1,696 research outputs found
Four distinct leadership roles in sport (as defined by Fransen, Vanbeselaere, et al., 2014).
No full textFour distinct leadership roles in sport (as defined by Fransen, Vanbeselaere, et al., 2014).</p
Automatic morphological parsing of Old Irish verbs using finite-state transducers
No full textThe topic of this paper constitutes the main part of a recently finished Ph.D. project carried out by the author which investigates how computational methods can be employed to map cognate verb forms in Early Irish (ca. 7th–12th centuries A.D.) and Modern Irish (ca. 1200 onwards). This paper discusses the development of a finite-state morphological transducer using foma (Hulden, 2009) for the Old Irish language (ca. 7th–9th centuries A.D.), focusing on verbs. Two main challenges are discussed. First, different practices of word segmentation have significant repercussions for the encoding of dependencies both on and beyond the word level. A second challenge is complex verb stem formation and considerable stem allomorphy. This has been tackled by operating with “monolithic stem” entries for each verb lemma, i.e., synchronic, invariable hard-coded stems, representing a semi-surface-level base form
A new stygobiont species of Halocaridinides Fujino & Shokita, 1975 (Crustacea, Decapoda, Caridea, Atyidae) from caves on Socotra Island (Yemen), with notes on the genus
No full textFransen, Charles H.J.M., Damme, K. Van (2018): A new stygobiont species of Halocaridinides Fujino & Shokita, 1975 (Crustacea, Decapoda, Caridea, Atyidae) from caves on Socotra Island (Yemen), with notes on the genus. Zootaxa 4442 (2): 241-261, DOI: 10.11646/zootaxa.4442.2.
A medicine for the Archduchess of Innsbruck
No full textBy Sietske Fransen, with Saskia Klerk. Two months ago Saskia Klerk discussed a recipe for the breaking of a bladder stone. It seems that the author of manuscript BPL3603 included this recipe into his collection because of the wonderful curative properties it proved to possess according to the eyewitness accounts documented in the text. On pages 117 and 118 of the same manuscript we find an ‘Excellent recipe against all ailments and diseases that have their origin in corrupt blood and bad humo..
An Anthology of Early British Motorcycle Travel Literature
No full textCollaborative book project in association with the International Journal of Motorcycle Studies (IJMS) and Riders for Health.
Along with a foreword by Steven E. Alford and Suzanne Ferriss, this volume contains three early twentieth-century British motorcycle travel narratives : Captain W. H. L. Watson’s Adventures of a Despatch Rider (1915), Lady Warren’s Through Algeria & Tunisia on a Motor-bicycle (1922) and C. K. Shepherd’s Across America by Motor-Cycle (1922). Interactive colour maps are available at :
Additionally, this publication follows a social enterprise model employed in a previous motorcycle travel project entitled Essex-Dakar with all profits helping support notable causes, in this case, Riders for Health.
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Book Review : Goodmann, T. (2009) The Road Worst Traveled. International Journal of Motorcycle Studies. [Internet]. vol.5, Issue 2: Fall. Available at : <http://ijms.nova.edu/Fall2009/IJMS_Rvw.Goodmann.html
Publication : Bart Fransen et Cyriel Stroo, "The Ghent Altarpiece. Research and Conservation of the Exterior", Bruxelles, Institut du Patrimoine artistique, 2020
No full textThe Ghent Altarpiece. Research and Conservation of the Exterior Sous la direction de Bart Fransen and Cyriel Stroo Avec les contributions de A.-S. Augustyniak, Chr. Ceulemans, A. Coudray, D. Deneffe, L. Depuydt, B. Devolder, H. Dubois, B. Fransen, A. Genbrugge, J.-A. Glatigny, K. Janssens, S. Jones, J. Ketels, L. Mortiaux, N. Laquière, M. Martens, Cl. Mehagnoul, M. Postec, J. Reyniers, Fr. Rosier, J. Sanyova, M. H. Smith, R. Spronk, Gr. Steyaert, C. Stroo, P. Vandenabeele, G. Vandersnickt, A...
Automatic morphological analysis and interlinking of historical Irish cognate verb forms
No full textThe main aim of the author’s research project is to use computational approaches
to gain more insight into the historical development of Irish verbs. One of the objectives is to investigate how a link between the electronic Dictionary of the Irish
language (eDIL),1 covering the period c. 700–c. 1700, but focussing on Early Irish
(7th–12th centuries), and the nascent Foclóir Stairiúil na Gaeilge ‘The Historical
Dictionary of Irish’,
2 covering the period 1600–2000, could be implemented. Such
a link will be hugely beneficial for scholars operating at the intersection of the medieval and modern period (see Table 1), who currently lack a comprehensive lexical resource for the “intermediate” early modern period.This paper stems from research carried out during a
Government of Ireland Postgraduate Scholarship (GOIPG/2017/1808) funded by
the Irish Research Council. The author would also like to acknowledge the
anonymous reviewer for helpful feedback and the editors for seeing this publication through.Peer reviewe
Nippontonia christellae Fransen, 2013, spec. nov.
