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REDESCRIPTION OF TETRAMORIUM ATLANTE CAGNIANT, 1970, NEW STATUS (HYMENOPTERA: FORMICIDAE: MYRMICINAE)
Lech Borowiec, Christophe Galkowski (2016): REDESCRIPTION OF TETRAMORIUM ATLANTE CAGNIANT, 1970, NEW STATUS (HYMENOPTERA: FORMICIDAE: MYRMICINAE). Annales Zoologici 66 (1): 43-52, DOI: 10.3161/00034541ANZ2016.66.1.00
Figures 1–2 in REDESCRIPTION OF TETRAMORIUM ATLANTE CAGNIANT, 1970, NEW STATUS (HYMENOPTERA: FORMICIDAE: MYRMICINAE)
Figures 1–2. Tetramorium atlante Cagniant, worker: (1) dorsal view; (2) lateral view. Scale bar = 1 mm.Published as part of Lech Borowiec & Christophe Galkowski, 2016, REDESCRIPTION OF TETRAMORIUM ATLANTE CAGNIANT, 1970, NEW STATUS (HYMENOPTERA: FORMICIDAE: MYRMICINAE), pp. 43-52 in Annales Zoologici 66 (1) on page 45, DOI: 10.3161/00034541ANZ2016.66.1.003, http://zenodo.org/record/26954
Oxyopomyrmex emeryi Santschi 1908
Oxyopomyrmex emeryi Santschi, 1908 (Figs. 1, 2, 3, 73) Oxyopomyrmex emery Santschi, 1908: 524, fig. 9 (w.); Cagniant 1968: 143; Délye 1971: 211 (g.m.); Bernard 1976: 114. Type locality: Dratamar, near Kairuan, Tunisia. Type material. Lectotype worker (top on the pin) (present designation): O. Emeryi || Tunisie | Kairouan | Dr Santschi || Sammlung | Dr. F. Santschi | Kairouan || CASENT | 0 101200 | ANTWEB || red label (NHMB); 27 paralectotype workers: the same data as lectotype (NHMB). Other material examined. 11 workers— Oxyopomyrmex | Emeryi Sants. | SANTSCHI det. 1913 || Kairouan | X 1913 || Sammlung | Dr. F. Santschi | Kairouan (NHMB); 2 workers—MUSEUM PARIS | Kairounan | 1911 | F. Santschi || Oxyopomyrmex | Emeryi | Santschi | type | Kairouan (MNHN); 1 worker —MUSEUM PARIS | Kairounan | Santschi 1910 || Oxyopomyrmex | Emeryi | Santschi | type (MNHN). Gyne. Description: see Délye (1971). Male. Description: see Délye (1971). Worker. Redescription. Measurements (n= 39): HL: 0.732 ± 0.017 (0.698-0.76); HW: 0.721 ± 0.018 (0.692- 0.758); SL: 0.537 ± 0.014 (0.514-0.559); EL: 0.307 ± 0.009 (0.291-0.319); EW: 0.167 ± 0.005 (0.162-0.179); ML: 0.923 ± 0.037 (0.849-1.005); PSL: 0.192 ± 0.012 (0.162-0.218); SDL: 0.136 ± 0.008 (0.123-0.156); PL: 0.339 ± 0.019 (0.313-0.363); PPL: 0.245 ± 0.01 (0.223-0.257); PH: 0.239 ± 0.009 (0.223-0.257); PPH: 0.232 ± 0.008 (0.221-0.246); PNW: 0.469 ± 0.011 (0.448-0.492); TL: 0.567 ± 0.022 (0.52-0.587); TW: 0.105 ± 0.006 (0.092- 0.117); PW: 0.188 ± 0.012 (0.162-0.201); PPW: 0.279 ± 0.012 (0.268-0.302); HI: 98.4 ± 1.2 (94.6-100.7); SI 1: 73.4 ± 1.4 (70.9 -76.0); EI: 54.4 ± 1.9 (51.8-57.7); SI 2: 74.3 ± 1.3 (72.0- 76.6); MI: 196.9 ± 5.1 (187.4-204.3); SPI: 140.6 ± 10.0 (118.0- 162.7); PI 1: 141.7 ± 5.6 (130.4-152.3); PI 2: 40.2 ± 1.9 (35.8-43.3); PPI 1: 104.2 ± 5.4 (94.9- 112.2); PPI 2: 59.7 ± 1.9 (57.1-63.5); HTI 1: 78.8 ± 2.0 (75.0- 82.1); HTI 2: 18.5 ± 1.1 (15.8-20.3). Head and abdomen dark brown. Thorax and legs brown to brick-red colour. Antennae dark brown, only apex of the scapes and first segments of funiculus paler (Figs. 1, 2, 3). Head oval, longer than wide (Fig. 3). Anterior margin of the clypeus smooth and straight. Eyes elongate, gently narrowing downward, 0.4 times as long as length of the head. Scape short, 0.7 times as long as width of the head, at base 0.83 times as wide as in apex, gradually widened, slightly bent downward. Funiculus short, 1.45 times as long as scape, the first segment elongate, triangular, 1.4 times as long as wide on apex, 1.9 times as long as second segment, length ratio of segments 100: 53: 53: 59: 47: 59: 71: 117: 129: 224, apical segments 1.6 times as wide as basal segments. Surface of the scape with a very fine microsculpture, shiny, covered with short, appressedor semierect setae (Figs. 1, 2, 3). Promesonotum 1.1 times as long as wide, gently and regularly convex in profile. Promesonotal suture distinct, the border between dorsal and posterior surfaces of the promesonotum gently curved in profile view. Propodeum quadrate, 0.9 times as long as wide, propodeal spines short, triangular but thin, rising obliquely upwards (Figs. 1, 2). Petiole rounded with a short peduncle, its anterior face slightly convex, node angulated in profile. Posterior face slightly rounded. Ventral margin of petiole straight or with a small ventral lobe (Fig. 2). Postpetiole regularly rounded in profile. In dorsal view postpetiole 1.1 times as long as wide, regularly widened from base to top, apical half with gently rounded sides (Fig. 2). Mandibles rounded, with outer and dorsal edges straight and smooth, inner margin with 7-8 teeth, apical tooth massive and long. Clypeus smooth and shiny on the entire surface. Frontal carinae short, extending to 1 / 3 length of the eye, antennal fossa area deeply impressed, shiny, frontal lobes with thin longitudinal rugae, microreticulate, shiny between rugosities. Frons shiny, with longitudinal striae, distinct reticulation. Area above eyes shiny and distinctly reticulate to microreticulate, ventral surface of head with indistinct microreticulation or microgranulate, gena smooth and shiny (Figs. 2, 3). Entire head bearing setae, posterior margin with sparse erect setae directed forward, sides of the head with a few appressed setae directed toward anterior margin, frontal area with sparse, appressed to semierect setae placed transversely, directed to the centre of the head, ventral surface of the head with a prominent psammophore and appressed to erect, long setae. Pronotum shiny, and finely punctate on the entire surface, only lateral surfaces micropunctae or smooth. Dorsal suface of pronotum shiny, punctate with transverse rugae. Mesonotum punctate on the entire surface, lateral surfaces with several transverse striae on posterior surface, propodeum punctate, with distinct oblique rugae below spiracles (Figs. 1, 2, 73). Dorsal suface of mesosoma with at least 5 erect setae on anterior half, mesonotum and propodeum with a few erect setae. Base of petiole and postpetiole punctate on the entire surface, nodes punctate, micropunctate on top, shiny, bearing several sparse setae. Gaster shiny with micropunctation or shagreened; bearing sparse, erect and semierect setae. Legs short, hind femora 0.8 times as long as mesosoma length, hind tibia 0.7 times as long as hind femora, hind tarsi 1.2 times as long as hind femora. Dorsal surface of femora with short, sparse, appressed setae, inner margin with a row of sparse, short setae, tibiae covered with short, appressed to semierect pubescence on the entire surface, with a row of slightly long and more erect setae on inner margins (Fig. 2). Biological data. Nests occur in humid sand between dunes, or under stones in sandy or clay soil. The entrance is always single and narrow, surrounded by a regular crater, 5 to 8 