276 research outputs found
FIG. 3 in First record of the genus Leptolophus Remy, 1965 (Mammalia, Perissodactyla) in the late Eocene (Priabonian) of Europe
No full textFIG. 3. — Left mandible (MCNA 9002) showing the m1-2 series of Leptolophus sp. from Zambrana (Alava, Basque Country); A, occlusal view; B, labial view (modified from Astibia et al. 2000). Scale bar: 5 cm.Published as part of Badiola, Ainara, Astibia, Humberto, Suberbiola, Xabier Pereda & Murelaga, Xabier, 2002, First record of the genus Leptolophus Remy, 1965 (Mammalia, Perissodactyla) in the late Eocene (Priabonian) of Europe, pp. 841-848 in Geodiversitas 24 (4) on page 845, DOI: 10.5281/zenodo.465064
FIG. 2 in First record of the genus Leptolophus Remy, 1965 (Mammalia, Perissodactyla) in the late Eocene (Priabonian) of Europe
No full textFIG. 2. — Upper teeth of Leptolophus sp. from Zambrana (Alava, Basque Country); A-C, right P4? (MCNA 9963); A, labial view; B, occlusal view; C, lingual view; D, E, left fragment of upper tooth (MCNA 9964); D, labial view; E, occlusal view. Scale bar: 5 cm.Published as part of Badiola, Ainara, Astibia, Humberto, Suberbiola, Xabier Pereda & Murelaga, Xabier, 2002, First record of the genus Leptolophus Remy, 1965 (Mammalia, Perissodactyla) in the late Eocene (Priabonian) of Europe, pp. 841-848 in Geodiversitas 24 (4) on page 844, DOI: 10.5281/zenodo.465064
Leptolophus Remy 1965
No full textGenus <i>Leptolophus</i> Remy, 1965 <p> TYPE SPECIES. — <i>Leptolophus stehlini</i> Remy, 1965, by original designation.</p>Published as part of <i>Badiola, Ainara, Astibia, Humberto, Suberbiola, Xabier Pereda & Murelaga, Xabier, 2002, First record of the genus Leptolophus Remy, 1965 (Mammalia, Perissodactyla) in the late Eocene (Priabonian) of Europe, pp. 841-848 in Geodiversitas 24 (4)</i> on page 842, DOI: <a href="http://zenodo.org/record/4650641">10.5281/zenodo.4650641</a>
FIG. 4. — A, diagram d in First record of the genus Leptolophus Remy, 1965 (Mammalia, Perissodactyla) in the late Eocene (Priabonian) of Europe
No full textFIG. 4. — A, diagram d/D (in mm) of the M1 of several Pachynolophidae from the Eocene of Europe, D, largest diagonal (from the parastyle to the hypocone) of the upper teeth, d, diagonal perpendicular to D; B, diagram l/L (in mm) of the M2 of several Pachynolophidae from the Eocene of Europe, l, maximum width, L, average length of the external wall of the ectoloph. Measurements from Casanovas 1975 and Casanovas et al. 1998 for Plagiolophus annectens (Owen, 1848); Cuesta 1994 for Plagiolophus mazateronensis Cuesta, 1994 and Leptolophus sp.; Remy 1998 for Leptolophus stehlini Remy, 1965 and Leptolophus nouleti Stehlin, 1904.Published as part of Badiola, Ainara, Astibia, Humberto, Suberbiola, Xabier Pereda & Murelaga, Xabier, 2002, First record of the genus Leptolophus Remy, 1965 (Mammalia, Perissodactyla) in the late Eocene (Priabonian) of Europe, pp. 841-848 in Geodiversitas 24 (4) on page 846, DOI: 10.5281/zenodo.465064
FIG. 3 in Hypsodont Myomiminae (Gliridae, Rodentia) from five new localities in the Lower Miocene Tudela Formation (Bardenas Reales, Ebro Basin, Spain) and their bearing on the age of the Agenian-Ramblian boundary
No full textFIG. 3. — Distribution chart of the Myomiminae species studied in this paper and in Daams (1990). The succession of the localities from Daams (1990) is, though in stratigraphical order, not calibrated.Published as part of <i>Ruiz-Sánchez, Francisco J., Murelaga, Xabier, Larrasoaña, Juan C., Freudenthal, Matthijs & Garcés, Miguel, 2012, Hypsodont Myomiminae (Gliridae, Rodentia) from five new localities in the Lower Miocene Tudela Formation (Bardenas Reales, Ebro Basin, Spain) and their bearing on the age of the Agenian-Ramblian boundary, pp. 645-663 in Geodiversitas 34 (3)</i> on page 651, DOI: 10.5252/g2012n3a10, <a href="http://zenodo.org/record/5375769">http://zenodo.org/record/5375769</a>