No full textNippontonia christellae spec. nov. (figs 1–9) Material examined. 1 ovigerous female holotype (pocl. 4.75 mm), RMNH.CRUS.D. 53861; 1 male paratype (pocl. 3.80 mm), RMNH.CRUS.D. 53837; 1 ovigerous female paratype (pocl. 4.81 mm), RMNH.CRUS.D. 53841: stn. SEM. 18: Malaysia, Sabah, Semporna area, Ligitan Island, Ligitan 4, 04014'06.5"N 118048 '26,5"E; 4.xii. 2010; 15 m depth; in the sponge Acanthostrongylophora ingens (Thiele, 1899) (id. N.J. de Voogd); collected by C.H.J.M. Fransen; photo 533, 540 – 552. Material for comparison. Nippontonia minirostris Bruce & Bauer, 1997. 1 paratype non-ovigerous female (pocl. 2.4 mm), RMNH.CRUS.D. 47746; Stn YMP- 1231, Japan, Ryukyu Is., Kerama group, Aka-jima, Nishihama; 22.iv. 1994; depth 15 m; in unidentified black sponge; collected by K. Nomura. 2 ovigerous females (pocl. 2.6, 2.7, and 2.8 mm), 2 males (pocl. 2.2 and 3.0 mm), and 1 juvenile (pocl. 1.8 mm), OUMNH 2010 -02-0056; 1 ovigerous female (pocl 2.4 mm), RMNH.CRUS.D. 54442: Taiwan, Green Island, General Rock, 22 º 40.567 ’N 121 º 23.618 ’E; from big black sponge; depth 10–15 m; 21.vii. 2009; leg. T. Naruse TAI 0 24. Description. Medium-sized shrimp (fig. 1) with subcylindrical, oblong body shape. Carapace smooth (fig. 1), glabrous. Rostrum (figs. 1, 2A, B) very short, not reaching beyond eyes, laterally compressed, triangular in dorsal view, with one small dorsal obtuse tooth, unarmed ventrally, lateral carina not developed. Supra-orbital, epigastric, and hepatic teeth absent. Minute antennal tooth present or absent. Orbit obsolescent. Inferior orbital angle (fig. 2 A) slightly produced, broadly rounded in dorsal view. Pterygostomial angle (fig. 2 A) strongly produced, rounded. Abdominal segments (fig. 1) smooth. Third segment not produced posterodorsally. Pleura all broadly rounded. Sixth segment as long as fifth, posteroventral angle feebly produced, posterolateral angle feebly acute. Telson (fig. 2 C) 1.5 times as long as sixth abdominal segment and 1.6 times longer than anterior width; lateral margins slightly converge posteriorly; two pairs of large sized submarginal dorsal spines present at 0.1 and 0.4 of telson length; posterior margin broadly rounded, about 0.66 of anterior width, with three pairs of spines. Lateral spines short, about third of dorsal spines. Intermediate spines well developed, about twice as long as lateral spines, 1.3 times length of submedian spines. Eyes (fig. 2 A, B) well developed. Large cornea globular, obliquely set on stalks, without accessory pigment spot. Eyestalks more than twice as long as proximal width, not swollen proximally in dorsal view. Antennular peduncle (fig. 2 D) slender, about twice as long as eyes. Proximal segment long, slender, 3.6 times longer than wide; stylocerite slender, acute, reaching to third of segment; lateral margin slightly convex, anterolateral margin not produced, with one strong distolateral tooth, distodorsal margin with distinct protrusion with row of setae; medial ventral margin with small but distinct acute tooth at mid length of segment. Statocyst normally developed, without statolith. Intermediate and distal segments short, together equal to 0.34 of proximal segment length. Upper flagellum biramous, with the first 7–8 segments fused. Shorter free ramus uni-segmented, longer free ramus with about 10 segments. Aesthetascs present in distal part of fused rami and uni-segmented short free ramus. The lower flagellum slender, about as long as the longer free ramus of the upper flagellum. Antennal basicerite (fig. 2 E) without lateral tooth. Ischiocerite and merocerite normal. Carpocerite slender, almost reaching to mid length of scaphocerite. Scaphocerite long, slender, with lamella strongly reduced, without setae. Lateral border slightly convex, ending in a very long robust acute distolateral tooth. Thoracic sternites very narrow, unarmed. Mandible (fig. 3 A, B) with cylindrical slender, strongly reduced molar process without any armature, distally two-segmented; incisor process distally expanded with row of about 20 posteromedial teeth; without palp. Maxillula (fig. 3 C) with bilobed palp; upper lacinia broad, furnished with several rows of stout simple and serrulate setae along median margin; lower lacinia short with long serrulate setae distally. Maxilla (fig. 3 D) with stout simple palp with one proximal seta. Basal endite well developed, not