centimeters in diameter. Nest depth is at least 40- 50 centimeters, and consists of one enormous central gallery and 5 to 10 small chambers. Workers are nocturnal, carrying the cuttings and debris of Helianthemum lippii (L.) (Délye 1971). Distribution. Algeria, Tunisia. Differential diagnosis. Gyne. According to the description (Délye 1971), Oxyopomyrmex emeryi, along with O. krueperi and O. santschii, belong to a group of species that have the genae covered with longitudinal striae without rugosity. In this group, O. emeryi is distinguished by punctation occurring between the longitudinal striation on the lateral surfaces of the thorax. The two other known species from this group are characterized by the lateral surfaces of the thorax bearing longitudinal striation with rugosity or a smooth surface between the striae. Male. Oxyopomyrmex emeryi is one of two known species that is devoid of propodeal spines (Délye 1971). In this group, O. emeryi differs from O. magnus by having the scutellum covered by longitudinal striae on the entire surface, whereas O. magnus has at least the center of the scutellum smooth and shiny, without striation. Worker. Oxyopomyrmex emeryi belongs to a group of species that have distinctly punctate pronotum. In this group, O. emeryi differs from O. polybotesi and O. pygmalioni in the lacking rugosity and longitudinal striae on dorsal surface of pronotum. In comparison with O. oculatus, O. emeryi differs in the occurrence of vertical striation on the dorsal surface of the pronotum and from O. negevensis it can be distinguished by the shining abdomen without a dense layer of appressed micropulpae on dorsal surface of the first tergite.Published as part of Sebastian Salata & Lech Borowiec, 2015, A taxonomic revision of the genus Oxyopomyrmex André, 1881 (Hymenoptera: Formicidae), pp. 1-66 in Zootaxa 4025 (1) on pages 8-11, DOI: 10.11646/zootaxa.4025.1.
Oxyopomyrmex pygmalioni Salata & Borowiec, 2015, sp. nov.
Oxyopomyrmex pygmalioni sp. nov. (Figs. 61, 62, 63, 84) Etymology. Named after the Cypriot king Pygmalion. According to Ovid’s narrative, Pygmalion was a sculptor who fell in love with an ivory statue which he had carved. Through Aphrodite’s blessing the sculpture changed to a woman and the king could fortunately married her. Oxyopomyrmex pygmalioni is the most densely and distinctly sculptured species, reminiscent of a raw sculpture. Type locality: Cape Drepano, Cyprus. Type material. Holotype worker: CYPRUS, Paphos distr., 21 m | Cape Drepano | 34 ° 54.027 N / 32 ° 19.159 E | 2 V 2012, L. Borowiec (DBET); 18 paratype workers: the same data as lectotype (DBET). Additional material: specimen on photo: Cypern— 17 | Prov. Paphos, Polis | vic. Argaka, 200 mH | Leg. Sanetra, 0 1.04. 94 || Oxyopomyrmex | oculatus | A. Schulz, det. (available from; https://www.antweb.org/specimen/ CASENT0101775, Accessed 2 April 2015). Gyne. Unknown. Male. Unknown. Worker. Description. Measurements (n= 19): HL: 0.698 ± 0.022 (0.659-0.726); HW: 0.662 ± 0.022 (0.603- 0.701); SL: 0.472 ± 0.015 (0.446-0.5); EL: 0.22 ± 0.005 (0.212-0.229); EW: 0.124 ± 0.003 (0.122-0.134); ML: 0.817 ± 0.037 (0.749-0.894); PSL: 0.17 ± 0.012 (0.14-0.19); SDL: 0.121 ± 0.008 (0.109-0.134); PL: 0.311 ± 0.022 (0.268-0.335); PPL: 0.224 ± 0.025 (0.179-0.246); PH: 0.223 ± 0.01 (0.201-0.235); PPH: 0.219 ± 0.006 (0.212- 0.229); PNW: 0.438 ± 0.014 (0.402-0.458); TL: 0.478 ± 0.02 (0.458-0.503); TW: 0.092 ± 0.003 (0.089-0.098); PW: 0.163 ± 0.01 (0.145-0.184); PPW: 0.238 ± 0.014 (0.212-0.268); HI: 95.0 ± 1.5 (91.5-96.8); SI 1: 71.3 ± 1.8 (68.3-75.2); EI: 56.4 ± 1.75 (55.2-60.9); SI 2: 71.3 ± 1.8 (68.3-72.2); MI: 186.7 ± 4.9 (176.2-197.7); SPI: 138.8 ± 10.7 (113.8-164.2); PI 1: 138.8 ± 8.3 (125.1-150.2); PI 2: 37.3 ± 2.6 (32.9-41.9); PPI 1: 107.6 ± 4.3 (100.0- 112.8); PPI 2: 54.4 ± 3.8 (46.2-62.2); HTI 1: 72.9 ± 2.6 (67.3-76.3); HTI 2: 19.3 ± 0.8 (17.9-20.1). Head, thorax and abdomen black. Antennae black, only apex of the scapes and first five segments of funiculus brown. Legs black, only apex of femora, tibiae and tarsi brown (Figs. 61, 62, 63). Head rectangular, longer than wide, lateral surfaces below eyes straight, slightly rounded on the posterior edges (Fig. 63). Anterior margin of the clypeus shiny with longitudinal striae, straight. Eyes elongate, strongly narrowing downward, reaching anteroventral margin of head, 0.3 times as long as length of the head. Scape short, 0.7 times as long as width of the head, at base 0.8 times as wide as in apex, gradually widened, slightly bent downward. Funiculus short, 1.5 times as long as scape, first segment elongate, triangular, 1.6 times as long as wide on apex, 2.25 times as long as second segment, length ratio of segments 100: 44: 33: 33: 39: 44: 56: 89: 111: 200, apical segments 1.9 times as wide as basal segments (Figs. 62, 63). Surface of the scape with very fine microsculpture, shiny, covered with short, dense, appressed setae. Promesonotum 1.1 times as long as wide, convex in profile. Promesonotal suture distinct, the border between dorsal and posterior surfaces of the promesonotum gently curved in profile view. Propodeum quadrate, 0.9 times as long as wide, propodeal spines short, triangular, erect (Fig. 62). Petiole rounded with short peduncle, its anterior face straight, node rounded on dorsal surface in profile. Posterior face straight. Ventral margin of petiole with small teeth-like projection or smooth (Fig. 62). Postpetiole regularly rounded in profile. Postpetiole 0.9 times as long as wide in dorsal view, regularly widened from base to top, apical half with gently rounded sides (Figs. 61, 62). Mandibles rounded, with outer and dorsal edges straight and smooth, inner margin with 7-8 teeth, the apical tooth massive and the longest. Clypeus shiny, micropunctate with longitudinal striae on entire surface. Frontal carinae short, extending to 1 / 3 length of eye; antennal fossa deeply impressed, microreticulate, frontal lobes rugulose with thick longitudinal striae, shiny between rugosities. Frons shiny, all surface with thick longitudinal striae and dense rugulose. Area above eyes shiny with thick longitudinal striae and densely rugulose. Ventral surface of the head with distinct striation and rugulose, gena shiny, rugulose with thick longitudinal striae (Figs. 62, 63). Entire head bearing setae, posterior margin with dense erect setae directed forward, lateral surfaces of the head with appressed setae directed toward anterior margin, frontal area with dense, appressed to semierect setae placed transversely, directed to the center of the head, ventral surface of the head with a prominent psammophore and appressed to erect long setae. Pronotum strongly punctate, rugose, lateral surfaces strongly punctate