1,721,746 research outputs found
Oxynoemacheilus isauricus Yogurtcuoglu, Kaya, Ozulug & Freyhof 2021
No full textOxynoemacheilus isauricus Yoğurtçuoğlu, Kaya, Özuluğ & Freyhof 2021 (Fig. 10) Oxynoemacheilus isauricus Yoğurtçuoğlu, Kaya, Özuluğ & Freyhof 2021:372, Figs. 2–4 (type locality: Lake Beyşehir Basin, stream Çeltek at Çeltek, south of Şarkikaraağac, 38.0124 31.3152) Material examined. IUSHM 2021-1425, holotype, 63 mm SL; IUSHM 2021-1463, paratypes, 48–53 mm SL; Turkey: Isparta prov.: Çeltek stream at Çeltek, south of Şarkikaraağaç, 38.0124 31.3152.— FFR 1534, 9, 46–56 mm SL; Turkey: Konya prov.: Akçay stream at Gökçehüyük, 3 km west of Seydişehir, 37.4396 31.8000. Diagnosis. Oxynoemacheilus isauricus is distinguished from all other species of the O. angorae group by having a very slender caudal peduncle (caudal peduncle depth 2.2–2.6 times in its length vs. 1.3–1.6 in O. anatolicus, 1.5–1.8 in O. angorae and O. germencicus, and 1.5–1.9 in O. eregliensis; see details for other species below). It is further differentiated from O. anatolicus, O. angorae and O. eregliensis by possession of a shorter head (head length 21–24% SL vs. 24–27 in O. anatolicus; 23–26 in O. angorae and 24–26 in O. eregliensis), a moderately deep emarginate caudal fin (vs. slightly emarginate or almost truncate) in which the shortest middle caudal-fin ray is 71–85% of the longest ray of the upper caudal-fin lobe (vs. 84–93 in O. anatolicus; 88–92 in O. angorae and 83–91 in O. eregliensis), and no depigmented stripe along the anterior part of the lateral line (vs. present in O. angorae). Oxynoemacheilus isauricus is further distinguished from O. germencicus by lacking scales on the belly (vs. present). Distribution. Oxynoemacheilus isauricus is known from Lake Beyşehir and Suğla basins in Central Anatolia Remarks. Based on DNA barcoding, O. isauricus is well separated by a minimum K2P sequence divergence of 7.5% and 8.0% in the mtDNA-COI barcode region from O. axylos and O. eregliensis, its closest relatives. It is also supported to be a distinct species by the PTP approach, and by the mPTP delimitation.Published as part of Yoğurtçuoğlu, Baran, Kaya, Cüneyt & Freyhof, Jörg, 2022, Revision of the Oxynoemacheilus angorae group with the description of two new species (Teleostei: Nemacheilidae), pp. 451-485 in Zootaxa 5133 (4) on pages 467-468, DOI: 10.11646/zootaxa.5133.4.1, http://zenodo.org/record/653088
Comments on the holotype of Alburnus kurui Mangit & Yerli 2018 and re-description of A. carianorum Freyhof, Kaya, Bayçelebi, Geiger & Turan, 2019 (Teleostei: Leuciscidae)
No full textFreyhof, Jörg, Kaya, Cüneyt, Bayçelebi, Esra, Geiger, Matthias, Turan, Davut (2019): Comments on the holotype of Alburnus kurui Mangit & Yerli 2018 and re-description of A. carianorum Freyhof, Kaya, Bayçelebi, Geiger & Turan, 2019 (Teleostei: Leuciscidae). Zootaxa 4550 (4): 594-596, DOI: 10.11646/zootaxa.4550.4.1
Romanogobio parvus Naseka & Freyhof 2004
No full textRomanogobio parvus : Russia: FSJF DNA-32, 2; Kuban at Ubezhenskaya, 44.914°N 41.278°E (GenBank accession numbers: MF960792, MF960793).Published as part of Friedrich, Thomas, Wiesner, Christian, Zangl, Lukas, Daill, Daniel, Freyhof, Jörg & Koblmüller, Stephan, 2018, Romanogobio skywalkeri, a new gudgeon (Teleostei: Gobionidae) from the upper Mur River, Austria, pp. 336-350 in Zootaxa 4403 (2) on page 348, DOI: 10.11646/zootaxa.4403.2.6, http://zenodo.org/record/121228
Oxynoemacheilus zarzianus Freyhof & Geiger, 2017, new species
