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Fig. 10 in Four new genera and five new species of 'Heterocypris' from Western Australia (Crustacea, Ostracoda, Cyprinotinae)
Fig. 10. Bilocypris mandoraensis gen. et sp. nov., Mandora Marsh (type locality). A. LVi, ♂, holotype (WAM 67226). B. RVi, ♂, holotype (WAM 67226). C. LVi, ♀, allotype (WAM 67601). D. RVi, ♀, allotype (WAM 67601). E. CpD, ♀, paratype (WAM 67604). F. CpV, ♀, paratype (WAM 67603). G. RVi, ♂, anterior detail, holotype (WAM 67226). Scales: A–F = 1 mm; G = 500 µm.Published as part of Halse, Stuart A. & Martens, Koen, 2019, Four new genera and five new species of 'Heterocypris' from Western Australia (Crustacea, Ostracoda, Cyprinotinae), pp. 1-35 in European Journal of Taxonomy 493 on page 23, DOI: 10.5852/ejt.2019.493, http://zenodo.org/record/256229
Fig. 7. A–B in Four new genera and five new species of 'Heterocypris' from Western Australia (Crustacea, Ostracoda, Cyprinotinae)
Fig. 7. A–B. Patcypris outback gen. et sp. nov., Lake Maitland (type locality), ♂, holotype (WAM 67184). A. CR attachment (note: the bifurcation to the right is the basis of the CR itself). B. T2. – C–G. Trilocypris horwitzi gen. et sp. nov., Lake Austin (type locality), ♂, holotype (WAM 67199). C. Hp. D. Rpp. E. Lpp. F. CR attachment. G. CR. Scale = 92 µm.Published as part of Halse, Stuart A. & Martens, Koen, 2019, Four new genera and five new species of 'Heterocypris' from Western Australia (Crustacea, Ostracoda, Cyprinotinae), pp. 1-35 in European Journal of Taxonomy 493 on page 16, DOI: 10.5852/ejt.2019.493, http://zenodo.org/record/256229
Fig. 2 in Four new genera and five new species of 'Heterocypris' from Western Australia (Crustacea, Ostracoda, Cyprinotinae)
Fig. 2. Patcypris outback gen. et sp. nov., Lake Maitland (type locality). A. LVi, ♂, holotype (WAM 67184). B. RVi, ♂, holotype (WAM 67184). C. CpRL, ♂, paratype, small specimen (WAM 67188). D. LVi, ♀, allotype (WAM 67185). E. RVi, ♀, allotype (WAM 67185). F. CpRL, ♂, paratype, detail of external rimmed pore on RV (WAM 67188). G. CpD, ♀, paratype (WAM 67187). H. CpD, ♂, paratype, detail anterior (WAM 67189). I. CpD, ♂, paratype, detail posterior (WAM 67189). J. CpD, ♂, paratype (WAM 67189). Scales: A–E, G, J = 1 mm; F = 10 µm; H–I =200 µm.Published as part of Halse, Stuart A. & Martens, Koen, 2019, Four new genera and five new species of 'Heterocypris' from Western Australia (Crustacea, Ostracoda, Cyprinotinae), pp. 1-35 in European Journal of Taxonomy 493 on page 8, DOI: 10.5852/ejt.2019.493, http://zenodo.org/record/256229
Trilocypris Halse & Martens 2019, gen. nov.