with fine longitudinal striae. Dorsal suface of pronotum strongly punctate and rugose at the edges to rugose at the central surface. Mesonotum strongly punctate on the top and lateral surfaces, dorsal surface of propodeum strongly punctate to punctate, below spiracles strongly punctate with fine longitudinal striae at the posterior edge (Figs. 61, 62, 84). Dorsal suface of mesosoma with at least 5 long setae on anterior half, mesonotum and anterior propodeum with a few long setae. Base of petiole and postpetiole on the entire surface punctate, nodes of the petiole punctate on the top and lateral surfaces, postpetiole node punctate, smooth with sparse punctation on the top, covered with several setae. Gaster shiny and smooth, bearing dense, erect to semierect setae (Figs. 61, 62). Legs short, hind femora 0.7 times as long as mesosoma length, hind tibia 0.8 times as long as hind femora, hind tarsi 1.1 times as long as hind femora. Dorsal surface of femora with short, sparse, semierect setae, inner margin with a row of the sparse, long, semierect setae, tibiae bearing long, appressed to semierect setae on the entire surface, inner margins with a row of appressed setae (Fig. 62). Biology. Borowiec collected specimens on grassy coastal terraces of cliffs. Despite low temperatures (16 ° C) and windy weather, the workers were active and were hand collected between clumps of grass. The following ant species were collected in the area: Messor sp. (hellenius complex), Plagiolepis pallescens sensu Radchenko, and Tapinoma simrothi Krausse. Distribution. Cyprus. Differential diagnosis. Worker. Oxyopomyrmex pygmalioni belongs to the group of species characterized by a distinctly punctate propodeum. It is distinguished from O. emeryi and O. oculatus by the longitudinal striae covering whole frontal surface of the head and the presence of rugosity on dorsal surface of the pronotum. From O. negevensis it differs by the lack of the transverse striation on dorsal surface of the pronotum, the shiny abdomen and by more dense and thicker hairiness at the scape and legs. See also differential diagnosis under O. polybotesi.Published as part of Sebastian Salata & Lech Borowiec, 2015, A taxonomic revision of the genus Oxyopomyrmex André, 1881 (Hymenoptera: Formicidae), pp. 1-66 in Zootaxa 4025 (1) on pages 53-55, DOI: 10.11646/zootaxa.4025.1.
Oxyopomyrmex polybotesi Salata & Borowiec, 2015, sp. nov.
Oxyopomyrmex polybotesi sp. nov. (Figs. 58, 59, 60, 83) Oxyopomyrmex oculatus: Forel 1911: 344; Kiran & Karaman 2012: 23 (misidentification) Etymology. Named after giant Polybotes. According to the Greek mythology, during the Gigantomachy Polybotes was crushed under Nisyros (type locality of the species), which was thrown at him by Poseidon. Type locality: Moni Evangelistrias, Nisyros Is., Greece. Type material. Holotype worker: GREECE, Dodecanese, 270 m | Nisyros, Moni Evangelistrias | 36,601 N / 27, 151 E | 1 VI 2001, M. Chatzaki (DBET); 45 paratype workers: the same data as holotype (DBET, SSC). Other material examined. 1 worker— Oxyopomyrmex | oculatus Andre? | Smyrne || Forel || Smyrne || MUSEUM PARIS | COLLECTION | ERNEST ANDRE | 1914 (MNHN). Gyne. Unknown. Male. Unknown. Worker. Description. Measurements (n= 30): HL: 0.705 ± 0.017 (0.681-0.732); HW: 0.66 ± 0.012 (0.648- 0.69); SL: 0.467 ± 0.013 (0.447-0.48); EL: 0.221 ± 0.01 (0.201-0.234); EW: 0.12 ± 0.008 (0.106-0.134); ML: 0.818 ± 0.02 (0.785-0.849); PSL: 0.18 ± 0.008 (0.162-0.19); SDL: 0.114 ± 0.009 (0.094-0.128); PL: 0.328 ± 0.013 (0.302-0.341); PPL: 0.224 ± 0.014 (0.201-0.24); PH: 0.216 ± 0.008 (0.201-0.223); PPH: 0.203 ± 0.012 (0.19- 0.223); PNW: 0.457 ± 0.007 (0.444-0.475); TL: 0.497 ± 0.011 (0.48-0.52); TW: 0.108 ± 0.005 (0.101-0.117); PW: 0.165 ± 0.01 (0.145-0.179); PPW: 0.244 ± 0.01 (0.229-0.268); HI: 93.6 ± 1.5 (91.4-96.8); SI 1: 66.2 ± 1.2 (64.5- 68.9); EI: 54.1 ± 5.0 (45.3-63.2); SI 2: 70.7 ± 1.4 (68.8-73.4); MI: 179.8 ± 4.3 (171.0- 187.8); SPI: 154.6 ± 14.7 (136.6 -184.0); PI 1: 151.6 ± 5.2 (142.5-158.2); PI 2: 36.7 ± 1.9 (34.2-39.8); PPI 1: 110.6 ± 9.2 (97.1-126.3); PPI 2: 54.0 ± 1.6 (51.3-56.4); HTI 1: 75.3 ± 1.6 (72.6-77.6); HTI 2: 21.7 ± 1.05 (20.4-23.8). Head, thorax and abdomen black. Antennal scapes smoked black to brown on the apex, funiculus segments 1- 7 brown, 7-10 smoked black. Mandibles black to brown. Femora and tibiae smoked black to brown at the edges, tarsi brown (Figs. 58, 59, 60). Head rectangular, longer than wide, lateral surfaces below eyes straight, slightly rounded on the posterior edges (Fig. 60). Anterior margin of the clypeus shiny with longitudinal striae, slightly curved downward at the central part. Eyes elongate, strongly narrowing downward, reaching anteroventral margin of head, 0.3 times as long as length of the head. Scape short, 0.7 times as long as width of the head, at base 0.7 times as wide as in apex, gradually widened, slightly bent downward. Funiculus short, 1.6 times as long as scape, first segment elongate, triangular, 2.0 times as long as wide on apex, 2.6 times as long as second segment, length ratio of segments 100: 39: 28: 39: 44: 44: 56: 100: 111: 156, apical segments 1.9 times as wide as basal segments (Figs. 59, 60). Surface of the scape with very fine microsculpture, shiny, covered with long, dense, semierect setae. Promesonotum as long as wide, convex in profile. Promesonotal suture distinct, the border between dorsal and posterior surfaces of the promesonotum curved in profile view. Propodeum quadrate, 0.9 times as long as wide, propodeal spines short, triangular, the upper edge parallel to the dorsal surface of propodeum (Fig. 59). Petiole rounded with short peduncle, its anterior face straight, node rounded on dorsal surface in profile. Posterior face straight. Ventral margin of petiole smooth (Fig. 59). Postpetiole regularly rounded in profile. Postpetiole 0.9 times as long as wide in dorsal view, regularly widened from base to top, apical half with gently rounded sides (Fig. 59). Mandibles rounded, with outer and dorsal edges straight and smooth, inner margin with 7-8 teeth, the apical tooth massive and long. Clypeus on entire surface shiny, micropunctate with longitudinal striae. Frontal carinae short, extending to 1 / 3 length of eye; antennal fossa deeply impressed, microreticulate, frontal lobes rugulose with thick longitudinal striae, shiny between rugosities. Frons shiny, all surface with thick longitudinal striae and dense rugulose. Area above eyes shiny with thick longitudinal striae and dense rugulose. Ventral surface of the head with distinct striation, gena shiny, rugulose with fine striation (Figs. 59, 60). Entire head bearing setae, posterior margin with sparse erect setae directed forward, lateral surfaces of the