A large testudinid with African affinities in the post‐Messinian (lower Pliocene) record of south‐eastern Spain
No full textHerein, we describe Alatochelon myrteum gen. et sp. nov., a large tortoise from the post-Messinian (lower Pliocene) of the area of Puerto de la Cadena (Region of Murcia), Spain. The new taxon cannot be attributed to Titanochelon, which represented the only lineage of large tortoises previously recognized in the Neogene record of Europe. Alatochelon myrteum shows African affinities, especially with the extant African spurred tortoise Centrochelys sulcata. Although close phylogenetic relationships have previously been recognized among some tortoises of both continents, the dispersal of this lineage had always been proposed as having occurred in only one direction: from Europe to Africa. The dispersal of the lineage including the new Spanish form and Centrochelys sulcata from Africa to Europe is proposed here. This proposal is compatible with those previously recognized for some lineages of mammals also found in Puerto de la Cadena, identified as African lineages that probably reached Europe during the Messinian Salinity Crisis event. An African origin is also proposed for the lineage of Titanochelon. Therefore, the two lineages of large derived testudinids (i.e. Geochelona) recognized in the European record experienced diachronic dispersal events from Africa to Europe: that to which Alatochelon belongs probably during the Messinian and the other much earlier, at the beginning of the Miocene or before.Fil: Pérez García, Adán. Universidad Nacional de Educacion A Distancia. Facultad de Ciencias.; EspañaFil: Vlachos, Evangelos. Museo Paleontológico Egidio Feruglio; Argentina. Consejo Nacional de Investigaciones Científicas y Técnicas. Centro Científico Tecnológico Conicet - Patagonia Norte; ArgentinaFil: Murelaga, Xabier. Universidad del País Vasco; Españ
FIG. 6 in The freshwater and terrestrial turtles from Monte Pila and Fuenmayor (La Rioja, northern Spain): new data on the lower Miocene turtle diversity of the Ebro Basin
No full textFIG. 6. — Plastral plates of several individuals of freshwater Ptychogasterinae Ptychogaster (Temnoclemmys) cf. bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola & Astibia, 1999, from the lower Miocene (MN2) of Monte Pila 1 (Lardero, La Rioja, Spain): A-V, epiplastra, in dorsal and ventral views; A, B, MP1b-sup1-a; C, D, MP1a-55; E, F, MP1a-1210a; G, H, MP1a-975a; I, J, MP1a-1145a; K, L, MP1a-1745; M, N, MP1b-481; O, P, MP1a-1679b; Q, R, MP1a-1189a; S, T, MP1a- 1557; U, V, MP1a-529b; W-AC, entoplastral, in ventral view; W, MP1a-1394b; X, MP1b-1597a; Y, MP1a-347a; Z, MP1a-672; AA, MP1a-449; AB, MP1a-180a; AC, MP1a-409. Abbreviations for the scutes (in upper case): GU, gular; HU, humeral; PC, pectoral. Scale bar: 3 cm.Published as part of Pérez-García, Adán, Suarez-Hernando, Oier, Hernández, Jose M., García, Salvador & Murelaga, Xabier, 2021, The freshwater and terrestrial turtles from Monte Pila and Fuenmayor (La Rioja, northern Spain): new data on the lower Miocene turtle diversity of the Ebro Basin, pp. 75-94 in Geodiversitas 43 (3) on page 82, DOI: 10.5252/geodiversitas2021v43a3, http://zenodo.org/record/454837
