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Taxonomic novelties and pollen morphological study in the genus Neo-uvaria (Annonaceae)
Chaowasku, Tanawat, KEssLER, Paul J.A., Punnadee, Suwit, Van Der Ham, Raymond W.J.M. (2011): Taxonomic novelties and pollen morphological study in the genus Neo-uvaria (Annonaceae). Phytotaxa 32: 27-42, DOI: 10.11646/phytotaxa.32.1.3, URL: http://biotaxa.org/Phytotaxa/article/view/phytotaxa.32.1.
Neo-uvaria undetermined Airy Shaw
Key to Neo-uvaria species in Thailand 1. Lower leaf surface sparsely covered with stellate indumentum. Flowers pedicellate, pedicels (in flower and in fruit) (3–)5–6(–8) mm long, sepals ca. 2.1 × 2.5 mm, outer petals 5.2 × 2.8–3.0 mm, inner petals 4.0–5.0 × 2.5 mm, stamens 12–15 per flower. Monocarps 3.1–3.3 × 2–2.5 cm..................................................................... N. sparsistellata - Lower leaf surface densely covered with stellate indumentum. Flowers (almost) sessile, sepals ca. 7.5 × 6.5 mm, outer petals 11.5–13.5 × 6.5–8.0 mm, inner petals 8.5–10.0 × 7.5–8.5 mm, stamens 16–20(–22) per flower. Monocarps 6.5–7.0 × 5.2–5.5 cm........................................................................................................................... N. telopeaPublished as part of Chaowasku, Tanawat, KEssLER, Paul J. A., Punnadee, Suwit & Van Der Ham, Raymond W. J. M., 2011, Taxonomic novelties and pollen morphological study in the genus Neo-uvaria (Annonaceae), pp. 27-42 in Phytotaxa 32 on page 29, DOI: 10.11646/phytotaxa.32.1.3, http://zenodo.org/record/489453
FIGURE 6 in Taxonomic novelties and pollen morphological study in the genus Neo-uvaria (Annonaceae)
FIGURE 6. Pollen of Neo-uvaria and Enicosanthum, TEM. A, B. Neo-uvaria foetida: (A) Cross-section of pollen grain, showing continuous exine and intine without recognizable germination zone(s); (B) Detail of cross-section, showing exine and intine sublayers. C, D. N. sparsistellata: (C) Cross-section of pollen grain, showing continuous exine and intine without recognizable germination zone(s); (D) Detail of cross-section, showing exine and intine sublayers. E, F. N. telopea: (E) Cross-section of pollen grain, showing continuous exine and intine without recognizable germination zone(s); (F) Detail of cross-section, showing exine and intine sublayers. G, H. Enicosanthum paradoxum: (G) Cross- section of pollen grain, showing continuous exine and intine without recognizable germination zone(s); (H) Detail of cross-section, showing exine and intine sublayers. Scale bars —5 µm (A, C, E, G), 1 µm (B, D, F, H); b = basal layer, en = endintine, ex = exintine, i = infratectum, t = tectum.Published as part of Chaowasku, Tanawat, KEssLER, Paul J.A., Punnadee, Suwit & Van Der Ham, Raymond W.J.M., 2011, Taxonomic novelties and pollen morphological study in the genus Neo-uvaria (Annonaceae), pp. 27-42 in Phytotaxa 32 on page 38, DOI: 10.11646/phytotaxa.32.1.3, http://zenodo.org/record/489453
Neo-uvaria merrillii Chaowasku 2011, comb. nov.
Neo-uvaria merrillii (C.B.Rob.) Chaowasku, comb. nov. Basionym:— Mitrephora merrillii C.B.Rob. (1908: 67). Griffithianthus merrillii (C.B.Rob.) W.H.Brown ex Merrill (1915: 231). Mitrephora ferruginea Merrill (1904: 16), nom. illeg. TYPE:—THE PHILIPPINES. Luzon, Bataan Province: Mt. Mariveles, January 1904, Merrill 3728 (PNH (destroyed), lectotype (here designated) NY!, isotypes BM!, P!), in fruit. Observations: —Originally, the syntypes of this species consist of disparate elements as stated by Merrill (1915) and Weerasooriya & Saunders (2010). Nevertheless, no one has designated a lectotype, therefore it is done here. Neo-uvaria merrillii primarily differs from N. acuminatissima in having usually (broadly) wedgeshaped leaf base (whilst usually obtuse to rounded in N. acuminatissima), larger sepals (4.5–5.0 × 4.0– 4.5 mm in N. merrillii vs. ca. 2.6 × 2.9 mm in N. acuminatissima) and wider monocarps (ca. 2.2 cm wide in N. merrillii vs. ca. 1.7 cm wide in N. acuminatissima).Published as part of Chaowasku, Tanawat, KEssLER, Paul J. A., Punnadee, Suwit & Van Der Ham, Raymond W. J. M., 2011, Taxonomic novelties and pollen morphological study in the genus Neo-uvaria (Annonaceae), pp. 27-42 in Phytotaxa 32 on pages 34-35, DOI: 10.11646/phytotaxa.32.1.3, http://zenodo.org/record/489453
Neo-uvaria viridifolia Chaowasku 2011, comb. nov.