head with appressed setae directed toward anterior margin, frontal area with dense appressed to semierect setae placed transversely, directed to the center of the head, ventral surface of the head with a prominent psammophore and appressed to erect long setae. Pronotum punctate with longitudinal striae, lateral surfaces strongly punctate with longitudinal striae. Dorsal suface of pronotum punctate with longitudinal striae, the central surface with striation and weak punctation or without punctation, shiny. Mesonotum strongly punctate on the dorsal surface, lateral surfaces punctate with longitudinal striae. Dorsal suface of propodeum strongly punctate to punctate, below spiracles strongly punctate with fine longitudinal striae at the posterior edge (Figs. 58, 59, 83). Dorsal suface of mesosoma with at least five long erect setae in theon anterior half, mesonotum and anterior propodeum with a few long setae. Base of petiole and postpetiole on the entire surface punctate, nodes of the petiole punctate on the dorsal and lateral surfaces, postpetiole node punctate, smooth with sparse punctation on the top, covered with several setae. Gaster shiny and shagreened, bearing dense, erect to semierect setae (Figs. 58, 59). Legs short, hind femora 0.7 times as long as mesosoma length, hind tibia 0.9 times as long as hind femora, hind tarsi 1.3 times as long as hind femora. Dorsal surface of femora with short, sparse, semierect setae, inner margin with a row of the sparse, long, semierect setae, tibiae bearing long, semierect setae on the entire surface, inner margins with a row of long semierect setae (Fig. 59). Biological data. Nothing is known about the biology of this species. In addition to the types, the following ant species were collected at the type locality: Aphaenogaster sporadis Santschi, Aphaenogaster subterraneoides Emery, Cataglyphis nodus (Brullé), Camponotus baldaccii Emery, Crematogaster ionia Forel, Lepisiota frauenfeldi (Mayr), Messor wasmanni Krausse, Plagiolepis pygmea sensu Radchenko, Tetramorium cf. semilaeve. Distribution. Greece: Dodecanese; Turkey. Differential diagnosis. Worker. Oxyopomyrmex polybotesi belongs to the species group that has a distinctly punctate propodeum. It is distinguished from O. emeryi and O. oculatus by the longitudinal striae covering whole frontal surface of the head and the presence of the rugosity and longitudinal striae on dorsal surface of the pronotum; from O. negevensis it differs in the lack of transverse striation on dorsal surface of the pronotum, the shiny abdomen and the more dense and thicker hairiness at the scape and legs. At first glance O. polybotesi is very similar to O. pygmalioni but it differs in following features: the shiny gena without rugosity (while O. pygmalioni has the gena dull and distinctly rugose), the occurrence of distinct longitudinal striae on the lateral surfaces of the pronotum (in O. pygmalioni the lateral surfaces of the pronotum are rugose with a slight longitudinal striae) and the dorsal surface of pronotum is shinier, weaker rugulose with longitudinal striae at the posterior edge (in O. pygmalioni the dorsal surface of pronotum is distinctly rugulose to rugose, and striation,if present, is oblique).Published as part of Sebastian Salata & Lech Borowiec, 2015, A taxonomic revision of the genus Oxyopomyrmex André, 1881 (Hymenoptera: Formicidae), pp. 1-66 in Zootaxa 4025 (1) on pages 50-53, DOI: 10.11646/zootaxa.4025.1.
Going Beyond Counting First Authors in Author Co-citation Analysis
The present study examines one of the fundamental aspects of author co-citation analysis (ACA) - the way co-citation
counts are defined. Co-citation counting provides the data on which all subsequent statistical analyses and mappings
are based, and we compare ACA results based on two different types of co-citation counting - the traditional type that
only counts the first one among a cited work's authors on the one hand and a non-traditional type that takes into
account the first 5 authors of a cited work on the other hand. Results indicate that the picture produced through this non-traditional author co-citation counting contains more coherent author groups and is therefore considerably clearer. However, this picture represents fewer specialties in the research field being studied than that produced through the traditional first-author co-citation counting when the same number of top-ranked authors is selected and analyzed. Reasons for these effects are discussed
Variations on the Author
“Variations on the Author” discusses two of Eduardo Coutinho’s recent films (Um Dia na Vida, from 2010, and Últimas Conversas, posthumously released in 2015) and their contribution to the general question of documentary authorship. The director’s filmography is characterized by a consistent yet self-effacing form of authorial self-inscription: Coutinho often features as an interviewer that rather than express opinions propels discourses; an interviewer that is good at listening. This mode of self-inscription characterizes him as an author who is not expressive but who is nonetheless markedly present on the screen. In Um Dia na Vida, however, Coutinho is completely absent form the image, while Últimas Conversas, on the contrary, includes a confessional prologue that moves the director from the margins to the center of his films. This article examines the ways in which these works stand out in the filmography of a director who offers new insights into the notion of cinematic authorship
Tetramorium atlante Cagniant 1970, new status
Tetramorium atlante Cagniant, 1970 new status (Figs 1 –13) Tetramorium caespitum st. punicum var. atlantis Santschi, 1918: 155 (terra typica: Tunisia), unavailable name. Tetramorium semilaeve subsp. atlante Cagniant, 1970: 430, 1997: 98. Material examined. Type material: syntype work- er on photo (AntWeb resources: Photo by Alexandra Westrich | URL: https://www.antweb.org/specimen/ CASENT0915045; accessed 1 April 2015): T. caespitum L. | st. punicum Sm | var. atlantis Sants | Type || Kairouan | (Santschi) || Tunisie | Kairouan | Dr. F. Santschi || 135 || Sammlung | Dr. F. Santschi | Kairouan || Type || NATURHIST. | MUSEUM | BASEL || ANTWEB | CASENT | 0 915045 (NHMB). Other material examined. 