<i>Iberoccitanemys atlanticum</i> n. comb. : nouvelles données sur la diversité et les répartitions paléobiogéographiques des tortues bothremydidés du Campanien et Maastrichtien de l’Europe
No full textLes Bothremydidae est le groupe de tortues le plus abondant dans les gisements du Campanien et Maastrichtien (Crétacé supérieur) du Sud-Ouest de l’Europe. Plusieurs membres de Foxemydina Gaffney, Tong & Meylan, 2006 ont été identifiés dans le Sud de la France et la moitié Nord-Est de l’Espagne. L’espèce la moins bien caractérisée est le taxon problématique ‘Polysternon’ atlanticum Lapparent de Broin & Murelaga, 1996. Il s’agit d’une espèce pour laquelle une diagnose adéquate permettant de confirmer sa validité spécifique n’est pas disponible, et pour laquelle l’attribution générique est considérée comme douteuse. Sa présence a été exclusivement reconnue dans sa localité type, la carrière du Campanien supérieur de Laño, dans le Comté de Treviño (province de Burgos, Nord de l’Espagne). Bien que les connaissances sur les Bothremydidae Baur, 1891 aient considérablement augmenté après la description de ‘Polysternon’ atlanticum, aucune nouvelle information sur cette espèce n’a été publiée depuis les années 1990. L’analyse de nombreux spécimens non publiés du bothremydidé de Laño nous permet de confirmer la validité de cette espèce. Il est non seulement identifié dans sa localité type, mais aussi dans d’autres régions espagnoles et dans le Sud de la France. La diversité des bothremydidés dans le Crétacé supérieur d’Europe est reconnue comme moindre que celle considérée jusqu’à présent. Ainsi, l’espèce ‘Iberoccitanemys convenarum’ (Laurent, Tong & Claude, 2002), qui a été définie à partir d’un individu dans le registre français, puis identifiée en Espagne, est reconnue comme synonyme de l’espèce de Laño. Une diagnose modifiée pour l’espèce du Campanien supérieur au Maastrichtien supérieur, Iberoccitanemys atlanticum (Lapparent de Broin & Murelaga, 1996) n. comb., est proposée.Bothremydidae is the most abundant clade of turtles in the Campanian and Maastrichtian (Upper Cretaceous) fossil record of southwestern Europe. Several members of Foxemydina Gaffney, Tong & Meylan, 2006 are known in an area that includes Southern France and the North-Eastern half of Spain. The problematic ‘Polysternon’ atlanticum is the worst characterized, lacking a diagnosis that allows its specific validity to be confirmed, and whose generic attribution has been recognized as doubtful. Its presence was exclusively proposed in its type locality, the upper Campanian quarry of Laño, in Treviño County (Burgos Province, North of Spain). Despite the fact that knowledge about Bothremydidae has markedly increased after the description of ‘Polysternon’ atlanticum Lapparent de Broin & Murelaga, 1996, no new information about this species has been published since the 1990s. The analysis of abundant unpublished material of the bothremydid from Laño allows us to confirm the validity of this species. As a consequence of this study, it is not only identified in its type locality, but also in other Spanish regions and in the south of France. The diversity of Bothremydidae Baur, 1891 in the Upper Cretaceous of Europe is lower than previously considered. Thus, the species ‘Iberoccitanemys convenarum’ (Laurent, Tong & Claude, 2002), originally defined for the French record, and subsequently also identified in Spain, is identified here as a synonym of the species described in Laño. An emended diagnosis for the upper Campanian to upper Maastrichtian, Iberoccitanemys atlanticum (Lapparent de Broin & Murelaga, 1996) n. comb., is proposed.</p
Xabier Zubiriren pentsamendua dela-ta