Trilocypris gen. nov. urn:lsid:zoobank.org:act: 80ACCA78-B7FD-4D37-BEA3-252915BAEBAD Type species Trilocypris horwitzi gen. et sp. nov. (here designated). Other species The genus is thus far monospecific. Diagnosis Valves high posteriorly, moderately arched, LV overlapping RV on all sides. RV with marginal tubercles (sometimes strongly developed) at least along anterior and postero-ventral margins and an outer list. RV with anterior sub-marginal selvage only; posteroventral selvage more clearly inwardly displaced. LV anteriorly with a blunt inner list parallel to valve margin. A1 with all segments of normal length / width ratio.Second segment of Mx1 palp slightly longer than basal width, not curved. Second segment on Lpp with long distal flagellum. Seta d2 on T2 absent. CR in both male and female proximally curved, but without swollen proximal side; attachment to CR uniramous, i.e., distally not bifurcated. Hemipenis in males with three distal lobes. Etymology This new genus is named after one of its most defining characters, namely the fact that the hemipenis in the male has three distal lobes (instead of two as in other Cyprinotinae). We shortened ‘trilobe’ to ‘trilo’ for convenience. Differential diagnosis Trilocypris gen. nov. has valve characteristics that are similar to those of Heterocypris, but differs from all other genera in the Cyprinotinae by the structure of the hemipenis, which has three distal lobes. This is unique in the subfamily Cyprinotinae, and possibly even in the entire family Cyprididae. The new genus can furthermore be distinguished from Cyprinotus by the absence of the dorsal hump on the right valve, from Hemicypris and Homocypris by the fact that the LV overlaps the RV (inverse in Hemicypris; valves symmetrical in Homocypris) and from Riocypris by, among other features, the absence of seta d2 on T2. Distribution Currently known from the Yilgarn region of Western Australia only. Remarks As usual in monospecific genera, it is difficult to determine which characters are valid at the generic level and which at the specific level. The diagnosis of this new genus may need to be adapted if more congeneric species are found.Published as part of Halse, Stuart A. & Martens, Koen, 2019, Four new genera and five new species of ' Heterocypris' from Western Australia (Crustacea, Ostracoda, Cyprinotinae), pp. 1-35 in European Journal of Taxonomy 493 on page 13, DOI: 10.5852/ejt.2019.493, http://zenodo.org/record/256229
Billcypris davisae Halse & Martens 2019, gen. et sp. nov.
Billcypris davisae gen. et sp. nov. urn:lsid:zoobank.org:act: 3904885C-9677-4739-8B10-5E83D36EB09A Figs 12–13 Diagnosis Valves highly arched, greatest height situated almost in middle. RV with anterior inwardly displaced selvage, more so in female than in male; marginal tubercles absent or very small and few. LV with postero-dorsal extension in male (less developed in females), also visible in carapace in right-lateral view. Lpp with elongated first and second segment, first segment with parallel margins, second segment without long distal flagellum. Hp with very large, sub-rectangular lobe ls and pointed lobe ms, the latter only ventrally developed. Etymology This species is named in honour of Prof. Jenny Davis (Charles Darwin University, Darwin, Australia), in recognition of her significant contributions to Australian limnology, especially of wetlands. Measurements See Table 1. Type locality Lake Cowan, Western Australia. Approximate coordinates: 31°42′04″ S, 121°54′14″ E. All type specimens collected by Shane Chaplin, 8 Apr. 1999 (voucher sample no. OSTR 064 B). Material examined Holotype AUSTRALIA • ♂; Western Australia, Lake Cowan; 31°42′04″ S, 121°54′14″ E; 8 Apr. 1999; Shane Chaplin leg.; voucher sample no. OSTR064 B; with soft parts dissected in glycerine on a sealed slide and with valves stored dry in a micropalaeontological slide; WAM 67217. Allotype AUSTRALIA • ♀; same collection data as for holotype; dissected and stored as the holotype; WAM 67218. Paratypes AUSTRALIA • 1 ♂; same collection data as for holotype; dissected and stored as the holotype; RBINS INV 138038 • 2 male and 2 female carapaces; same collection data as for