4 gynes, 5 males, 10 workers: Morocco | Meknès-Tafilale | Aïn Vittel n. Ifrane, 1600 m | 35.5409 ° / - 5.113 °, 10 VII 2010 | C. Galkowski || Collection L. Borowiec | Formicidae | LBC-MOR00010 (DBET). Redescription. Worker (Figs 1–3). Measurements and indices (n=10): CL: 0.77 ± 0.03 (0.737–0.844); POC: 0.3 ± 0.02 (0.268–0.324); CW: 0.755 ± 0.04 (0.704– 0.866); FR: 0.27 ± 0.017 (0.257–0.313); FL: 0.271 ± 0.016 (0.257–0.313); SL: 0.535 ± 0.02 (0.506–0.575); OMD: 0.192 ± 0.015 (0.17–0.212); EL: 0.146 ± 0.08 (0.128–0.291); EH: 0.098 ± 0.01 (0.078–0.101); ML: 0.827 ± 0.039 (0.782–0.927); SPSP: 0.148 ± 0.013 (0.112–0.17); SPL: 0.106 ± 0.011 (0.089–0.126); PEL: 0.191 ± 0.014 (0.179–0.229); NOL: 0.153 ± 0.011 (0.134–0.179); PPL: 0.176 ± 0.007 (0.162–0.19); PEH: 0.25 ± 0.013 (0.235–0.282); NOH: 0.177 ± 0.016 (0.165–0.223); PPH: 0.243 ± 0.011 (0.229–0.268); MW: 0.471 ± 0.024 (0.441–0.531); PEW: 0.235 ± 0.013 (0.215–0.267); PPW: 0.261 ± 0.01 (0.246–0.285); CS: 0.763 ± 0.035 (0.721–0.855); EYE: 0.16 ± 0.01 (0.138– 0.167); CL/CW: 1.021 ± 0.02 (0.975–1.047); FR/CS: 0.353 ± 0.009 (0.341–0.367); FL/FR: 1.009 ± 0.01 (1.0–1.043); SL/CS: 0.703 ± 0.016 (0.672–0.741); MW/CS: 0.618 ± 0.008 (0.6–0.632); PEW/PPW: 0.902 ± 0.03 (0.857– 0.937); NOH/NOL: 1.163 ± 0.08 (1.057–1.336); NOH/ PEL: 0.928 ± 0.05 (0.852–1.0); NOL/PEL: 0.801 ± 0.04 (0.729–0.872); PEH/NOL: 1.637 ± 0.08 (1.538–1.722); PEW/PEH: 0.943 ± 0.03 (0.914–0.979); CS/PEW: 3.245 ± 0.07 (3.14–3.351); CS/PPW: 2.927 ± 0.08 (2.816– 3.065); CW/MW: 1.619 ± 0.02 (1.582–1.666). Small to medium size, CS 0.763 [0.721–0.855]. Whole body brown to dark brown and appendages yellow, the palest specimens never yellow. Head nearly square CL/CW 1.021 [0.975–1.047], with almost parallel to slightly rounded sides, straight or slightly concave occipital margin and regularly rounded occipital corners. Eyes small, EYE 0.16 [0.138–0.167]. Frons moderately wide, FR/CS 0.353 [0.341–0.367], frontal lobes as wide as frons, FL/FR 1.009 [1.0–1.043]. Scape short, SL/CS 0.703 [0.672–0.741], without dorsal carina basally, surface smooth and shiny. Promesonotal dorsum slightly convex, metanotal groove shallow, but distinct. Propodeal teeth short, spiniform, apex of spine located approximately at 2/3 height of mesosoma (Fig. 2). Dorsal surface of petiole flat, NOH/NOL 1.163 [1.057– 1.336], petiole relatively high, PEH/NOL 1.637 [1.538–1.722], postpetiole distinctly transverse. General appearance moderately rugose, ground surface finely sculptured. Head dorsum mostly longitudinally rugose and shiny between rugae, rugae extend occipital margin of head, occiput mostly smooth and shiny, sides in anterior half longitudinally rugose and shiny between rugae. In most specimens short band without rugosities runs between frontal rugose area and rugosities along ocular area on each side (Fig. 3) but smooth area never exceeds 1/7 length of anterior surface of head; in extremely sculptured specimens almost entire frontal surface of head with long rugae with very small smooth patch between interrupted rugae and occiput with fine rugosities. Mesosoma dorsum longitudinally rugose and microreticulate but never reticulate, only occasionally rugae on pronotum partly interrupt- ed with indistinct microreticulation but pronotum nev- er with distinct smooth and shiny areas (Fig. 1). Sides of pronotum and meso- and metapleuron usually coarsely microreticulate, sometimes reticulation tends to form transverse lines but surface never appears striate or rugose (Figs 2). Dorsum of petiolar node smooth and shiny with sides carinate, lateral surface microreticulate. Dorsum of postpetiole smooth and shiny, sides microreticulate. First gastral tergite smooth and shiny. Whole dorsum, including head, covered with sparse setae, the longest on pronotum and the shortest on frons. Ventral surface of head with sparse short and 2–3 moderately long setae not forming a psammophore. Gyne (Figs 4–7). Measurements and indicates (n=3): CL: 1.05 ± 0.013 (1.039–1.065); POC: 0.388 ± 0.013 (0.374–.4); CW: 1.17 ± 0.02 (1.148–1.187); FR: 0.409 ± 0.007 (0.4–0.413); FL: 0.389 ± 0.01 (0.379– 0.398); SL: 0.727 ± 0.022 (0.704–0.749); OMD: 0.21 ± 0.016 (0.201–0.229); EL: 0.287 ± 0.007 (0.279–0.291); EH: 0.227 ± 0.02 (0.212–0.246); ML: 1.849 ± 0.001 (1.848–1.85); SPSP: 0.294 ± 0.02 (0.268–0.313); SPL: 0.216 ± 0.006 (0.212–0.223); PEL: 0.29 ± 0.02 (0.279– 0.313); NOL: 0.214 ± 0.008 (0.207–0.223); PPL: 0.315 ± 0.011 (0.302–0.324); PEH: 0.458 ± 0.011 (0.446–0.469); NOH: 0.287 ± 0.025 (0.257–0.302); PPH: 0.436 ± 0.007 (0.425–0.446); MW: 1.125 ± 0.01 (1.11–1.135); PEW: 0.39 ± 0.01 (0.38–0.4); PPW: 0.488 ± 0.02 (0.475–0.508); CS: 1.11 ± 0.014 (1.094–1.12); EYE: 0.232 ± 0.012 (0.224–0.246); CL/CW: 0.898 ± 0.015 (0.88–0.907); FR/CS: 0.368 ± 0.01 (0.357–0.378); FL/FR: 0.953 ± 0.04 (0.918–0.995); SL/CS: 0.655 ± 0.014 (0.644–0.671); MW/CS: 1.014 ± 0.09 (0.995–1.032); PEW/PPW: 0.801 ± 0.014 (0.787–0.815); NOH/NOL: 1.345 ± 0.168 (1.152– 1.459); NOH/PEL: 0.989 ± 0.08 (0.921–1.082); NOL/PEL: 0.74 ± 0.07 (0.661–0.799); PEH/NOL: 2.139 ± 0.03 (2.103–2.16); PEW/PEH: 0.853 ± 0.04 (0.83–0.897); CS/ PEW: 2.845 ± 0.106 (2.734–2.946); CS/PPW: 2.278 ± 0.12 (2.152–2.357); CW/MW: 0.987 ± 0.02 (0.967–1.005); WAIST: 0.792 ± 0.035 (0.764–0.83). Moderate size, CS 1.11 [1.094–1.12]. Head and mesosoma dark brown, abdomen brown, appendages yellowish. Head wider than long, CL/CW 0.898 [0.88– 0.907], with straight subparallel sides, shallowly emarginate occipital margin and regularly rounded occipital corners. Frons moderately wide, FR/CS 0.368 [0.357– 0.378], frontal lobes as wide as frons, FL/FR 0.953 [0.918–0.995]. Scape short, SL/CS 0.655 [0.644–0.671], without dorsal carina basally, smooth and shiny. Head as wide as scutum, MW/CS 1.014 [0.995–1.032]. Propodeal teeth short, triangular. Dorsal crest of petiolar node in frontal view slightly convex. Petiolar node dorsum steeply rounded backward. Petiole and postpetiole relatively narrow, WAIST 0.792 [0.764–0.83]. General appearance partly sculptured. Head dorsum, occiput and sides distinctly rugulose, ground surface shiny or indistinctly microreticulate. Frons longitudinally rugulose (Fig. 4). Mesosoma flat, pronotal sides visible from above. Anterior margin, sides, and median part of scutum smooth and shiny, lateral to shiny medi- an band punctate and longitudinally striate, striae extending from base to 2/3 length of scutum, in the strongest sculptured specimens median smooth band very narrow and whole base of scutum striate. Scutellum mostly smooth and shiny, only corners or corners and sides with oblique striation and punctate (Fig. 5). Sides of pronotum ruguloso-reticulate and feebly microreticulate, anepisternum in upper part with fine longitudinal rugae and rest smooth and shiny, katepisternum mostly smooth and shiny but posterior corners and sometimes whole posterior third with fine longitudinal rugae and microreticulation (Fig. 7). Whole surface of petiolar node distinctly reticulate, posterior surface granulate without or with indistinct transverse rugae. Postpetiole distinctly transverse, sides rounded (Fig. 7), dorsum of postpetiole smooth, sides granulate and reticulate. First gastral tergite smooth and shiny. Whole