bilobed, broad with about 18 simple setae along anterior and medial margins. Coxal endite not developed, median region slightly convex. Scaphognathite as usual for the genus. First maxilliped (fig. 4 A) with simple non-setose palp. Basal endite well developed, subrectangular, distinctly separated from coxal endite, fringed medially with fine setulose and simple setae. Coxal endite convex with fine setulose and simple setae medially. Caridean lobe distinct but not well developed, with plumose marginal setae. Flagellum of exopod short with few lateral and distal plumose setae. Epipod well developed, indistinctly bilobed. Second maxilliped (fig. 4 B) with rectangular dactylar segment, about 4 times longer than wide, slightly convex medially, bearing row of stout biserrulate spines. Propodal segment longer than dactylar segment, with rounded distomedial angle with long serrulate setae in medial half. Carpus short and merus normal. Ischium completely fused to basis. Basis with long exopod with plumose setae in distal fifth. Coxa slightly produced medially, with oblong epipod laterally. Third maxilliped (fig. 4 C) with moderately broad antepenultimate segment, about 3 times longer than proximal width. Basis completely fused with ischiomerus. Medial margin with simple setae. Penultimate segment slender, 3.8 times longer than wide, 0.39 of length of antepenultimate segment, with rows of long, robust setulose setae on medial border. Terminal segment 3.0 times longer than wide, almost half length of penultimate segment, with long, robust serrulate and simple setae medially and distally. Exopod long, just overreaching distal margin of antepenultimate segment; with long plumose setae in distal third. Coxa not produced medially, lateral plate well developed, convex; oblong epipod laterally. Without arthrobranch. First pereiopod (fig. 4 D) moderately robust, overreaching scaphocerite by chela. Chela with palm subcylindrical, slightly bowed, about twice as longer as wide. Fingers (fig. 4 E) 0.8 of palm length, subspatulate, with brushes of setae. Cutting edges poorly developed, entire. Tip of dactylus and fixed finger tridentate, central tooth robust, articulate. Cleaning setae present proximally on palm and distoventral end of carpus. Carpus 1.4 times length of chela, 5.0 times longer than wide. Merus about as long as carpus, 1.4 times as long as ischium. Basis normal. Coxa with medial setose process. Second pereiopods (fig. 5) large, symmetrical in shape, unequal in size. Major cheliped (fig. 5 A) with ischium 0.64 times length of merus, distally armed with few small acute tubercles. Merus as long as carpus, 2.5 times longer than central depth, with minute, acute tubercles along medial margin. Carpus 0.66 of palm length, strongly excavate dorsally. Palm subcylindrical, somewhat compressed, unarmed, strongly swollen posteriorly, 1.5 times longer than proximal depth. Fingers (fig. 5 B, C) 0.62 of palm length, cutting edges entire, tips strongly hooked, with several brushes of setae. Minor cheliped (fig. 5 D) as major cheliped. Fingers of chela (fig. 5 E) as long as palm. Dactylus slender, with medial longitudinal carina, tip strongly hooked, distal 2 / 5 th of cutting edge entire, proximal 3 / 5 th with minutely tuberculate surface. Fixed finger oblong triangular with distal fifth of cutting edge entire, proximal 4 / 5 th of cutting edge with minutely tuberculate surface, tip strongly hooked. The ambulatory pereiopods (fig. 6) relatively robust, similar. Third pereiopod (fig. 6 A) reaching with dactylus to distal margin of antennular peduncle. Dactylus (fig. 6 B, C) short, about twice as long as proximal depth, strongly recurved, slightly compressed, uniformly tapering, with distinct slender unguis; with row of 4–6 obtuse teeth in middle part of flexor margin, distalmost tooth largest, corpus without setae. Propodus about five times longer than wide, about 6.5 times length of dactylus, with row of 6–9 ventral spines including distoventral one. Carpus, merus and ischium 0.87, 1.4 and 0.82 of propodus length, unarmed. Fourth pereiopod (fig. 6 D) propodus with 7–9 spines along ventral margin, dactylus (fig. 6 E, F) with row of 4–6 obtuse teeth in middle of flexor margin of corpus. Fifth pereiopod (fig. 6 G) propodus with two distoventral spines, distoventral cleaning brush not developed, dactylus (fig. 6 H, I) with one obtuse