No full textCon motivo de la muerte del filosofo Xabier Zubiri, y debido a los diferentes homenajes realizados en su honor, se ha llegado a conocer mucho mejor su obra. El autor en este articulo nos describe algunas de sus características y la sitúa dentro de los diferentes movimientos de la época haciendo algunas comparacionesAfter the death of the philosopher Xabier Zubiri, and due to the different homages tributed in his honour, his work has become much better known. In this article, the author describes some of his main characteristics and places Zubiri within the various philosophical currents of the era, making some comparison
Ptychogaster (Temnoclemmys) bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola & Astibia 1999
No full textPtychogaster (Temnoclemmys) cf. bardenensis Murelaga, Lapparent de Broin, Pereda Suberbiola & Astibia, 1999 (Figs 3-10) MATERIAL EXAMINED. — Numerous disjointed and isolated plates, several partial carapace and plastra, and some relatively complete shells from Monte Pila (e.g., MP 1a-6, MP 1a-14, MP 1a-48, MP 1a- 55, MP 1a-56, MP 1a-81, MP 1a-109, MP 1a-169, MP 1a-180a, MP 1a-228, MP 1a-252, MP 1a-263, MP 1a-277a, MP 1a-281, MP 1a-341, MP 1a-347a, MP 1a-367, MP 1b-369, MP 1b-380b, MP 1a-385, MP 1a-409, MP 1a-449, MP 1a-474, MP 1a-475, MP 1a- 511, MP 1a-528, MP 1a-529b, MP 1a-543, MP 1a-638, MP 1a-643, MP 1a-659, MP 1a-672, MP 1a-679, MP 1a-681, MP 1a-694, MP 1a- 707, MP 1a-757, MP 1a-865, MP 1a-879, MP 1a-911, MP 1a-918, MP 1a-975a, MP 1a-1025, MP 1a-1066, MP 1a-1068b, MP 1a-1070, MP 1a-1145a, MP 1a-1189a, MP 1a-1194, MP 1a-1210a, MP 1a-1221, MP 1a-1227, MP 1a-1242, MP 1a-1305, MP 1a-1333, MP 1a-1342, MP 1a-1345, MP 1a-1350, MP 1a-1364, MP 1a-1394b, MP 1a-1417, MP 1a-1421, MP 1a-1422, MP 1a-1423, MP 1a-1557, MP 1a-1570, MP 1a-1594, MP 1a-1679b, MP 1a-1724, MP 1a-1745, MP 1b-95, MP 1b-169, MP 1b-415, MP 1b-456, MP 1b-467, MP 1b-481, MP 1b-556, MP 1b-585, MP 1b-1597a, MP 1b-sup1-a, MP 2-93, MP 2-175, MP 2-306, MP 2-307, MP 2-308, MP 3-1; Figs 3-9) and Fuenmayor (e.g., FM1-1, FM1-2, FM1-3, FM1-4, FM2-1, MCNA 16032, MCNA 16031; Fig. 10). LOCALITY AND HORIZON. — Most specimens come from the Monte Pila sites, municipality of Lardero, Autonomous Community of La Rioja, Spain. Levels MP 1, MP 2 and MP 3 (Figs 3-9). Other specimens came from the site of Fuenmayor, municipality of Fuenmayor, Autonomous Community of La Rioja, Spain (Fig. 10). Western part of the Ebro Basin. Transitional facies between the Najera and Haro formations. Local zone Y, biozone MN 2, lower Miocene (see Figs 1, 2 and the section Geographic and geological setting). DESCRIPTION The maximum length of the shells of the larger specimens of the freshwater turtles identified at both Monte Pila and Fuenmayor is close to 20 cm (Figs 3-10). However, elements corresponding to individuals of very varied sizes are identified, some of them with a length of less than half that of the largest specimens. Some of the most complete shells, as well as the articulated elements of other specimens recognized as adult individuals due to their size, show fusion of many of their sutures, so that the margins of each plate cannot be recognized in them (Figs 3; 5). The thickness of the plates is greater in the larger individuals. Although the width/length ratio of the shell is variable, this form shows a relatively wide carapace, being recognized as slightly longer than wide in some cases (Fig.3G). The carapaces are low. The abundance of specimens allows assessment of the variability in morphology and dimensions of several elements. For example, the relatively wide nuchal plates are almost