dorsum, including head, covered with short, sparse setae. Ventral surface of head with several short setae, as long as to 1.5 times longer than frontal setae, arising posteriorly to buccal cavity. Male (Figs 8 –13). Measurements and indicates (n=5): CL: 0.694 ± 0.013 (0.673–0.715); POC: 0.284 ± 0.01 (0.268–.302); CW: 0.836 ± 0.014 (0.81–0.854); FR: 0.258 ± 0.009 (0.246–0.274); FL: 0.292 ± 0.006 (0.285– 0.302); SL: 0.311 ± 0.008 (0.302–0.324); OMD: 0.076 ± 0.007 (0.061–0.084); EL: 0.315 ± 0.008 (0.302–0.324); EH: 0.258 ± 0.005 (0.249–0.263); ML: 1.81 ± 0.113 (1.617–1.935); SPSP: 0.243 ± 0.01 (0.235–0.263); SPL: 0.216 ± 0.011 (0.201–0.232); PEL: 0.32 ± 0.011 (0.302– 0.335); NOL: 0.244 ± 0.019 (0.212–0.268); PPL: 0.304 ± 0.018 (0.279–0.324); PEH: 0.333 ± 0.01 (0.313–0.346); NOH: 0.206 ± 0.015 (0.19–0.235); PPH: 0.464 ± 0.029 (0.425–0.503); MW: 1.108 ± 0.049 (1.067–1.2); PEW: 0.36 ± 0.005 (0.351–0.366); PPW: 0.518 ± 0.02 (0.492– 0.547); CS: 0.765 ± 0.014 (0.742–0.785); EYE: 0.374 ± 0.002 (0.372–0.379); CL/CW: 0.831 ± 0.004 (0.824– 0.837); FR/CS: 0.337 ± 0.01 (0.32–0.349); FL/FR: 1.134 ± 0.06 (1.044–1.228); SL/CS: 0.407 ± 0.009 (0.395– 0.421); MW/CS: 1.449 ± 0.06 (1.389–1.565); PEW/PPW: 0.696 ± 0.03 (0.655–0.744); NOH/NOL: 0.85 ± 0.093 (0.739–0.955); NOH/PEL: 0.644 ± 0.05 (0.6–0.725); NOL/PEL: 0.762 ± 0.044 (0.702–0.821); PEH/NOL: 1.372 ± 0.08 (1.25–1.476); PEW/PEH: 1.084 ± 0.05 (1.014– 1.169); CS/PEW: 2.185 ± 0.06 (2.042–2.191); CS/PPW: 1.478 ± 0.06 (1.402–1.561); CW/MW: 0.691 ± 0.027 (0.639–0.719). Whole body dark brown, appendages yellowish. Head behind eyes almost trapezoidal, occipital margin slightly convex, occipital corners subangulate. Scutum distinctly wider than head. Propodeum in profile with distinct angulation in position of propodeal teeth. Dorsal crest of petiolar node with obtuse transversal edge, slightly emarginated in frontal view. Head distinctly granulate, dull. Sides of pronotum microreticulate with fine transverse striation. Scutum between sutures in anterior part with oblique striation and microreticulate only along middle runs shiny band, laterally smooth and shiny, rest microreticulate with longitudinal and oblique striation. Scutellum at base and laterally with longitudinal striation, rest smooth and shiny. Anepisternum in anterior half smooth and shiny in posterior half with oblique striation, katepisternum in both dorsal angles distincty striate and microreticulate. Whole surface of petiolar node microgranulate and microreticulate, dull, postpetiole mostly microreticulate only top partly smooth and shiny. Whole surface of propodeum microreticulate, dull. First gastral tergite smooth and shiny. Male genitalia stout (Figs 10 –13), in lateral distinctly slightly constricted before apex with obtuse inner angle, top shortly and sparsely pubescent, ventral and dorsal margins of parameres shallowly incised, top of inner margin of paramere before apical denticle straight, without dentiform plate extending beyond the sharp edge of paramere (Fig. 13). Differental diagnosis. Tetramorium atlante and T. semilaeve at first glance look similar, and workers are especially difficult to identify. Males and gynes, however, have constant differences in the morphological characters and biometric data (see Table 1). Workers of T. atlante differ from workers of T. semilaeve (in parentheses characters for T. semilaeve) in brown to dark brown body (usually pale yellow to yellowish brown, occasionally brown), head sculpture with longitudinal rugosities, more distinct spread over the frontal surface with very small lateral areas without striation (rugosities less distinct on sides of frontal surface, usually with longitudinal areas without striation). Nevertheless, dark and strongly sculptured workers of T. semilaeve are extremely similar to T. atlante and a proper identification requires a nest sample with sexual forms. Workers of T. depressum Forel and T. punctatum Santschi, two other well-defined species of T. semilaeve complex, differ in reduced sculpture of head with at least half frontal surface smooth and shiny. Gynes of T. atlante distinctly differ from gynes of T. semilaeve (characters for T. semilaeve in parentheses) in larger size with ML 1.848–1.850, MW 1.11– 1.35, CW 1.148–1.187 and SPL 0.212–0.223 (1.053– 1.813, 1.0–1.14, 0.978–1.161 and 0.173–0.201 ), scutum at base and laterally with striation extending from base to at least half length of scutum (scutum completely smooth and shiny or with short striation at base never extending to half length of scutum), scutellum with distinct striation in angles (completely smooth and shiny or with rudiments of striation), anepisternum in posterior half with distinct striation and microreticulation (mostly smooth and shiny), katepisternum along posterior margin with broad area of distinct sculpture (narrow area or reduced to posterior corners), postpetiole less transverse and more sculptured with smooth and shiny area reduced to the middle of top (postpetiole more transverse, less sculpture with whole dorsal surface smooth and shiny). Males of T. atlante distinctly differ from males of T. semilaeve in higher petiole with NOH 0.19–0.235 and PEH 0.313–0.346 (0.156–0.179 and 0.257–0.313), lower SL/CS ratio 0.395–0.421 (0.436–0.478), head behind eyes trapezoidal (rounded), scutum between sutures mostly striate and microreticulate (mostly smooth and shiny), scutellum at whole base microreticulate and striate (completely smooth and shiny or in basal corners with fine sculpture), propodeum slightly angulate (round), anepisternum in posterior half with oblique striation (smooth and shiny, at most with few striae close to upper margin), katepisternum in both upper corners with broad microreticulate and striate area (narrow sculptured areas), male genitalia in lateral view distinctly constricted before apex (shallowly constricted) and top of inner margin of paramere without dentiform plate extending beyond the sharp edge of paramere (with dentiform plate). Biological notes. Although locus typicus Kairouan in Tunisia is placed only 70 m a.s.l. Cagniant (1970, 1997) noted than T. atlante prefers mountainous habitats. Its known localities are mostly between 1600 m (our material) to 2300 m (Haut Atlas). Ants were collected in agricultural habitat such as farmlands, pastures, and in scrubs, luminous forests, roadsides. Preference for mountain habitats additionally differentiates T. atlante from T. semilaeve, which, according to Borowiec et al. (2015), prefers littoral habitats located usually below 700 m a.s.l. and only occasionally reach to the elevation of just over 1000 m a.s.l.Published as part of Lech Borowiec & Christophe Galkowski, 2016, REDESCRIPTION OF TETRAMORIUM ATLANTE CAGNIANT, 1970, NEW STATUS (HYMENOPTERA: FORMICIDAE: MYRMICINAE), pp. 43-52 in Annales Zoologici 66 (1) on pages 44-50, DOI: 10.3161/00034541ANZ2016.66.1.003, http://zenodo.org/record/26954