tooth in middle of flexor margin of corpus. Female first pleopod (fig. 7 A) with exopod more than twice length of endopod. Second pleopod with normal appendix interna. Male first pleopod (fig. 7 B) with exopod twice as long as endopod. Endopod (fig. 7 C) with row of simple short setae medially, long serrulate setae distally and plumose setae laterally. Male second pleopod (fig. 7 D) with corpus of appendix masculina (fig. 7 E) not developed, with two long strongly serrulate setae extending beyond normal appendix interna. Uropods (fig. 2 C) extending beyond tip of telson. Protopodite unarmed laterally. Exopod with lateral border with 9–11 strong teeth, increasing in size distally. Very large mobile spine present distolaterally, about 1.8 times longer than dorsal telson spines. Eggs 0.6 mm in diameter. Colouration. Generally dotted with brown to dark green chromatophores over body and appendages (figs. 8, 9). Eyestalks with longitudinal broad brown-green bands. Chelipeds with brown-green reticulate pattern or large dots. Hepatopancreas dark brown to black. Eggs without pigment spot bright orange (fig. 9 upper), with spots brownish (fig. 9 lower). Etymology. The species is named after the author’s sister-in-law, Christel van Eijnatten, with respect and admiration for her inspiring perseverance and positivism in conquering life again after it was almost taken from her. The specific name christellae is a noun in the genitive singular. Host. Acanthostrongylophora ingens (Thiele, 1899) (Porifera: Petrosiidae: Haplosclerida) (fig. 8), identified by N.J. de Voogd. The type species of the genus has been collected from an unidentified black sponge. Distribution. The species is only know from its type locality: Ligitan Island, Semporna area, Sabah, Malaysia. Systematic position Morphological data. The present three specimens differ from N. minirostris in the following characters: 1) the specimens are about twice as large in size as the 9 specimens from the type series as well as the material from Taiwan of N. minirostris; 2) the scaphocerite is almost three times as long as the eye while slightly longer than the eye in N. minirostris; 3) the proximal segment of the antennular peduncle is distinctly longer than the eye while falling short of the eye in the types of N. minirostris and falling short to slightly overreching the eye in the Taiwan material; 4) the outer flagellum of the antennula has the proximal 7–8 segments fused while the proximal 3–6 in N. minirostris; 5) the reduced molar process of the mandible is 2 -segmented while unsegmented in the N. minirostris holotype; 6) the flexor margin of the dactylus of the third pereiopod has a row of 4–6 obtuse teeth in the middle part of the flexor margin of which the distalmost tooth is the largest while one or two acute teeth are present in N. minirostris. The rostrum of the holotype of N. minirostris was depicted in being short and obtuse but without an indication of a subdistal tooth. A subdistal obtuse tooth like in the new species has been observed though in the paratype (RMNH.CRUS.D. 47746) as well as in most specimens of the Taiwan N. minirostris material. In the largest male from Taiwan the subdistal tooth is distinct and acute. The anterolateral margin of the carapace in the holotype of N. minirostris is described as being ‘bluntly rounded’ and figured as such (Bruce & Bauer, 1997: fig. 2 A). In the paratype specimen of N. minirostris (RMNH.CRUS.D. 47746) as well as the Taiwan material however, a strongly produced anterolateral angle is present. Molecular data. The COI barcoding gene was compared with a subset of other sponge-associated pontoniine genera (Table I). No close similarity was found as genetic distances were more than 18 %. The morphologically very disimilar Thaumastocaris came up as the sister taxon in a phylogenetic analysis (fig. 10) indicating the distant relation to the other sponge-associated genera included in this analysis. Support for the branches within the ingroup is low.Published as part of Fransen, Charles H. J. M., 2013, A new species of the sponge-associated pontoniine shrimp genus Nippontonia Bruce & Bauer, 1997 (Decapoda, Caridea, Palaemonidae) from Sabah, Malaysia, pp. 343-357 in Zootaxa 3694 (4) on pages 345-355, DOI: 10.11646/zootaxa.3694.4.3, http://zenodo.org/record/21877
Halocaridinides socotraensis Fransen & Damme 2018, sp. nov.