Neo-uvaria viridifolia (Elmer) Chaowasku, comb. nov. Basionym:— Mitrephora viridifolia Elmer (1913: 1716). TYPE:—THE PHILIPPINES. Mindanao, Agusan Province: Cabadbaran (Mt. Urdaneta), October 1912, Elmer 14184 (PNH (destroyed), lectotype (here designated) NY!, isotypes BM!, BP, DS, E!, G!, GH, L!, NA, P!, U!), in flower. Observations: — N. viridifolia is considerably different from N. acuminatissima, especially in the much sparser stellate indumentum on the lower leaf surface. Moreover, its pedicels are longer than those of N. acuminatissima [(4–) 5–6 mm long in N. viridifolia vs. nearly absent in N. acuminatissima]. The leaf base of N. viridifolia is usually (broadly) wedge-shaped in contrast to N. acuminatissima which usually has obtuse to rounded leaf base. N. acuminatissima was very rarely collected concerning the flowers. Of all specimens investigated, only the collection Sinclair et al. 9250 [this collection seems to have generally smaller leaves with more obtuse base and wider distance of the adjacent tertiary veins compared to those of Elmer 21112 and the type of N. acuminatissima, however, all other morphology of the three collections is similar] has flowers which are smaller than those of N. viridifolia [outer petals ca. 3.6 × 2.5 mm, inner petals ca. 3.8 × 3.0 mm in N. acuminatissima vs. outer petals (7.0–)10.2 × (3.6–) 4.1 mm, inner petals (6.7–)9.5 × 3.4–3.7 mm in N. viridifolia].Published as part of Chaowasku, Tanawat, KEssLER, Paul J. A., Punnadee, Suwit & Van Der Ham, Raymond W. J. M., 2011, Taxonomic novelties and pollen morphological study in the genus Neo-uvaria (Annonaceae), pp. 27-42 in Phytotaxa 32 on page 35, DOI: 10.11646/phytotaxa.32.1.3, http://zenodo.org/record/489453
Neo-uvaria sparsistellata : Chaowasku 2011, sp. nov.
Neo-uvaria sparsistellata Chaowasku, sp. nov. (Figs. 1E, F; 2; 3) Neo-uvaria viridifolia proxima, praecipue petalis minoribus, staminibus carpellisque paucioribus differt. TYPE: — THAILAND. Phatthalung Province: Si Ban Phot District, Khao Pu / Khao Ya National Park, headquarters, August 2005, Gardner et al. ST 1894 (holotype L!, isotypes BKF, K), in flower and fruit (quite young). Medium-sized trees, ca. 15 m tall, ca. 24 cm in dbh, all parts generally covered with stellate hairs intermixed with simple hairs. Young twigs tomentose. Petioles 2.0– 4.5 mm long, densely tomentose. Leaves elliptic, 10.2–32.3 × 3.3–10.1 cm, base (broadly) wedge-shaped, apex generally (narrowly) acuminate, lamina (very) sparsely (appressed-)puberulous above, indumentum mostly on the lower half near the midrib, sparsely puberulous below, upper surface of midrib slightly sunken, (sparsely) (appressed-)tomentose, lower surface of midrib raised, (sparsely) tomentose, secondary veins 15–21 pairs per leaf, angle with midrib 35°–43°. Flowers solitary, axillary, pedicels (in flower and in fruit) (3–)5–6(–8) mm long, densely (appressed-)tomentose, bracts ca. 2 per flower, rather inconspicuous, at the base of the pedicels. Sepals broadly triangular, ca. 2.1 × 2.5 mm, persistent in fruit, outside and margin densely tomentose, inside glabrous. Outer petals elliptic-ovate, 5.2 × 2.8–3.0 mm, outside and margin tomentose, inside glabrous at the base, indumentum of the rest same as outside. Inner petals ovate, 4.0–5.0 × 2.5 mm, apically thickened, indumentum same as outer petals. Stamens 12–15 per flower, ca. 1.4 mm long, connective tissue flat-topped. Carpels 4–6 per flower, stigmas subgloboseellipsoid, ovaries appressed-tomentose, ovules 1 per ovary, basal. Torus more or less flat, sparsely puberulous, slightly enlarged in fruit. Monocarps 1–3 per fruit, sessile, ellipsoid, 3.1–3.3 × 2.0– 2.5 cm, surface (sparsely) tomentose-villous. Seeds 1 per monocarp, ellipsoid, 2.6–2.8 × 1.6–2.1 cm. Distribution: —Peninsular Thailand [Phatthalung Province (Fig. 3)] [only known from two collections (Chaowasku 99 and Gardner et al. ST 1894) collected from the same individual]. Habitats and Phenology: —Occurring in aggrading evergreen/deciduous forests amongst limestone outcrops. Elevation ca. 120 m. Flowering August, December. Fruiting