holotype; used for SEM and stored dry in a micropalaeontological slide; WAM 67219 to 67222. Other material AUSTRALIA • 1 ♂; Western Australia, Lake Lefroy; 31°17′35″ S, 121°41′16″ E [approximate coordinates]; 8 Apr. 1999; Shane Chaplin leg.; dissected and stored as the holotype; WAM 67224 • 6 ♂♂, 7 ♀♀; same collection data as for preceding; in 70% EtOH as bulk vouchers; WAM 67225. Description Male RV (Fig. 12B–C, K). Arched, greatest height situated in, or just behind, middle; anterior margin rounded, dorsal margin straight and posteriorly sloping over a short distance, posterior margin relatively straight in dorsal section and broadly rounded ventrally, ventral margin almost straight. With both anterior and posteroventral sub-marginal selvage, posterior one somewhat more inwardly displaced than anterior one. Anterior and posteroventral margins with some remnants of marginal tubercles. Calcified inner lamellae narrow, more so posteriorly. LV (Fig. 12A, J). With a shape generally comparable to that of RV, slightly larger; anterior calcified inner lamella slightly wider than in RV, anteriorly with blunt inner list running parallel to most of valve margin, posteriorly without inner list. Posterodorsal extension of valve present. CARAPACE. In dorsal view (Fig. 12F) without anterior rostrum; greatest width situated behind middle. In lateral view (Fig. 12H) with posterodorsal extension on LV prominent. PREHENSILE PALPS ON T1. Asymmetrical, chaetotaxy of endopodite as typical for family (not shown). Rpp (Fig. 13B, F) with second segment relatively narrow and curved, distal margin either straight (Fig. 13B) or rounded (Fig. 13F) (possibly depending on position of limb on slide); first segment broad. Lpp (Fig. 13C, E) with nearly rectangular first segment. Second segment sickle- shaped and narrow, of variable length, and without distal flagellum-like expansion. ZENKER ORGAN. As typical of family, i.e., ca 3–5 × as long as wide and with numerous spinous whorls. HEMIPENIS (Fig. 13A, D). With lobe ls large and sub-rectangular, with blunt point extending ventrally beyond edge of hemipenis; ms triangular and bluntly pointed. CAUDAL RAMUS (Fig. 13H). Curved and proximally hardly expanded; its attachment (Fig. 13G) uniramous (note: the bifurcation on the left side on Fig. 13G is actually the proximal part of the CR itself). Female VALVES (Fig. 12 D–E). With shape similar to those of male, but with posteroventral extension on LV much smaller and even absent in some specimens. Valve margin anatomy as in male, except for selvage in RV being more inwardly displaced, both anteriorly and posteriorly. CARAPACE. In dorsal view (Fig. 12L) as in male. In lateral view (Fig. 12G, I) with posteroventral lump of LV still visible in some specimens, absent in others. A1, Md, Mx1, T2, T3 and attachment of CR as in male. A2 (not shown). With distal chaetotaxy showing sexual dimorphism as typical of family. T1 (not shown). With palp not segmented, relatively short and broad. Differential diagnosis This species can at once be distinguished from all others in the subfamily Cyprinotinae by the very large lobe ls and the pointed lobe ms on the hemipenis. The posterodorsal extension on the LV, especially in the male, is also characteristic, but similar features have been seen in other, as yet undescribed, species of West Australian Heterocypris. Ecology and distribution Billcypris davisae gen. et sp. nov. appears to be a halophilic species, collected from Lakes Cowan and Lefroy near Norseman, Western Australia. Both are shallow hypersaline playas that are typically dry but may be shallowly inundated for a few months in wet years. Both lakes are surrounded by smaller playas that are inundated by smaller rainfall events and tend to be less saline. No information on salinities at the time of collecting is available.Published as part of Halse, Stuart A. & Martens, Koen, 2019, Four new genera and five new species of ' Heterocypris' from Western Australia (Crustacea, Ostracoda, Cyprinotinae), pp. 1-35 in European Journal of Taxonomy 493 on pages 26-30, DOI: 10.5852/ejt.2019.493, http://zenodo.org/record/256229
Bilocypris mandoraensis Halse & Martens 2019, gen. et sp. nov.