No full textHalocaridinides socotraensis sp. nov. (Figs. 2–10) Material examined. Socotra Island (Yemen): Ehrer Cave, 12°33’0.54”N 54°27’35.64”E, alt. 250m (type locality). 1 holotype male pocl. 3.9mm (nr. E19) RMNH.CRUS.D.57274; 18 paratypes (2 males, 16 non-ovigerous females, nrs E1–18), RMNH.CRUS.D.57275: from stagnant pools near entrance, 27–28.xii.2003; collected by Kay Van Damme. 34 paratypes (24 males, 10 females of which one ovigerous, nrs E21–55), RMNH.CRUS.D.57276: pop. 1 (waterfall); 27–28.xii.2003; collected by Kay Van Damme. 6 paratypes (1 male, 5 females of which 4 ovigerous, nrs E56–59, E70, 71), RMNH.CRUS.D.57277: pop. 2 (waterfall); 27–28.xii.2003; collected by Kay Van Damme. 6 paratypes (2 males, 4 non-ovigerous females, nrs E60–65), RMNH.CRUS.D.57278: pop. 3 (endbasin); 27–28.xii.2003; collected by Kay Van Damme. Fosreher Cave, 12°37’41.1”N 53°30’42.2”E, alt. 624m, subterranean lake at - 100m. 8 paratypes (1 male, 7 non-ovigerous females, nrs F1–8) RMNH.CRUS.D.57279: - 80 m; 2.i.2004; collected by E. Claes, SKP-team. Ghiniba Cave, 12°26’01.3”N 53°56’11.5”E, alt. 395m, margin of subterranean freshwater river. 1 non-ovigerous female pocl. 3.0mm (third pereiopods missing) (nr. G1), RMNH.CRUS.D.57280: 6.i.2003; collected by Kay Van Damme. 1 non-ovigerous female pocl. 4.38mm (nr. G2), RMNH. CRUS.D.57281: 9.i.2004; collected by L. Vernaeve, SKP-team. Near Qadub, 12°38’14.9”N 53°57’27.2”E, light brackish coastal well. 1 non-ovigerous female mounted in a glass slide, from a zooplankton sample, damaged (not measured), pers. coll. KVD: 12.ii.1999; collected by Kay Van Damme. Comparative material. Halocaridinides trigonophthalma (Fujino & Shokita, 1975) . 3 specimens, RMNH. CRUS.D.40922: Japan, Akasaki-do, long narrow cave with very slowly flowing stream; width 0.6m; 0.25m; bottom clay & stones and hard limestone rock; tourist lights intermittently on in summer; sample mainly from stagnant pool at end of cave second exit; 12x 0.6 m; 23.xii.1986; 19°C, 0 2 saturation 57%, conductivity (EC) 650 S/cm; leg. et don. N.W. Broodbakker. 2 specimens, RMNH. CRUS.D.37247: Japan, Ryukyu Islands, N coast of Yoron Island; 25.xii.1986; leg. et don. N.W. Broodbakker. Many specimens, RMNH. CRUS.D.33133: Palau Distict, Caroline Islands, Angaur Island, Doresi Pond, 6°54’44”N 13°48’20”E; 14.vii.1980; leg. et don. J.A. Maciolek. Halocaridinides fowleri (Gordon, in Gordon & Monod, 1968) . 1 male, ZMA. CRUS.D.103701: Zanzibar, from Pango Managola cave near Makunduchi; 12.ix.1983; leg. et don. P. Beron & V. Beshkov. Description. Body cylindrical. Carapace smooth. Rostrum (Fig. 4A, B) not developed, without teeth, anterior margin of carapace rounded in dorsal view, blunt in lateral view, reaching level of rounded inferior orbital angle. Supraorbital, antennal and pterygostomian spines absent. Anterior lateral margin straight; pterygostomian angle broadly rounded (Fig. 4A, B). Abdomen smooth; pleura of abdominal segments posteroventrally rounded; sixth abdominal segment 1.5 times as long as fifth. Telson (Fig. 4C) as long as sixth abdominal segment, twice as long as its proximal width; lateral margin convex in proximal half, slightly concave in distal half; distal margin 0.6 times as wide as proximal width, straight; two pairs of dorsal spines at about 0.5 and 0.75 of telson length, about 0.0 8 times as long as telson, proximal pair submarginal, distal pair marginal; posterior margin with four pairs of spines, lateral spines as long as dorsal spines, intermediate pair long, three times as long as lateral spines, two pairs of submedian spines about twice as long as lateral spines. Eyes (Fig. 4A, B) triangular, cornea unpigmented, extending to the proximal third of basal segment of antennular peduncle. Antennula (Fig. 4A, B) with peduncle and flagella well-developed. Basal segment without distolateral tooth, distal margin not developed; medioventral tooth absent; stylocerite half as long as basal segment, with distal acute tip, slightly convex lateral margin with row of small setae, median margin sinuous, proximally rather broad; distodorsal margin of basal segment with row of short setae. Intermediate segment somewhat longer than wide, distodorsal margin with few short setae. Distal segment about as long as wide. Upper flagellum slightly longer than carapace, slender; lower flagellum about 1.3 times longer than upper flagellum, slender. Antenna (Fig. 4A, B). Basicerite of antennal peduncle with small distolateral blunt tooth; ischiocerite and mericerite normal; carpocerite cylindrical, reaching distal margin of intermediate segment of antennular peduncle; anterior lamina of scaphocerite almost reaching distal margin of distal segment of antennular peduncle, distolateral tooth distinct but small, not reaching anterior margin of lamina, lateral margin straight, anterior