No full textOxynoemacheilus zarzianus, new species (Figs. 3–6) Holotype. ZFMK Ich-103667, 67 mm SL; Iraq: stream Kunamasi in Sevanja, 35°47.35'N 45°24.18'E; J. Freyhof, 6 June 2012. Paratypes. FSJF 3352, 28, 39–69 mm SL; same data as holotype. — FSJF 3348, 16, 46–68 mm SL; Iraq: stream in Merga village, 36°03.09'N 45°05.67'E. — FSJF 3651, 18, 54–75 mm SL: Iraq: stream Kunamasi in Kunamasi, 35°47'48"N 45°24'49"E. Additional material (non types). FSJF 3372, 30, 43–71 mm SL; Iraq: stream Suraw near Suraw village, 35°45.76'N 45°59.09'E. Material for molecular genetic analysis: FSJF DNA- 2236; Iraq: stream Kunamasi in Sevanja, 35°47.35'N 45°24.18'E; GenBank accession KY849797.— FSJF DNA- 2203; Iraq: stream in Merga village, 36°03.09'N 45°05.67'E; GenBank accessions KY849792, KY849793, KY849794.— FSJF DNA- 2225; Iraq: stream Suraw near Suraw village, 35°45.76'N 45°59.09'E; GenBank accessions KY849795, KY849796, KY884997. Diagnosis. Oxynoemacheilus zarzianus is distinguished from the other species of Oxynoemacheilus in the Tigris drainage by a combination of characters, none of them unique. Oxynoemacheilus zarzianus belongs to a group of species (O. chomanicus, O. frenatus, O. gyndes, O. hazarensis, O. kentritesensis, O. kiabii, O. zagrosensis), which lack a suborbital groove in males (vs. present in O. bergianus, O. euphraticus, O. hanae, O. longipinnis, O. karunensis, O. kurdistanicus and O. parvinae) and have a slightly emarginate or truncate caudal fin (vs. deeply emarginate or forked in O. bergianus, O. euphraticus, O. hanae, O. longipinnis, O. karunensis, O. kurdistanicus and O. parvinae). Oxynoemacheilus zarzianus is most similar to O. chomanicus (Fig. 2), which also occurs in the Lesser Zab drainage. It is distinguished from that species by having a convex, very rarely straight dorsal profile (vs. straight), a wider interorbital distance (1.1–1.4 times in snout length vs. 1.4–1.5) and a deeper caudal peduncle (caudalpeduncle depth 1.0–1.2 times in caudal-peduncle length vs. 1.3–1.4). It is distinguished from O. zagrosensis, an additional species from the Lesser Zab drainage, by having th e posterior process of the bony air-bladder capsule directed posteriorly (vs. directed laterally). The new species is distinguished from O. kentritesensis, from the Botan River in the Turkish Tigris drainage, by having a short maxillary barbel, which is often rudimentary, reaching the vertical of the anterior eye margin or the vertical of the middle of the eye in some individuals (vs. reaching beyond middle of the eye; maxillary barbel 12–26% HL vs. 27–34), a wider interorbital distance (31–40% HL vs. 27–32; 1.1–1.4 times in snout length vs. 1.5–1.8) and always 8+8 branched caudal-fin rays (vs. 10+9–9+8). Furthermore, the colour pattern on the flank behind the dorsal-fin base is usually mottled or marmorate in O. zarzianus. The marmorate pattern is usually organised in vertically elongated blotches. In O. kentritesensis, there are distinct, but irregularly shaped and set bars on the flank behind the dorsal-fin base. Oxynoemacheilus zarzianus is superficially similar to O. frenatus from the upper Tigris drainage by having the colour pattern on the flank often interrupted by an unpigmented zone along the anterior part of the lateral line (vs. not interrupted on anterior part of flank in other similar species). It is distinguished from O. frenatus by having a complete lateral line (vs. incomplete) and a median incision in the upper lip (vs. absent or with shallow groove in the middle of the upper lip). It is distinguished from O. hazarensis, O. gyndes and O. kiabii by having scales on the back and flank in front of the anus (vs. absent) and a complete lateral line (vs. incomplete). The new species is further distinguished from O. kiabii by the presence of a central pore in the supratemporal