Bilocypris mandoraensis gen. et sp. nov. urn:lsid:zoobank.org:act: 41706371-0D1C-486E-91D5-B287EE5C3375 Figs 10–11 Diagnosis Valves arched, greatest height situated in middle. RV with anterior and posterior marginal selvage. Marginal tubercles present along anterior, postero-ventral and posterior margins. Lpp with elongated first and second segment; first segment with sub-parallel margins; second segment sickle-shaped and with flagellum-like distal part. Rpp with second segment elongated and somewhat skewed. Hp with lobe ls boot-shaped, with ventral part sharply pointed, and with distal margin almost straight; lobe ms very broad, more clearly bilobed than in preceding species; distal part of bc short and pointed. Etymology Named after the type locality, Mandora Marsh, Pilbara, Western Australia. Measurements See Table 1. Type locality Fern Spring, Mandora Marsh, Pilbara, West Australia. Coordinates: 19°45′59.9″ S, 121°23′34.1″ E. Collected by Adrian Pinder, 4 Sep. 2015. Nearby Melaleuca Spring (coordinates: 19°46′10.7″ S, 121°23′21.3″ E) also sampled. Material examined Holotype AUSTRALIA • ♂; Western Australia, Pilbara, Mandora Marsh, Fern Spring; 19°45′59.9″ S, 121°23′34.1″ E; 4 Sep. 2015; Adrian Pinder leg.; dissected with soft parts stored in glycerine on a sealed slide and valves stored dry in a micropalaeontological slide; WAM 67226. Allotype AUSTRALIA • ♀; Western Australia, Pilbara, Mandora Marsh, Melaleuca Spring; 19°46′10.7″ S, 121°23′21.3″ E; 4 Sep. 2015; Adrian Pinder leg.; dissected and stored as the holotype; WAM 67601. Paratypes AUSTRALIA • 5 ♀♀; same collection data as for allotype; WAM 67227, 67602 to 67605. Description Male RV (Fig. 10B, G). Arched, greatest height situated in middle; anterior margin and dorsal margin rounded, posterior margin nearly straight, ventral margin weakly sinuous in middle. With both anterior and posteroventral sub-marginal selvages, anterior and posteroventral margins set with medium-sized marginal tubercles. Anterior calcified inner lamella relatively wide, posterior lamella narrow; both lamellae without inners lists. LV (Fig. 10A). Generally with shape as in RV, slightly larger. Anterior calcified inner lamella slightly wider than in RV, blunt anterior inner list running parallel to most of valve margin, posteriorly with blunt inner list in top part of calcified inner lamella. CARAPACE. In dorsal and ventral views with a weak anterior rostrum; greatest width situated in middle. In lateral view with anterior LV/RV overlap most prominent. PREHENSILE PALPS ON T1. Asymmetrical, chaetotaxy of endopodite as typical for family (not shown). Rpp (Fig. 11B) with second segment elongated and somewhat skewed, without additional protrusion on dorso-proximal corner, distal sensory organ medium long. Lpp (Fig. 11C) with nearly rectangular first segment (except for skewed proximal margin) and with ventro-distal expansion broad and rounded; second segment sickle-shaped and narrow, with long and thin distal flagellum-like expansion. ZENKER ORGAN. As typical of family, i.e., ca 3–5 × as long as wide and with numerous spinous whorls. HEMIPENIS (Fig. 11A). With ls shaped as a birdhead, ventrally sharply pointed, with distal margin nearly straight; lobe ms very broad, more clearly bilobed than in preceding species; distal part of bc short and pointed. CAUDAL RAMUS (Fig. 11E). Curved; its attachment (Fig. 11F) uniramous. Female VALVES (Fig. 10C–D). With shape similar to that of male. Valve margin anatomy as in the male, except for more inwardly displaced posteroventral selvage in LV. CARAPACE. In dorsal (Fig. 10E) and ventral (Fig. 10F) views with slight anterior rostrum; greatest width situated in middle. A1, Md, Mx1, T2, T3 and attachment of CR as in male. A2 (not shown). With distal chaetotaxy showing sexual dimorphism as typical of family. T1 (not shown). With palp not segmented, relatively elongated. GENITAL LOBES. Large and with rounded distal and distally sloping lateral margins (Fig. 11G–H). Differential diagnosis Bilocypris mandoraensis gen. et sp. nov. is closely related to B. fortescuensis gen. et sp. nov. (see above), but can be