margin rounded. Epistome with blunt anterior median carina; labrum oval. Paragnath small, alae small, oblique rectangular, distomedial margin with row of setae; corpus small, unarmed. Second to eighth thoracic sternites narrow, without special features. Mandible (Fig. 4D) with short incisor process terminating in some irregular teeth and small denticles; bristle of long setae between incisor and molar process; molar process distally concave with rows of very short setae forming transverse ridges. Maxillula (Fig. 4E) upper lacinia broad rectangular with two rows of strong spines medially; lower lacinia broadly rounded medially, with anterior lobe, with one submarginal row of short setae and several marginal rows of simple and serrate setae; palp distally bilobed with two subdistal simple setae. Maxilla (Fig. 5A) with basal endite well developed, bilobed; distal lobe much smaller than broad proximal lobe, with row of slender setae and few spines along distal and medial margin, distalmost spines distally serrate; median margin of proximal lobe with many rows of moderately long, slender, distally serrate setae; coxal endite semicircular, fringed by row of many slender setae along median margin; scaphognathite with narrow posterior lobe and broad anterior lobe, anterior lobe as long as distal lobe of basal endite; palp simple, short, subdistal seta not observed. First maxilliped (Fig. 5B) with small coxal endite with few plumose setae; basal endite long with rows of simple setae on ventral surface and row of long grid setae along median margin; caridean lobe well developed, flagellum short, both fringed by plumose setae; palp as long as caridean lobe, broad, fringed by simple setae distally. Second maxilliped (Fig. 5C) with dactylar segment triangular, median margin with long serrate setae; anterior medial margin of propodal segment produced with row of long robust setae medially; carpal segment short; meral segment with few long plumose setae medially; ischial and basal segments fused, with long plumose setae along medial margin, long plumose seta proximolaterally; exopod well developed, slightly broadened proximally with few long simple setae proximally and long plumose setae distally; coxal segment with oblong triangular epipod, without podobranch. Third maxilliped (Fig. 5D) pediform, exceeding antennal scale by ultimate segment; ultimate segment slightly shorter than penultimate segment, with one stout terminal spine and about 5 subterminal spines, with several transverse rows of strong serrate setae on medial surface; penultimate segment as long as ischiomeral segment, with row of 7 strong spines along medial margin; ischiomeral segment fused to basal segment, slightly broader than penultimate segment, with pair of long plumose setae proximomedially; exopod well developed, reaching exceeding distal margin of ischiomeral segment, with simple and plumose setae distally; coxal segment with epipod laterally and patch of plumose setae medially. Branchial formula as follows (Table 1): Pereiopods. Epipods on the pereiopods are straplike grabbing the single very long robust setobranch of the next pereiopod. First pereiopod (Fig. 6A) with chelae short and broad, fingers about three times as long as palm; tips of fingers with tufts of long brush-like setae; carpus 1.5 times as long as chela, distally broad, deeply excavate; merus shorter than carpus, unarmed; ischium slightly shorter than merus, unarmed; basis short; coxa with strap-like epipod and long single setobranch. Second pereiopod (Fig. 6B) longer and more slender than first pereiopod; chalae as in first pereiopod; carpus twice as long as chela; merus 0.5 times as long as carpus; ischium slightly shorter than merus; basis and coxa as in first pereiopod. Third pereiopod (Fig. 6C, D) with dactylus with 6 spinules along flexor margin of corpus; propodus 3 times as long as dactylus with 11 spinules along ventral margin; carpus 0.85 times propodus length, with 6 spinules along ventral margin; merus straight, slightly longer than propodus, with one spine at proximal fourth of length; ischium slightly less than half length of merus, unarmed; basis and coxa as in first pereiopod. Fourth pereiopod (Fig. 7A, B) with dactylus with 7 spinules along flexor margin of corpus; propodus 3 times as long as dactylus with 11 spinules along ventral margin; carpus 0.75 times propodus length, with 5 spinules along ventral margin; merus straight, as long as propodus, with two spines at half and proximal fourth of length; ischium and basis as in first pereiopod; coxa without epipod, with setobranch. Fifth pereiopod (Fig. 7C, D) with dactylus with row of 16 spinules along flexor margin of corpus; propodus with many small spines along ventral margin; propodus 4 times as long as dactylus with about 14 spinules along