August, December. Field notes: —Bark grey; inner bark brownish-yellow. Petals white maturing yellow. Monocarps pale green with white and pale brown hairs. Etymology: —The epithet refers to the sparse stellate indumentum on the lower leaf surface. Vernacular names: —Ma-Khiew (Thai). Observations: —This new species seems very close to Neo-uvaria viridifolia from the Philippines. They both possess a (very) sparse stellate indumentum on the lower leaf surface whereas it is (much) denser in the other species. In addition, a single stellate hair on the lower leaf surface of both species is relatively (much) smaller than that of the other species. N. sparsistellata, however, principally differs from N. viridifolia in having smaller petals (outer petals 5.2 × 2.8–3.0 mm, inner petals 4.0–5.0 × 2.5 mm in N. sparsistellata vs. outer petals (7.0–)10.2 × (3.6–) 4.1 mm, inner petals (6.7–)9.5 × 3.4–3.7 mm in N. viridifolia), fewer stamens per flower (12–15 in N. sparsistellata vs. ca. 31 in N. viridifolia) and carpels per flower (4–6 in N. sparsistellata vs. ca. 12 in N. viridifolia). Besides, N. sparsistellata always has solitary flowers while N. viridifolia often have two (or three) flowers per inflorescence.Published as part of Chaowasku, Tanawat, KEssLER, Paul J. A., Punnadee, Suwit & Van Der Ham, Raymond W. J. M., 2011, Taxonomic novelties and pollen morphological study in the genus Neo-uvaria (Annonaceae), pp. 27-42 in Phytotaxa 32 on page 29, DOI: 10.11646/phytotaxa.32.1.3, http://zenodo.org/record/489453
Neo-uvaria telopea Chaowasku 2011, sp. nov.
Neo-uvaria telopea Chaowasku, sp. nov. (Figs. 1A–C; 3; 4) Neo-uvaria foetida e Malaysia peninsulari proxima, precipue foliis plerumque maioribus, venis tertiariis adiacentibus plus distantibus, pilis (plus) densibus, floribus maioribus, staminum numero minore differt. TYPE: — THAILAND. Ranong Province: Klong Nakha Wildlife Sanctuary, September 2008, Chaowasku 77 (holotype L!, isotype BKF!), in fruit. Medium-sized trees, ca. 11 m tall, ca. 12 cm in dbh, all parts generally covered with stellate hairs intermixed with simple hairs. Young twigs densely velvety-villous. Petioles 5–7 mm long, densely velvety-villous. Leaves elliptic, (9.5–)13.0–21.5 × (4.0–) 4.7–9.1 cm, base broadly wedge-shaped to obtuse, apex caudateacuminate, lamina almost glabrous above, (densely) villous below, upper surface of midrib slightly sunken, velvety, lower surface of midrib raised, densely (appressed-)velvety-villous, secondary veins 17–18 pairs per leaf, angle with midrib 45º–50º. Flowers solitary, axillary, (almost) sessile, bracts usually 2 per flower, ovate. Sepals broadly ovate to slightly triangular, ca. 7.5 × 6.5 mm, outside and margin (appressed-)velvetytomentose, inside (appressed-)tomentose. Outer petals elliptic, 11.5–13.5 × 6.5–8.0 mm, indumentum on outer side and margin similar to sepals, inside shortly cobwebbed with sparser hairs, base (almost) glabrous. Inner petals slightly ovate to broadly elliptic, 8.5–10.0 × 7.5–8.5 mm, apically thickened, outside (appressed-) velvety-tomentose in the middle, sparser towards the margin which is shortly cobwebbed, inside shortly cobwebbed, sparser towards the base which is (almost) glabrous. Stamens 16–20(–22) per flower, 1.6–1.7 mm long, connective tissue flat-topped. Carpels 5–8 per flower, stigmas ellipsoid-cylindrical, ovaries densely villous, ovules 1(–2) per ovary, basal. Torus more or less flat, villous, enlarged in fruit. Monocarps 1–5 per fruit, sessile, ellipsoid-ovoid, 6.5–7.0 × 5.2–5.5 cm, surface (appressed-)velvety-tomentose. Seeds 1 per monocarp, ellipsoid(-ovoid), 5.0 × 3.4–3.5 cm. Distribution: —Peninsular Thailand [Ranong Province (Fig. 3)] [only known from two collections (Chaowasku 77 and Gardner & Sidisunthorn ST 1992) collected from the same individual]. Habitats and Phenology: —Occurring in understory of semi-disturbed lowland evergreen forests. Elevation ca. 120 m. Flowering December (collection Gardner & Sidisunthorn ST 1992). Fruiting September (collection Chaowasku 77). Field notes: —Crown monopodial with horizontal branching. Bark dark brown, smooth