distinguished from it by its larger size, the more pronounced bilobed nature of the hemipenis lobe ms, the shape of lobe ls (spatulate in B. fortescuensis gen. et sp. nov., pointed boot-shaped in B. mandoraensis gen. et sp. nov.) and the more elongated and skewed shape of the second segment of the Rpp (robust subtriangular in B. fortescuensis gen. et sp. nov.). Ecology and distribution Bilocypris mandoraensis gen. et sp. nov. has been collected from two small, hyposaline, mound springs within Mandora Marsh, which is a very large saline area near the coast of the southern Kimberley region, Western Australia. The marsh is mostly dry, although it has both marine and groundwater influences and occasionally experiences widespread flooding after cyclonic rain (Storey et al. 2011). Bilocypris mandoraensis gen. et sp. nov. was collected at salinities of 7400 and 6280 mg L-1 TDS in 2015 (Quinlan et al. 2016). The same sites had salinities of 4700 and 5800 mg L- 1 in 1999.Published as part of Halse, Stuart A. & Martens, Koen, 2019, Four new genera and five new species of ' Heterocypris' from Western Australia (Crustacea, Ostracoda, Cyprinotinae), pp. 1-35 in European Journal of Taxonomy 493 on pages 22-25, DOI: 10.5852/ejt.2019.493, http://zenodo.org/record/256229
Trilocypris horwitzi Halse & Martens 2019, gen. et sp. nov.
Trilocypris horwitzi gen. et sp. nov. urn:lsid:zoobank.org:act: 39E852F5-6D42-4412-8004-F721C6861F87 Figs 6, 7C–G Diagnosis Valves higher in posterior half than anterior, but with greatest height situated in middle. RV with both anterior and posterior sub-marginal selvage, posterior one somewhat more inwardly displaced than anterior one; anterior and posteroventral margins set with medium-sized marginal tubercles. LV anteriorly with blunt inner list (more like an elevation), posteriorly with inner list in top part of calcified inner lamella. Carapace in dorsal view with anterior rostrum. Rpp without additional protrusion on sub-triangular second segment. Lpp with almost rectangular first segment, but with rounded expansion in distal part of ventral margin; second segment sickle-shaped and with long, thin apical flagellum. Hemipenis with lobe ls boot-shaped; lobe ms asymmetrically rounded; additional lobe os rounded and pustulose; distal part of bc small and sickle-shaped, distally pointed. Etymology This new species is named in honour of Prof. Pierre Horwitz (Edith Cowan University, Perth, Australia), who collected the present material, in recognition of his outstanding contributions to Western Australian limnology. Measurements See Table 1. Type locality Lake Austin, Western Australia. Approximate coordinates: 27°37′24″ S, 117°54′48″ E. Collected by P. Horwitz, 1 Oct. 1997. Material examined Holotype AUSTRALIA • ♂; Western Australia, Lake Austin; 27°37′24″ S, 117°54′48″ E [approximate coordinates]; 1 Oct. 1997; P. Horwitz leg.; with soft parts dissected in glycerine on a sealed slide and with valves stored dry in a micropalaeontological slide; WAM 67199. Allotype AUSTRALIA • ♀; same collection data as for holotype; dissected and stored as the holotype; WAM 67200. Paratypes AUSTRALIA • 1 ♂; same collection data as for holotype; dissected and stored as the holotype; WAM 67201 • 3 female carapaces; same collection data as for holotype; used for SEM and stored in a micropalaeontological slide; RBINS INV 138035, WAM 67202 to 67203 • 4 ♀♀, 2 ♂♂; same collection data as for holotype; in EtOH as bulk paratypes; WAM 67204. Other material This species has recently been collected from a small number of salt lakes around the type locality. Some of these specimens show slight morphological differences from the specimens described here. Description Male RV (Fig. 6B). With anterior margin rounded and with greatest height in middle; dorsal margin sloping in anterior third, then straight, posterior margin straight, then broadly rounded in ventral half, ventral margin slightly sinuous in middle. RV with both anterior and posteroventral sub-marginal selvages, posterior one somewhat more inwardly displaced than anterior one, anterior and posteroventral margins set with medium-sized