canal (vs. absence), three pores in the supratemporal canal (vs. 4–6 pores) and a shorter head (length 22–25% SL vs. 24–30). Oxynoemacheilus zarzianus is further distinguished from O. hazarensis by having a deeper caudal peduncle (caudal peduncle 1.0–1.2 times longer than deep vs. 1.6–1.9). Description. For general appearance see Figs. 3–6; morphometric data are provided in Table 2. Medium sized and stout species with a blunt head. Dorsal profile convex, rarely straight between head and dorsal-fin origin. Body deepest at dorsal-fin origin or at about midline between nape and dorsal-fin origin, depth decreasing at dorsal-fin base and then remains almost equal towards caudal-fin base. A prominent hump before dorsal-fin origin in some individuals. Greatest body width at pectoral-fin base. Section of head roundish, flattened on ventral surface. Interorbital distance wide, 1.1–1.4 times in snout length. Caudal peduncle compressed laterally, 1.0–1.2 (mean 1.1) times longer than deep. Axillary lobe at base of pelvic fin absent or a very shallow pad. Pelvic-fin origin below second or third branched dorsal-fin ray. Anal-fin origin behind middle between base of last dorsal-fin ray and caudal-fin origins. Pectoral fin reaching approximately 50–60% of distance from pectoral-fin origin to pelvic-fin origin. Pelvic fin not reaching vertical of dorsal-fin tip, not reaching to anus. Anus about 0.9–1.3 eye diameter in front of anal-fin origin. Anal fin not reaching caudal-fin base. High to shallow dorsal and shallow ventral adipose crest on caudal peduncle. Dorsal crest reaching to vertical of anal-fin base. Margin of dorsal fin convex. Caudal fin slightly emarginate. Largest known specimen 75 mm SL. Dorsal fin with 8½ (n=15) branched rays. Anal fin with 5½ branched rays. Caudal fin with 8+8 (n=15) branched rays. Pectoral fin with 10–12 and pelvic fin with 6–8 rays. Body covered by scales except anterior part of back, scales isolated and embedded in skin on flank before dorsal-fin origin. Lateral line complete. One lateral pore and 1 central pore in supratemporal canal. In one out of 20 individuals examined for this character, there are two central pores in supratemporal canal. Anterior nostril opening at end of a low, pointed and flap-like tube. Posterior tip of anterior nostril overlapping posterior nostril when folded backwards. No suborbital groove in males. Mouth small, arched (Fig. 7). Lips thick, with marked furrows. A deep median interruption in lower lip. A median incision in upper lip. Processus dentiformis narrow and pointed. No median notch in lower jaw. Barbels short, inner rostral barbel reaching to base of maxillary barbel or to a point between base of outer rostral and maxillary barbel; outer one reaching to base of maxillary barbel. Maxillary barbel very short, rudimentary in some individuals, reaching vertical of anterior margin or middle of eye in others. No suborbital groove or flap in males. Coloration. Body yellowish in life and preserved individuals. Head with very fine and dense brown mottling on top, on cheeks and lateral side of head, without colour pattern ventrally. No pigmentation below a line from pectoral-fin base to anus. Back brown or pale-brown with 3–4 fuzzy, irregularly shaped and set dark-brown blotches, marmorate of mottles in some individuals. Colour pattern on flank not interrupted or with a narrow interruption along anterior part of lateral line. A large, irregularly shaped, dark-brown blotch at dorsal-fin-origin and below posterior half or dorsal-fin base. Flank mottled, behind dorsal-fin origin marmorate or with 3–10 darkbrown, irregularly shaped and set, horizontally elongated blotches, about