, long toxactinium, membranous dome on dorsal surface of head, 24–27 anal-fin rays, 37–40 vertebrae, serrated ctenactinium, non-projecting lower jaw beyond upper jaw, no first dorsal fin, and 7–9 second dorsal-fin rays. In particular, no other phallostethid genera are known that bear 24 or more anal-fin rays (vs. 22 or less anal-fin rays in the other phallostethids). Within the genus, the new species resembles the Bornean species Phallostethus lehi in sharing 11–13 pectoral-fin rays, but differs in having seven serrae on second ctenactinium in adult (vs. eight in P. l e h i), 25–26 caudal vertebrae (vs. 28), and 6–19 paradentary teeth (vs. 28 or more). All six examined males of the new species are dextral, immediately distinguishing them from sinistral males in P. l e h i. The new species is also distinguished from Phallostethus dunckeri, known only from Johor, Malay Peninsula but presumed to be extinct (Parenti, 1996), by having seven serrae on second ctenactinium in adults (vs. five in P. d u n c k e r i), 11–13 pectoral-fin rays (vs. 9–10), and 25–26 caudal vertebrae (vs. 27). Sexual dimorphism in the pleural ribs was reported from Phallostethus lehi by Parenti (1996); according to her, the species has the first pair of pleural ribs on the fifth vertebrae in males, the fourth vertebrae in females. This dimorphism is also found in Phallostethus cuulong. Furthermore, P. cuulong appears to show sexual dimorphism in the number of precaudal vertebrae: all six males examined have 13–14 precaudal vertebrae, as against 11–12 in all three females examined. Although Parenti & Louie (1998) reported similar sexual dimorphism in vertebral counts from four species of Neostethus, hitherto it has never been known from the other species of Phallostethus.Published as part of Shibukawa, Koichi, Tran, Dinh Dac & Tran, Loi Xuan, 2012, Phallostethus cuulong, a new species of priapiumfish (Actinopterygii: Atheriniformes: Phallostethidae) from the Vietnamese Mekong, pp. 45-51 in Zootaxa 3363 on pages 46-51, DOI: 10.5281/zenodo.28164
Ranatra heoki Tran & Poggi 2019, sp.n.
<i>Ranatra heoki</i> sp.n. <p>(Figs. 17–27)</p> <p> <b>Material examined.</b> HOLOTYPE: male, Malaysia, Sarawak, 38 km towards Simunjan fr. Kuching – Sri Anam Rd, coll. HH Tan, 4 Sep. 1996, THH96100 (ZRC). Paratype: 1 male, same locality data as holotype (ZRC).</p> <p> <b>Description.</b> General colouration: mostly brown; eyes dark brown; vertex yellowish brown; all coxae brown and mottled with yellowish brown; fore femur, fore tibia, fore tarsus, middle and hind femora yellowish brown and mottled with brown; middle and hind tibiae and tarsi annulated brown and yellow; ventral surface brown with sparse small whitish mottles.</p> <p>Measurements: Males: body length 25.5–26.0 (holotype: 25.5); length of siphon 14.0–14.5 (holotype: 14.0); width of head 2.50–2.53 (holotype: 2.53); width of eye 0.84–0.86 (holotype: 0.84); interocular width 0.78–0.84 (holotype: 0.84); anterior width of pronotum 1.94; humeral width of pronotum 2.27; length of anterior lobe of pronotum along midline 5.00; length of posterior lobe along midline 2.00; fore leg: lengths of coxa 4.50, femur 7.45, tibia 3.00, tarsi 0.72; middle leg: lengths of femur 10.50, tibia 9.40, tarsi 1.75; hind leg: lengths of femur 10.80, tibia 13.25, tarsi 1.90.</p> <p>Head (Figs. 17, 18): Vertex raised above eyes into conical tubercle, the height of the tubercle about a half the height of eye on lateral view; width of eyes about same as interocular width; clypeus, higher than and surpassing lora, anteriorly with a low, conical tubercle; lora bearing long, pale setae along dorsal side, similar setae sparsely on vertex. Antenna: second segment with long, finger-like projection; finger-like projection about three quarters length of third segment.</p> <p>Thorax: Prothorax in lateral view distinctly longer than fore coxa (about 1.8× the length of fore coxa) and slightly longer than fore femur; anterior lobe 2.2–2.3× as long as posterior lobe when measured along midline; anterior margin of pronotum with a tubercle on each side of midline, distinctly raised when viewed laterally; ratio of humeral width / anterior width about 1.17; posterior lobe with humeri broadly rounded (Fig. 18). Scutellum with length 1.59–1.68× width, posterior section with narrowly rounded apex. Prosternum with low median carina on anterior part, anterior margin straight. Mesosternum with a pair of low tubercles on anterolateral margin, posterior projection between middle coxae truncate, weakly grooved along midline. Metasternum with anterior part weakly grooved along midline, posterior part slightly raised medially and grooved laterally, posterior margin deeply emarginated (Fig. 19). Space between middle coxae narrower than that between hind coxae. Hemelytra uniformly textured, mostly brown, with membrane only reaching mid-length of abdominal tergum VI.</p> <p>Legs: Fore femur (Figs. 20–23): slender, widest at basal part, ratio of maximum width at basal part / maximum width at distal part: 1.15–1.17 (holotype: 1.17); ventral margin with a long median carina bearing dense short setae and a tooth on mesal surface situated distally to median carina; distal part with a very small black tooth on lateral surface of ventral margin, before pre-apical constriction on ventral margin, distal tooth shorter than setae on ventral side of femur; ratio of width of femur across median tooth (excluding tuft of setae) / width of femur at basal half: 1.00–1.06 (holotype: 1.06); ratio of width of femur across median tooth / width at base of tooth at distal side: 1.43– 1.55 (holotype: 1.43); ratio of width of femur across median tooth / width of femur across median carina (excluding setae, on proximal side of median tooth): 1.15–1.25 (holotype: 1.25). Middle femur subequal in length to hind femur; hind femur not reaching posterior margin of abdominal sternum VI when folded back parallel to body. Middle tibia shorter than middle femur; hind tibia longer than hind femur; middle and hind femora both bearing sparsely distributed long, thin, pale hairs along their lengths; middle and hind tibiae both bearing dense fringe of long hairs on posterior margins along their entire lengths.