ventral margin; carpus 0.6 times propodus length, unarmed; merus straight, 0.8 times propodus length, with one spine in proximal fifth of length; ischium and basis as in first pereiopod; coxa without epipod nor setobranch. First pleopod (Fig. 7E) with endopod as long as exopod, narrowing distally, with blunt tip, without setae. Second pleopod (Fig. 7F) with appendix masculina slightly more than half as long as appendix interna, triangular, with basis broad, tapering distally, with setae along its entire median margin. Uropods (Fig. 4C) with short unarmed protopodite; exopod overreaching telson with half of distal lamina, distolateral tooth strong, only one strong mobile spine at diaeresis; endopod overreaching telson, slightly shorter than exopod. Ovigerous females (specimens nr E21, E56–57, E67) with small clutches of 10– 15 eggs; eggs relatively large, 1.0 x 1.5mm in size. See Figs. 2–3 for general female morphologies. Size. Ehrer Cave: postorbital carapace length (pocl) in largest ovigerous females is 4.06mm; pocl in largest non-ovigerous female is 5.50mm; pocl in largest male is 4.09mm; pocl of smallest male is 2.97mm; most males and females with pocl between 3 and 4 mm. Fosreher Cave: pocl of largest non-ovigerous female is 3.75mm; pocl of only male is 3.69mm. Ghiniba Cave: pocl of two non-ovigerous females respectively 4.38 and 3.06mm. Live colouration. The specimens are a uniform milky white to beige (Fig. 2A). Eggs in females white to yellow. Etymology. The new species was named after Socotra Island (Yemen), its type locality. Morphological variation. The most striking variation (or cryptic speciation?) is found in the robustness of the first and second chelipeds. Differences in both the robustness of the segments of the chelipeds and of the chelae itself are present. The variation covers a relatively continuous range (Fig. 11) and the two extremes at both ends of the morphometric spectrum are: 1) tenuis type (Figs. 2–7), specimens with slender P1 and P2 and with chela with the fingers 2–3 times the palm length, and the chela 3-5 times longer than the palm height, the fingers are usually gaping, the carpus is long and relatively narrow distally. Bristles on the propodus are relatively long, the longest about two thirds of the propodus length (Fig. 6A); 2) robustus type (Figs. 8–10), robust chela with fingers about as long as the palm, and the chela 2.0–2.5 times the palm height, the fingers are not gaping, the carpus is short and relatively broad distally. Bristles on the propodus are relatively shorter, up to half the propodus length (fig. 10A). These two types have no taxonomical value at this point. There has been no correlation found of these types with either sex or size of the specimens (Table 2; Fig. 11). Animals with both slender and robust chelae were found in Erher Cave in stagnant pool and waterfall localities, as well as in Fosreher Cave and Ghiniba Cave. Other features are rather constant. The number of mobile spines at the uropodal diaeresis is always one. The position of the proximal dorsal pair of spines on the telson is between 0.5 and 0.6 of the telson length, that of the distal pair 0.7–0.8. Only few specimens have the distal pair somewhat closer to the distal margin (Fig. 9A). All female specimens have short forward-directing spinules placed on the flexor margins of the dactyli of the third, fourth and fifth pereiopods and short spines on the distoventral margin of the propodi (Fig. 10D, F). Most males (n=20) have long spinules placed perpendicular on the corpus and long spines on the distoventral margin of the propodi as well. The other males (eight) have these features as in the females. These morphological differences in males are not correlated with size nor with location. The length of the appendix masculina ranges from 0.5–1.5 times the length of the appendix interna. There is no relation between the size of the appendix masculina with that of the appendix interna and the variation in dactylar morphology in males. ......continued on the next page Distribution. Socotra Island, Yemen. Found in four localities (Fig. 1): northwest (Fosreher Cave in the Maleh or Ma’alah Plateau), north central (coastal well near Qadub), northeast (Erher Cave in the northern escarpments of the Momi Plateau) and south central (Ghiniba Cave in the Shibehon Plateau). Most common in Erher Cave. Ecology. Subterranean freshwater and slight brackish environments in limestone karst. In Erher, populations were found in three separate locations in the cave: large densities were found in temporary shallow stagnant muddy pools near the entrance that were created by overflow during the rainy seasons. The pools near the entrance receive some light from the cave entrance, yet animals remained unpigmented, and bright white. Lower densities in the