marginal tubercles. Calcified inner lamellae narrow, more so posteriorly; both lamellae without inner lists. LV (Fig. 6A). Generally with shape comparable to that of RV, slightly larger. In inner view, anterior calcified inner lamella slightly wider than in RV. Anteriorly with blunt inner list running parallel to most of valve margin, posteriorly with blunt inner list in top part of calcified inner lamella. CARAPACE. In dorsal and ventral views (Fig. 6 F, H–J) with weak anterior rostrum; greatest width situated in middle. In lateral view with prominent anterior LV/RV overlap (illustrated for females in Fig. 6C). PREHENSILE PALPS ON T1. Asymmetrical, chaetotaxy of endopodite as typical for family (not shown). Rpp (Fig. 7D) with subtriangular second segment as in previous species, but without additional process on dorso-proximal corner. Lpp (Fig. 7E) with nearly rectangular first segment, except for rounded distal part of ventral margin. Second segment sickle-shaped and narrow, with distal flagellum-like expansion. ZENKER ORGAN. As typical of family, i.e., ca 3–5 × as long as wide and with numerous spinous whorls. HEMIPENIS (Fig. 7C). With ls boot-shaped, with ventrally directed blunt point; ms evenly rounded, additional lobe os rounded and pustulose. Extremity of internal bc small and sickle-shaped, with pointed tip. CAUDAL RAMUS (Fig. 7G). Curved and proximally less expanded than in previous species. Its attachment (Fig. 7F) uniramous. Female VALVES (Fig. 6D). With shape similar to that of male. Valve margin anatomy as in male. CARAPACE (Fig. 6G). Somewhat wider than in male, but also with weak rostrum. A1, Md, Mx1, T2, T3 and attachment of CR as in male. A2 (not shown). With distal chaetotaxy showing sexual dimorphism as typical of family. T1 (not shown). With palp not segmented, relatively elongated. Differential diagnosis As for genus. Specifically, the structure and shape of the three hemipenis lobes are, to the best of our knowledge, different from that in all other species in the Cyprinotinae. Ecology and distribution Trilocypris horwitzi gen. et sp. nov. is known only from Lake Austin, a large, shallow saline pan situated more than 400 km from the coast, and some smaller pans in its immediate vicinity. Trilocypris horwitzi gen. et sp. nov. was hatched from Lake Austin sediment and collected from the hatching aquaria at salinities between 0.3 and 67.1 mS cm-1 (approximately 200 and 45 0 0 0 mg L-1). At the time the sediment was collected, Lake Austin was flooded and water salinity in the lake and pans varied from 23 0 0 0 mg L-1 to supersaturation. Some additional individuals of Trilocypris horwitzi gen. et sp. nov. were collected, but the salinity where they occurred was not recorded.Published as part of Halse, Stuart A. & Martens, Koen, 2019, Four new genera and five new species of ' Heterocypris' from Western Australia (Crustacea, Ostracoda, Cyprinotinae), pp. 1-35 in European Journal of Taxonomy 493 on pages 13-17, DOI: 10.5852/ejt.2019.493, http://zenodo.org/record/256229
Going Beyond Counting First Authors in Author Co-citation Analysis
The present study examines one of the fundamental aspects of author co-citation analysis (ACA) - the way co-citation
counts are defined. Co-citation counting provides the data on which all subsequent statistical analyses and mappings
are based, and we compare ACA results based on two different types of co-citation counting - the traditional type that
only counts the first one among a cited work's authors on the one hand and a non-traditional type that takes into
account the first 5 authors of a cited work on the other hand. Results indicate that the picture produced through this non-traditional author co-citation counting contains more coherent author groups and is therefore considerably clearer. However, this picture represents fewer specialties in the research field being studied than that produced through the traditional first-author co-citation counting when the same number of top-ranked authors is selected and analyzed. Reasons for these effects are discussed
Bilocypris fortescuensis Halse & Martens 2019, gen. et sp. nov.