as long as wide in some individuals, but slightly wider than long in others (Fig. 4 A-L). All of them show a thickening in the visceral region. The neural series anteriorly contacts the nuchal plate, and posteriorly the suprapygals. The neurals are wide in relation to their length (Fig. 4 M-O; 10C, D). The morphology of these plates is variable. However, most of them are hexagonal, with the shortest margins being postero-laterally located. Although the anterior suprapygal is subquadrangular, the posterior is hexagonal, noticeably wider than long (Fig. 4 Q-W). The pygal is also subquadrangular (Fig. 4 X-Z). This taxon has a single cervical scute (Figs 3A; 4 A-L; 9F). Its lateral margins are subrounded. Its width/length ratio shows a wide range of variability, being slightly wider than long in some specimens, but almost twice as long as wide in others. The overlap of the cervical on the visceral surface of the nuchal is relatively long. The vertebral scutes are hexagonal, almost as wide as they are long (Fig. 3 A-G). The first and last ones are slightly wider than the second to fourth vertebrals, which are all similar in width. However, the vertebral series is narrow, the first scute being narrower than the nuchal in many cases, not overlapping the postero-lateral margins of this plate. The fifth vertebral overlaps the anterior region of the pygal plate (Fig. 4 X-Z). The contact between the hyoplastra and the hypoplastra, and that between the hypoplastra and the peripheral series, corresponds to a ligamentous junction, so that a hinge is developed between these plates, allowing mobility of the posterior plastral region (Figs 3F; 5 A-F; 7B-M; 8; 9D-E; 10A, B). The lateral margins of both plastral lobes are subrounded. The anterior lobe is very wide. Its anterior margin varies from subrounded to almost straight. Although the posterior lobe is clearly expanded in some specimens, it is much narrower in others. Thus, the development of the anal notch is variable in width and length, as is the shape of its margins (Figs 3F; 7 N-U; 8; 10A, B). The epiplastral lip is concave (Figs 5; 6 A-V; 9A, B; 10A, B, G-L). Both the length and the thickness of this lip are observed to be highly variable. The dorsal thickening of this lip shows a pair of anterior processes, which exceed the margin of the anterior plastral lobe, at the level of the contacts between the gulars and the humerals. The gular scutes overlap the anterior region of the entoplastron in some specimens, but not in others (Fig. 6 W-AC). In the same way, the humero-pectoral sulci may be located next to the posterior margin of the entoplastron in some specimens, but overlap this plate in others, ranging from a very short overlap of the pectorals on the entoplastron in some individu- als, to a relatively long overlap, greater than a quarter of the length of this plate, in others. The abdominal scutes overlap the posterior region of the hyoplastra (Figs 5 A-F; 7B-I; 9D-E; 10A). The anal scutes are restricted to the xiphiplastra, their anterior margin being well away from the suture between those plates and the hypoplastra (Figs 7 N-U; 8; 9C; 10A, N).Published as part of Pérez-García, Adán, Suarez-Hernando, Oier, Hernández, Jose M., García, Salvador & Murelaga, Xabier, 2021, The freshwater and terrestrial turtles from Monte Pila and Fuenmayor (La Rioja, northern Spain): new data on the lower Miocene turtle diversity of the Ebro Basin, pp. 75-94 in Geodiversitas 43 (3) on pages 78-85, DOI: 10.5252/geodiversitas2021v43a3, http://zenodo.org/record/454837
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