Oxyopomyrmex insularis Santschi 1908
Oxyopomyrmex insularis Santschi, 1908 (Figs. 4, 5, 6, 7, 8, 9, 74) Oxyopomyrmex insularis Santschi, 1908: 523, fig. 7 (w.m.); Wheeler 1927: 105; Wellenius 1955: 6; Báez & Ortega 1978: 190; Barquín 1981: 103. Oxyopomyrmex insularis var. major Santschi, 1923: 326; Wheeler 1927: 105; Wellenius 1955: 6; Báez & Ortega 1978: 190; Barquín 1981: 103 (as syn. of O. insularis). Type locality: Medano, Tenerife, Canary Is, Spain. Type material. Lectotype worker (present designation): O. insularis | Sants || type || Tenerife | Medano | 3.1. 1906 || Sammlung | Dr. F. Santschi | Kairouan || ANTWEB | CASENT | 0 913243 (NHMB),; 1 paralectotype worker: red label | Tenerife | Medano | 3.1. 1906 || Sammlung | Dr. F. Santschi | Kairouan (NHMB); 1 paralectotype worker: O. insularis | Sants || red label || Tenerife | Medano | 3.1. 1906 || TENERIFE | Medano | Cabrera.y. Diaz | 1.III. 1906 || Sammlung | Dr. F. Santschi | Kairouan (NHMB) 1 paralectotype male: the same data as lectotype (NHMB). Oxyopomyrmex insularis var. major: 1 worker — Oxyopomyrmex | insularis type | v. major | Sants. || Tenerife | El Medano | 3.1. 1906 | A. Cabrera || type || Sammlung | Dr. F. Santschi | Kairouan || ANTWEB | CASENT | 0 913244 (NHMB). Other material examined. 10 workers —SPAIN Tenerife, El | Médano, 10 m 28 ° 02’ | N/ 16 ° 32 ’ W, 13.12. 2002 | leg. X. Espadaler (SSC); 16 workers —Collection L. Borowiec | Formicidae | LBC-ES0043 || Oxyopomyrmex | insularis | det. L. Borowiec || SPAIN, Canary Is., Tenerife | Puertito de Güimar, 29 m | 28,31312 / 16,36161 | 1 I 2015, L. Borowiec (DBET); 3 workers —CANARY IS. | Tenerife | El Medano | 13.XII. 2000 | X. Espadaler (BMNH). Gyne. Unknown. Male. Redescription. Measurements (n= 1): HL: 0.603; HW: 0.581; SL: 0.436; EL: 0.246; EW: 0.179; ML: 1.52; MH: 0.976; PSL: 0.251; SDL: 0.223; PL: 0.469; PPL: 0.346; PH: 0.268; PPH: 0.279; PNW: 0.771; TL: 0.905; TW: 0.095; PW: 0.246; PPW: 0.358; HI: 96.4; SI 1: 72.3; EI: 72.8; SI 2: 75.0; MI: 197.1; SPI: 122.6; PI 1: 175.0; PI 2: 31.9; PPI 1: 124.0; PPI 2: 46.4; HTI 1: 61.2; HTI 2: 10.5. Whole body uniformly brown. Antennal scapes brown to yellowish brown at the apex, first segment of funiculus pale brown, segments 2–11 yellowish brown. Mandibles brown to yellowish brown. Legs brown to pale brown, tibiae pale brown, tarsi pale brown to yellowish brown (Figs. 4, 5, 6). Head oval, longer than wide, lateral surfaces below eyes and on the posterior edges gently rounded (Fig. 6). Anterior margin of the clypeus slightly convex in central part. Eyes oval, 0.4 times as long as length of the head. Ocelli large. Scape short, 0.8 times as long as width of the head, at base 0.75 times as wide as in the apex, straight. Funiculus short, 1.7 times as long as scape, the first segment elongate, triangular, 1.4 times as long as wide on the apex, 1.4 times as long as second segment, length ratio of segments: 100: 71: 57: 57: 64: 64: 78: 86: 100: 114: 207, apical segments as wide as the basal segments (Figs. 4, 5). Surface of the scape with very fine microsculpture, shiny, covered with short, dense, appressed to semierect setae. Mesosoma 2.5 times as long as head, relatively high and robust, very feeble convex in profile with rounded pronotal corners. Scutum 1.2 times as wide as long, posterior margin regularly semicircular. Propodeum located considerably lower than mesosomal plate, propodeal spines triangular, short with a wide base. Petiole rounded with a long peduncle, its anterior face slightly concave, node sharply rounded with deep cavity in the central part and two nodules at the outer edges, posterior face strongly concave. Ventral margin of the petiole straight, without lobe. Postpetiole regularly rounded in profile. Postpetiole regularly widened from base to top in the dorsal view, apical half with a gently rounded sides (Figs. 4, 5). Mandibles elongate with longitudinally striae, shiny, inner margin with 4-5 teeth, the apical tooth massive and long. Clypeus rugulose but shiny. Frontal carinae curved outward to merge with the rugae surrounding antennal sockets; antennal fossa impressed, shiny and rugulose, frontal lobes rugulose with thin longitudinal striae, shiny between rugosities. Frons rugulose with longitudinal striae in the central part to rugulose towards eyes, area above the eyes and ventral surface of the head rugulose, gena rugulose with longitudinal thin striae (Figs. 5, 6). Entire head bearing setae, posterior margin with dense, very long semierect to erect setae directed forward, lateral surfaces of the head with dense, long and semierect setae directed toward anterior margin, frontal area with dense, semierect to erect, long setae placed transversely, directed toward center of the head, ventral surface of the head with a prominent psammophore and appressed to erect long setae. Sides of the pronotum punctate to rugulose with thin longitudinal striae, anepisternum smooth and shiny with fine microreticulation, katepisternum smooth to microreticulate with fine longitudinal striae, metanepisternum and metakatepisternum rugulose with fine longitudinal striae. Scutum rugulose with fine longitudinal striae except three longitudinal stripes located at the centre and outer edges of the scutum which are smooth and shiny with sparse, very weak longitudinal striae. Central stripe wide, narrowing toward centre, reaching only half of the length of the scutum. Scutellum punctate with longitudinal striae at the outer surfaces, central part smooth and shiny (Figs. 4, 5). Propodeum on lateral surfaces punctate, area between propodeal spines shiny and smooth. Dorsal suface of the propodeum punctate with transverse striation, between and below the spines punctate to smooth and shiny. Dorsal suface of the mesosoma on anterior half with sparse, long, erect setae, anterior propodeum without setae. Base of petiole and postpetiole punctate on the entire surface, nodes of the petiole punctate on lateral surfaces, dorsal surface of the petiole node punctate, postpetiole node punctate, punctate to micropunctate on the dorsal surface, shiny. Gaster shiny with sparse micropunctation, bearing sparse, long, semierect to erect setae. Legs short, hind femora 0.7 times as long as mesosoma length, hind tibia 0.9 times as long as hind femora, hind tarsi 1.5 times as long as hind femora. Dorsal surface of the femora with several, short, semierect setae, inner margin with a row of sparse, semierect setae, tibiae covered with long, semierect setae on the entire surface, inner margins with a row of semierect