Bilocypris fortescuensis gen. et sp. nov. urn:lsid:zoobank.org:act: 9354F295-A5CF-4DB4-A1B1-9DA925F677E9 Figs 8–9 Diagnosis Valves arched, greatest height situated in middle. RV with anterior and posterior marginal selvage. Large marginal tubercles present along anterior, postero-ventral and posterior margins. Lpp with elongated first and second segment; first segment with sub-parallel margins; second segment sickle-shaped and with flagellum-like distal part. Hemipenis with lobe ls spatulate rather than boot-shaped, and with distal margin almost straight; lobe ms bilobed; distal part of bc long, stout and pointed. Etymology Named after the type locality, Fortescue Marsh, Pilbara, Western Australia. Measurements See Table 1. Type locality Fortescue Marsh, Pilbara, Western Australia. Approximate coordinates: 22°30′48″ S, 119°46′41″ E. Collected by Jane McRae and Adrian Pinder, 16 Aug. 2003 (sample PSW002). Subsequent collections made by S. Halse, I. Schön and K. Martens, 24 Apr. 2006 (sample KIES 11). Material examined Holotype AUSTRALIA • ♂; Western Australia, Pilbara, Fortescue Marsh; 22°30′48″ S, 119°46′41″ E [approximate coordinates]; 16 Aug. 2003; Jane McRae and Adrian Pinder leg.; sample PSW002; dissected, with soft parts stored in glycerine on a sealed slide and valves stored dry in a micropalaeontological slide; WAM 67205. Allotype AUSTRALIA • ♀; same collection data as for holotype; dissected and stored as the holotype; WAM 67206. Paratypes AUSTRALIA • 1 ♂; same collection data as for holotype; dissected and stored as the holotype; WAM 67207 • 2 female carapaces; same collection data as for holotype; used for SEM and stored in a micropalaeontological slide; WAM 67208 to 67209 • 1 ♂, 3 ♀♀; same collection data as for holotype; in EtOH as bulk paratypes; WAM 67211 • 1 ♂; same locality as for holotype; 24 Apr. 2006; S. Halse, I. Schön and K. Martens leg.; sample KIES11; with valves stored dry in micropalaeontological slide and with soft body successfully used for DNA extraction; RBINS INV 138036 • 1 ♀; same collection data as for preceding; used and stored as preceding; WAM 67214 • 2 ♂♂; same collection data as for preceding; carapaces used for SEM and stored dry in micropalaeontological slides; WAM 67212 to 67213 • 3 ♀♀; same collection data as for preceding; used and stored as preceding; WAM 67215 to 67216, RBINS INV 138037. The bulk sample of sample KIES11 was lost. Description Male RV (Fig. 8B). Arched, greatest height situated in middle; anterior margin rounded, posterior margin nearly straight, ventral margin weakly sinuous in middle. With both anterior and posteroventral submarginal selvages, anterior and posteroventral margins set with medium-sized marginal tubercles. Anterior calcified inner lamella relatively wide, posterior lamella narrow; both lamellae without inner lists. LV (Fig. 8A). Generally with shape as in RV, slightly larger. Anterior calcified inner lamella slightly wider than in RV, blunt anterior inner list running parallel to most of valve margin, posteriorly with blunt inner list in top part of calcified inner lamella. CARAPACE. In dorsal (Fig. 8J, M) and ventral views (see Fig. 8C, E, of female carapace) with weak anterior rostrum; greatest width situated in middle. In lateral view (Fig. 8H) with anterior LV/RV overlap