clear, shallow permanent water were found in Erher Cave at the base of the large waterfall (creating a strong flow; this can be seen as riverine) and only few animals in the clear, deep permanent lake (end basin) at the end of the cave (no flow). The water in Erher derives from the Momi plateau above and is fresh, measuring between 400–600 µS/cm (256–384 ppm) and a pH of 7.6–7.8. The waterfall is used as local water source (De Geest & Van Damme 2006), transporting water through tubes to the nearby village (Irisseyl). In Ghiniba Cave only two specimens were found in flowing water in the large and clear permanent cave river at the end of the dry section of the cave in the main gallery. The river in Ghiniba contains freshwater of the same low conductivity as in Erher. Conductivity of the water in Fosreher Cave was not measured, yet this can be considered as freshwater as well and was used as drinking water by the team (not brackish in taste). Only few animals (eight) were found in the latter locality, which like Ghiniba functions as a sump for the runoff from the surrounding plateau. Water in the coastal well (near Qadub) was slightly brackish (4200 µS/cm, corresponding to ca. 2690 ppm) and near neutral pH (7.5), here only one specimen was found. Reproductive biology not studied, ovigerous females have large egg clutches containing many white to bright yellow eggs. Remarks. The new species differs from H. trigonophthalma in having only one spine at the uropodal diaeresis in all specimens while H. trigonophthalma has 1–9 spines (Fujino & Shokita 1975, Naruse et al. 2003, present material); in the strongly reduced rostrum, in the absence of pigmentation in the eye; in the appendix masculina becoming not longer than 1.5 times the length of the appendix interna while more than three times as long in H. trigonophthalma (present material). H. socotraensis sp. nov. differs from H. fowleri in having only one spine at the uropodal diaeresis in all specimens while H. fowleri has a variable number of 1–3 spines there. In H. fowleri the rostrum is acute in lateral aspect whereas it is blunt in the new species. The endopod of the first pleopod in males of the new species is almost as long as the exopod and devoid of marginal setae while in the only male specimen known of H. fowleri it is half as long as the exopod bearing several setae along the median margin. The recently described genus from Madagascar, Monsamnis Richard, De Grave & Clark, 2012 also has only a single spine in the uropodal diaeresis (Richard et al. 2012), like the new species of Halocaridinides from Socotra. However, in Monsamnis all pereipods have exopods (fifth reduced), whereas in Halocaridinides exopods are absent on all pereiopods. Monsamnis is known from freshwater mountain streams on Madagascar.Published as part of Fransen, Charles H. J. M. & Damme, K. Van, 2018, A new stygobiont species of Halocaridinides Fujino & Shokita, 1975 (Crustacea, Decapoda, Caridea, Atyidae) from caves on Socotra Island (Yemen), with notes on the genus, pp. 241-261 in Zootaxa 4442 (2) on pages 244-257, DOI: 10.11646/zootaxa.4442.2.3, http://zenodo.org/record/130308
Halocaridinides Fujino & Shokita 1975
No full textGenus Halocaridinides Fujino & Shokita, 1975 Halocaridinides Fujino & Shokita, 1975: 106. Palauatya Hart, 1980: 481. Amended definition. Rostrum unarmed. Supraorbital, antennal and pterygostomian spines absent. Telson broad, with two pairs of dorsal spines. Eyes reduced. Carpus of first two pereiopods excavated anteriorly. Palm of chelipeds distinct. Exopods absent from all pereiopods. Epipods on first three pereiopods. Pleurobranchs on first four pereiopods. First male pleopod without appendix interna. Uropodal exopod ending in a tooth. Diaeresis with one to several movable spines. Type species. Halocaridina (Halocaridinides) trigonophthalma Fujino & Shokita, 1975. Remarks. As there is no concensus on the delimitation of subfamilies within the Atyidae, subdivision has been abandoned (De Grave et al. 2009, De Grave & Fransen 2011). Von Rintelen et al. (2012) did not recover these subfamilies in their phylogeny of atyid genera. Instead of using subfamilies they recognized five groups of genera based on their phylogenetic reconstruction. The anchialine Typhlatya group comprises Halocaridinides, Halocaridina Holthuis, 1963, Typhlatya Creaser, 1936, Stygiocaris Holthuis, 1960 and Antecaridina Edmondson, 1954.Published as part of Fransen, Charles H. J. M. & Damme, K. Van, 2018, A new stygobiont species of Halocaridinides Fujino & Shokita, 1975 (Crustacea, Decapoda, Caridea, Atyidae) from caves on Socotra Island (Yemen), with notes on the genus, pp. 241-261 in Zootaxa 4442 (2) on page 243, DOI: 10.11646/zootaxa.4442.2.3, http://zenodo.org/record/130308
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