setae (Fig. 5). Worker. Redescription. Measurements (n= 17): HL: 0.706 ± 0.026 (0.659-0.737); HW: 0.667 ± 0.03 (0.603- 0.704); SL: 0.524 ± 0.019 (0.492-0.558); EL: 0.254 ± 0.014 (0.229-0.282); EW: 0.135 ± 0.007 (0.123-0.145); ML: 0.879 ± 0.048 (0.771-0.961); PSL: 0.189 ± 0.01 (0.17-0.207); SDL: 0.125 ± 0.009 (0.109-0.145); PL: 0.341 ± 0.023 (0.302-0.369); PPL: 0.27 ± 0.027 (0.212-0.324); PH: 0.236 ± 0.014 (0.212-0.254); PPH: 0.238 ± 0.013 (0.218-0.257); PNW: 0.439 ± 0.027 (0.391-0.48); TL: 0.557 ± 0.032 (0.469-0.603); TW: 0.1 ± 0.006 (0.089-0.109); PW: 0.171 ± 0.009 (0.156-0.19); PPW: 0.265 ± 0.02 (0.232-0.313); HI: 94.4 ± 1.8 (91.5-98.3); SI 1: 74.0 ± 1.5 (71.6-77.1); EI: 53.1 ± 2.7 (47.9-58.5); SI 2: 78.2 ± 1.8 (75.9-82.2); MI: 201.7 ± 3.3 (196.8-208.5); SPI: 152.5 ± 8.9 (131.0- 164.3); PI 1: 144.5 ± 7.1 (130.9-156.6); PI 2: 39.4 ± 2.3 (36.1 -43.0); PPI 1: 113.7 ± 12.7 (95.1-146.6); PPI 2: 60.2 ± 2.2 (57.9-65.2); HTI 1: 83.2 ± 2.6 (77.8-88.5); HTI 2: 17.8 ± 1.4 (15.6-21.1). Head and abdomen black. Thorax black to dark brown. Antennal scapes black, apex of the scapes and funiculus brown to yellowish brown. Legs dark brown, knees and tarsi brown to yellowish brown. Mandibles dark brown to yellowish brown (Figs. 7, 8, 9). Head oval, longer than wide (Fig. 9). Anterior margin of the clypeus smooth and slightly curved in central part. Eyes elongate, gently narrowing downward, reaching anteroventral margin of head, 0.4 times as long as length of the head. Scape short, 0.7 times as long as width of the head, at base 0.6 times as wide as in apex, gradually widened, slightly bent downward. Funiculus short, 1.6 times as long as scape, first segment elongate, triangular, 2.2 times as long as wide on apex, 2.75 times as long as second segment, length ratio of segments 100: 36: 36: 36: 32: 45: 54: 72: 81: 163, apical segments 1.6 times as wide as basal segments (Figs. 7, 8). Surface of the scape with very fine microsculpture, shiny, covered with short and semierect setae. Promesonotum 1.1 times as long as wide, gently, regularly convex in profile. Promesonotal suture indistinct, in profile the border between dorsal and posterior surfaces of the promesonotum gently curved. Propodeum quadrate, 1.1 times as long as wide, propodeal spines triangular, rising obliquely upwards, peaks gently curved downward (Fig. 8). Petiole rounded with short peduncle, its anterior face slightly concaved, node angulated in profile. Posterior face slightly concave. Ventral margin of petiole with distinct ventral lobe (Fig. 8). Postpetiole in profile regularly rounded. In dorsal view postpetiole as long as wide, regularly widened from base to top, apical half with gently rounded sides (Fig. 7). Mandibles rounded, with outer and dorsal edges straight and smooth, inner margin with 7-8 teeth, the apical tooth massive and long. Clypeus on entire surface smooth and shiny. Frontal carinae short, extending to 1 / 3 length of eye, antennal fossa area deeply impressed, shiny and smooth, frontal lobes with thin longitudinal rugae, microreticulate but shiny between rugosities. Frons shiny, in central part with longitudinal striae, on sides with distinct reticulation. Area above eyes shiny and finely rugulose, ventral surface of the head with indistinct microreticulation or microgranulate, gena smooth and shiny (Figs. 8, 9). Entire head bearing setae, posterior margin with a few erect setae directed forward, lateral surfaces of the head with sparse appressed to semierect setae directed toward anterior margin, frontal area with sparse appressed to semierect setae placed transversely, directed to the center of the head, ventral surface of the head with a prominent psammophore and appressed to erect long setae (Fig. 9). Pronotum shiny, on the entire surface finely rugulose with longitudinal striae. Dorsal suface of pronotum shiny, finely rugulose with longitudinal or transverse striation at the outer surface, lateral surfaces finely rugulose with longitudinal striae. Mesonotum on entire surface with fine longitudinal striae, dorsal surface finely rugulose, shining, lateral surfaces with several longitudinal striae, on the posterior surface distinctly rugulose. Propodeum shiny, finely rugulose with longitudinal striae on the dorsal and lateral surfaces (Figs. 7, 8, 74). Dorsal suface of mesosoma on anterior half with at least 5 long, erect setae, mesonotum and anterior propodeum with a few erect to semierect setae. Base of petiole and postpetiole on the entire surface rugulose, nodes rugulose, on the dorsal surface shiny and microreticulate to smooth, covered with several sparse, setae. Gaster shiny, bearing erect setae. Legs short, hind femora 0.9 times as long as mesosoma length, hind tibia 0.7 times as long as hind femora, hind tarsi 1.3 times as long as hind femora. Dorsal surface of femora with short, sparse, appressed setae, inner margin with a row of sparse, long and semierect setae, tibiae covered with long, semierect pubescence on the entire surface, inner margins with a row of slightly long and more erect setae (Fig. 8). Biological data. Borowiec collected this species in the desert wadi located near El Médano. Despite the low temperature (15 ° C) and windy, arid climate of the wadi, which is vegetated by a sparse, halophilic vegetation, worker ants were active, and could be caught between clumps of saline herbs. Other ant species collected at the site were: Camponotus rufoglaucus feae Emery, Monomorium subopacum (Smith), Plagiolepis maura Santschi, Solenopsis canariensis Forel, and Tetramorium depressum Forel. Distribution. Spain: Canary Is. Differential diagnosis. Male. Oxyopomyrmex insularis is the only species known to have the posterior margin of the head distinctly rugulose and completely devoid of striation. Only the center of the frons has a few, fine and oblique wrinkles. These two features easily distinguish the male of O. insularis from other Oxyopomyrmex males. Worker. Oxyopomyrmex insularis belongs to a species group that has a longitudinal striae on the head limited to the center of the frons. Oxyopomyrmex insularis differs from O. emeryi and O. oculatus in lacking punctation on dorsal and lateral surfaces of the pronotum and by occurrence of smooth and shiny surface at the center of the dorsal surface of its pronotum and mesonotum; from O. nitidior it differs in the lack of rugosity on dorsal surface of its pronotum and the lack of shiny and smooth surface at the center of its frons.Published as part of Sebastian Salata & Lech Borowiec, 2015, A taxonomic revision of the genus Oxyopomyrmex André, 1881 (Hymenoptera: Formicidae), pp. 1-66 in Zootaxa 4025 (1) on pages 11-15, DOI: 10.11646/zootaxa.4025.1.
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