with very shallow horizontal cracks with raised edges; middle bark indistinct; inner bark cream, fibrous. Petals pale green aging greenish-yellow, thick and fleshy; staminal mass pale yellow, blackened at male anthesis (Fig. 1B). Monocarps brown with velvety hairs. Etymology: —The epithet refers to the huge size of the monocarps (Fig. 1C), which therefore can be easily seen from afar. Vernacular names: —Ma-Neng (Thai). Observations: —Odour of rotten fish was emitted from (nearly) dried monocarps. This smell was also detected in those of N. foetida (pers. obs. TC). This new species seems to be closely related to N. foetida occurring in Peninsular Malaysia. It chiefly differs in having generally larger leaves [(9.5–)13.0–21.5 × (4.0–) 4.7–9.1 cm in N. telopea vs. 7.7–15.7(–18.0) × 2.5–5.0(–6.0) cm in N. foetida] with wider distance of the adjacent tertiary veins, denser indumentum on nearly all parts, larger flowers [sepals ca. 7.5 × 6.5 mm, outer petals 11.5–13.5 × 6.5–8.0 mm, inner petals 8.5–10.0 × 7.5–8.5 mm in N. telopea vs. sepals ca. 4.0 × 3.6 mm, outer petals ca. 9.2 × 5.4 mm, inner petals, ca. 6.7 × 5.4 mm in N. foetida], fewer stamens per flower [16–20(– 22) in N. telopea vs. (26–) 27 in N. foetida], and generally fewer carpels per flower [5–8 in N. telopea vs. 7–11 in N. foetida]. Vegetatively, N. telopea resembles N. acuminatissima in the density of the indumentum, but the latter species has much smaller monocarps and flowers. The fruits of N. parallelivenia are unknown but its flowers are noticeably different from those of N. telopea, especially the much smaller sepals and the nonthickened apex of the inner petals of the former.Published as part of Chaowasku, Tanawat, KEssLER, Paul J. A., Punnadee, Suwit & Van Der Ham, Raymond W. J. M., 2011, Taxonomic novelties and pollen morphological study in the genus Neo-uvaria (Annonaceae), pp. 27-42 in Phytotaxa 32 on pages 32-34, DOI: 10.11646/phytotaxa.32.1.3, http://zenodo.org/record/489453
Going Beyond Counting First Authors in Author Co-citation Analysis
The present study examines one of the fundamental aspects of author co-citation analysis (ACA) - the way co-citation
counts are defined. Co-citation counting provides the data on which all subsequent statistical analyses and mappings
are based, and we compare ACA results based on two different types of co-citation counting - the traditional type that
only counts the first one among a cited work's authors on the one hand and a non-traditional type that takes into
account the first 5 authors of a cited work on the other hand. Results indicate that the picture produced through this non-traditional author co-citation counting contains more coherent author groups and is therefore considerably clearer. However, this picture represents fewer specialties in the research field being studied than that produced through the traditional first-author co-citation counting when the same number of top-ranked authors is selected and analyzed. Reasons for these effects are discussed
Variations on the Author
“Variations on the Author” discusses two of Eduardo Coutinho’s recent films (Um Dia na Vida, from 2010, and Últimas Conversas, posthumously released in 2015) and their contribution to the general question of documentary authorship. The director’s filmography is characterized by a consistent yet self-effacing form of authorial self-inscription: Coutinho often features as an interviewer that rather than express opinions propels discourses; an interviewer that is good at listening. This mode of self-inscription characterizes him as an author who is not expressive but who is nonetheless markedly present on the screen. In Um Dia na Vida, however, Coutinho is completely absent form the image, while Últimas Conversas, on the contrary, includes a confessional prologue that moves the director from the margins to the center of his films. This article examines the ways in which these works stand out in the filmography of a director who offers new insights into the notion of cinematic authorship
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