</p> <p>Abdomen: Operculum of male (Fig. 24) about as long as connexivum, medially keeled, apex pointed. Respiratory siphon with only sparse long, thin hairs along its length, not formed into a fringe.</p> <p>Male genitalia: Paramere (Figs. 26, 27): dorsoventrally thick, dorsal surface almost straight; strongly constricted at distal quarter before a broadly curved apical hook, inner side of the curve with short setae; apex of hook rounded; ventral side of paramere before apical hook bearing a sub-triangular elevation, with long setae. Phallotheca moderately sclerotised, as in Fig. 25.</p> <p>Female: Unknown.</p> <p> <b>Etymology.</b> The new species is dedicated to Dr. Tan Heok Hui (Lee Kong Chian Natural History Museum), for his significant contribution to the water bug collection in the LKCNHM, including numerous samples of <i>Ranatra</i>.</p> <p> <b>Discussion.</b> <i>Ranatra heoki</i> <b>sp.n.</b> also belongs to the <i>R. gracilis</i> species group (<i>sensu</i> Lansbury, 1972). It can be easily distinguished from other taxa of this species group by the following characteristics: (a) the height of the tubercle on the vertex is about a half the height of an eye in lateral view; (b) the clypeus has a small, but distinct tubercle above the lora (Fig. 17); and (c) the paramere is dorsoventrally thick, with the dorsal surface almost straight, and strongly constricted at the distal quarter before a broadly curved apical hook, and its subapical process is sub-triangular, bearing long setae (Figs. 26, 27).</p> <p> Among taxa of the <i>R. gracilis</i> group, the general appearance of the paramere of this new species is most similar to that of <i>R. schuhi</i> from Myanmar. However, it can be easily distinguished from <i>R. schuhi</i> by the following characteristics: <i>Ranatra heoki</i> <b>sp.n.</b> has a shorter body, with length excluding siphon ca. 25 mm (in <i>R. schuhi</i>, body length is ca. 32 mm). The clypeus of the new species has a small tubercle above the lora (in <i>R. schuhi</i>, the clypeus above the lora is flat and without a tubercle). The ratio of length of the anterior pronotal lobe to length of the posterior protonal lobe is greater in <i>R. heoki</i> <b>sp.n.</b> (2.2–2.3: 1.0) than in <i>R. schuhi</i> (ca. 1.9: 1.0). In the new species, the pair of tubercles at the anterior margin of the mesosternum is less prominent than that in <i>R. schuhi</i>. The hemelytra of <i>R. heoki</i> <b>sp.n.</b> is uniformly textured, mostly brown and shorter, and the membrane reaches only midlength of abdominal tergum VI (hemelytra of <i>R. schuhi</i> is reddish brown with yellow maculated marks and longer, and the membrane reaches to the posterior margin of abdominal tergum VI). The fore femur of <i>R. heoki</i> <b>sp.n.</b> does not widen distally, the maximum width anterior to the median tooth is less than the maximum width posterior to the median tooth (in <i>R. schuhi</i>, the fore femur widens distally, with the maximum width anterior to the median tooth greater than the maximum width posterior to the median tooth). In <i>R. heoki</i> <b>sp.n.</b>, the median tooth of fore femur is more prominent, the width of the femur across the median tooth is slightly greater than the maximum width of the basal part of the femur (in <i>R. schuhi</i>, the width of the femur across the median tooth is subequal to the maximum width of the basal part of the femur). The hind femur of <i>R. heoki</i> <b>sp.n.</b> is shorter and does not reach the posterior margin of abdominal sternum VI when folded back parallel to abdomen (the hind femur of <i>R. schuhi</i> is longer, reaching the anterior half of the operculum). In <i>R. heoki</i> <b>sp.n.</b>, the respiratory siphon has only sparse setae along its ventral margin, without a fringe of dense erect setae (in <i>R. schuhi</i>, the respiratory siphon has a fringe of erect setae along the ventral margin in the basal half).</p> <p> The parameres of these two species have a similar general appearance, but they differ in the following details. The paramere of <i>R. heoki</i> <b>sp.n.</b> is dorsoventrally thicker; its dorsal surface is almost straight, not sinuate, and is constricted in the distal quarter; its distal hook is shorter, with a more open curve, directing its apex ventrad; and the subapical process is more acute, and sub-triangular. In contrast, the paramere of <i>R. schuhi</i> is sinuate in the basal part and is constricted in the distal third; its distal hook is longer and recurved, directing its apex anteroventrad; and the subapical process is round.</p> <p>Female specimen is not available for comparison.</p> <p> <b>Distribution.</b> Borneo: Sarawak.</p>Published as part of <i>Tran, A. D. & Poggi, R., 2019, The genus Ranatra Fabricius (Heteroptera: Nepidae) in Borneo, with a redescription of Ranatra spinifrons Montandon and the description of a new species, pp. 236-246 in Zootaxa 4555 (2)</i> on pages 241-244, DOI: 10.11646/zootaxa.4555.2.4, <a href="http://zenodo.org/record/2624329">http://zenodo.org/record/2624329</a>
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