prominent (illustrated for females in Fig. 8G, I). PREHENSILE PALPS ON T1. Asymmetrical, chaetotaxy of endopodite as typical for family (not shown). Rpp (Fig. 9C) with second segment as in previous species, also without the additional protrusion on dorso-proximal corner. First segment expanding distally, with two ventro-distal lobes, each supporting a sensory organ. Lpp (Fig. 9B) with nearly rectangular first segment, ventro-distal lobes narrow and bluntly pointed; second segment sickle-shape and narrow, with long distal, flagellum-like expansion. ZENKER ORGAN. As typical of family, i.e., ca 3–5 × as long as wide and with numerous spinous whorls. HEMIPENIS (Fig. 9A). With ls spatulate, with distal margin nearly straight; ms bilobed as a result of distal indentation, a small additional rounded ‘lobe’ present on ventral side of ms; distal part of internal bc stout and elongated, with pointed tip. CAUDAL RAMUS (Fig. 9E). Curved; its attachment (Fig. 9D) uniramous. Female VALVES (Fig. 8D–E). With shape similar to that of male. Valve margin anatomy as in male, except for more inwardly displaced posteroventral selvage in LV. CARAPACE. In ventral (Fig. 8K) and dorsal (Fig. 8L) views as in male. A1, Md, Mx1, T2, T3 and attachment of CR as in male. A2 (not shown). With distal chaetotaxy showing sexual dimorphism as typical of family. T1 (not shown). With palp not segmented, relatively elongated. GENITAL LOBES. Large and with rounded distal and sub-parallel lateral margin. Differential diagnosis Bilocypris fortescuensis gen. et sp. nov. is closely related to B. mandoraensis gen. et sp. nov. (see below), but can be distinguished from it by its smaller size, less pronounced bilobed nature of the lobe ms of the hemipenis, the shape of lobe ls (spatulate in B. fortescuensis gen. et sp. nov., pointed boot-shaped in B. mandoraensis gen. et sp. nov.) and the shape of the second segment of the Rpp (robust and subtriangular in B. fortescuensis gen. et sp. nov., more elongated, with a skewed shape in B. mandoraensis gen. et sp. nov.). Ecology and distribution Bilocypris fortescuensis gen. et sp. nov. has been collected only from Fortescue Marsh, a very large mostly hyposaline playa in the central Pilbara region of Western Australia. The marsh floods every few years and holds water for up to six months. Bilocypris fortescuensis gen. et sp. nov. has been recorded at salinities of 3400 and 12 0 0 0 mg L-1 TDS (Pinder et al. 2010). Remarks No male soft parts of the KIES11 material could be checked. However, the valve morphology is so similar, almost identical, to that of the PSW002 material collected at the same location three years earlier that the conspecificity of the animals in the two samples cannot be doubted. There is a hint of a third lobe occurring ventrally on the ms but, because it is more of a ridge than a lobe we do not interpret it as homologous to full-sized third lobe observed in Trilocypris horwitzi gen. et sp. nov.Published as part of Halse, Stuart A. & Martens, Koen, 2019, Four new genera and five new species of ' Heterocypris' from Western Australia (Crustacea, Ostracoda, Cyprinotinae), pp. 1-35 in European Journal of Taxonomy 493 on pages 18-22, DOI: 10.5852/ejt.2019.493, http://zenodo.org/record/256229
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