as wide as or narrower than interspaces in many individuals. Upper part of caudal peduncle often with 3–5 irregularly shaped blotches, some blotches fused with midlateral blotches in some individuals. A wide, dark-brown, irregularly shaped bar at caudal-fin base or two large dark-brown blotches at upper and lowermost caudal-fin base. Last unbranched dorsal-fin ray with a black spot at base of ray. Dorsal-fin with many elongated, dark-brown blotches on rays, forming two bands in some individuals, fuzzy in some others. Caudal-fins with many elongated, dark-brown blotches on rays, forming 2–3 wide, irregularly shaped, dark-brown bands in most individuals, fuzzy in some others. Anal-, pelvic- and pectoral fins with many elongated blotches on rays. Distribution. Oxynoemacheilus zarzianus was found at three places in the Lesser Zab drainage. One spring fed stream (Fig. 8) in the Qal-ah Chulan River drainage, one headwater stream in the Qal-ah Chulan River drainage and in one spring east of the Dukan reservoir. The Qal-ah Chulan is a tributary of the Lesser Zab River, entering the Lesser Zab from the south at about Hawat in Iraq and the Dukan reservoir is a dam lake of the Lesser Zab River. Etymology. Oxynoemacheilus zarzianus is named for the Zarzian culture, which is an archaeological culture of late Paleolithic and Mesolithic in Kurdistan. An adjective. Remarks. The fishes from the stream Suraw (FSJF 3372) a headwater stream in the Qal-ah Chulan River drainage are distinguished from the two other populations by at least 2.6% K2P distance based on the cytochrome b sequences. However, these fishes fit perfectly the diagnosis of O. zarzianus and no morphological difference was found. Therefore, they are identified as O. zarzianus. Four species of Oxynoemacheilus are actually known from the Lesser Zab River drainage. Oxynoemacheilus kurdistanicus is widespread. This is a slender species with a deeply emarginate caudal fin and very narrow caudal peduncle. The occurrence of three (O. chomanicus, O. zarzianus, O. zagrosensis) superficially similar, deep bodied species with a slightly emarginate caudal fin and a very similar colour pattern in the Lesser Zab drainage is the surprising result of this study. The fishes from the Suraw represent even an additional, remarkably isolated population of O. zarzianus. There is indication, that these deep bodied loaches are specialists to headwater streams. In the upper Choman River at the Iraq-Iran border area, JF was not able to find O. chomanicus in the Choman River itself, but only in a very small tributary stream. All over the Lesser Zab drainage in Iraq, usually only O. kurdistanicus is present, but at most sites, no Oxynoemacheilus could be found at all. We suspect that the deep bodied Oxynoemacheilus in the Lesser Zab drainage all have very limited distribution areas, being restricted to springs, cold, spring fed streams or high altitude streams. This very specific habitat choice seems to lead to highly isolated populations and our molecular data support the view that even within one tributary of the Lesser Zab (Qalah Chulan River), isolated populations occur (Suraw & Kunamasi). Our molecular data also support the view that this isolation is not of recent origin but must have lasted for substantial period of time. While deep bodied Oxynoemacheilus species are well known for local endemics, the diversity in the Lesser Zab drainage is an exceptional case. The occurrence of 15 different species of Oxynoemacheilus within the Tigris drainage is a surprising situation. If we consider that seven additional species occur in the Euphrates drainage, including Lake Van basin (O. araxensis, O. argyrogramma, O. ercisianus, O. erdali, O. kaynaki, O. paucilepis, O. samanticus) this raises the number of Oxynoemacheilus species within this combined drainage system to twenty-two. We are not aware of any other genus of fishes with so many species within one drainage system in the Western Palaearctic and we are not aware of any other river being inhabited by so many different Oxynoemacheilus species. In the old world, similar or even much higher species numbers of one genus within one drainage system are found in the Indus, Ganges, Mekong and some other major rivers in South and South-East Asia (Freyhof & Serov 2001, Kottelat 1990, Kottelat 2000, Kottelat 2012). Here also, Nemacheilid loaches are known to be very diverse and restricted to one or few headwater streams within these large drainage systems. Like no other group of freshwater fishes, Nemacheilid loaches seem to become easily isolated in headwaters of rivers. Comparative material. Listed by Freyhof et al. (2012), Freyhof (2016), Freyhof & Abdullah (2017), Freyhof & Özuluğ (2017) and Freyhof et al. (2017).Published as part of Freyhof, Jörg & Geiger, Matthias, 2017, Oxynoemacheilus zarzianus, a new loach from the Lesser Zab River drainage in Iraqi Kurdistan (Teleostei: Nemacheilidae), pp. 258-270 in Zootaxa 4273 (2) on pages 261-268, DOI: 10.11646/zootaxa.4273.2.6, http://zenodo.org/record/80200
Cobitis avicennae Mousavi-Sabet, Vatandoust, Esmaeili, Geiger & Freyhof 2015
No full textCobitis avicennae Mousavi-Sabet, Vatandoust, Esmaeili, Geiger & Freyhof, 2015 (Fig. 4) Cobitis avicennae Mousavi-Sabet, Vatandoust, Esmaeili, Geiger & Freyhof, 2015: 562, figs. 3–4 (type locality: Iran: Hamedan prov.: Gamasiab River at Dehno, a tributary to Karkhek, 34.170 48.355). Material examined. FSJF 3225, 18, 26–37 mm SL; Iran: Hamedan prov.: Gamasiab River at Solgi Sharak, a tributary to Karkheh, 34.281 48.157.— FSJF 3231, 10, 31–56 mm SL; Iran: Kermanshah prov.: Sepidbarg River about 3 km west of Javanrud, 34.805 46.458. Material used in molecular genetic analysis. FSJF DNA-1994; Iran: Kermanshah prov.: Sepidbarg River about 3 km west of Javanrud, 34.805 46.458. (GenBank accession numbers: KP 050516, KP 050525).— FSJF DNA 2712; Iran: Kermanshah prov.: s tream about 1 km south-east of Dinawar, 34.573 47.456. (BOLD accession number: EUFWF 2713-18). Diagnosis. Cobitis avicennae is distinguished from other Cobitis species in the Persian Gulf basin by having one lamina circularis in the male (vs. two in C. elazigensis and C. linea) and distinct, large, dark-brown, usually horizontally elongated blotches in Z4 along the anterior part of the flank, roundish or vertically elongated on the caudal peduncle (vs. indistinct minute dark-grey roundish dots in Z 4 in C. kellei). Distribution. Cobitis avicennae is found in tributaries of the Karkheh and Karun Rivers, which are lower tributaries of the Tigris in Iran. Cobitis from the Little Zab River in Iraqi Kurdistan (Coad 2010) might also belong to this species. Remarks. Molecular data presented by Perdices et al. (2018) and Mousavi-Sabet et al. (2015) and shown in Figure 1, place C. avicennae close to C. faridpaki and C. saniae from the southern Caspian Sea basin and it might be speculated that its ancestors entered the Persian Gulf basin from the Caspian Sea basin by river captures. See below for details to distinguish C avicennae from C. faridpaki and C. saniae. Our own molecular data as well as Perdices et al. (2018) place C. avicennae in the C. taenia species group (Fig. 1). Based on DNA barcoding C. avicennae is well separated from all other Cobitis included in this study and by a minimum K2P distance of 5.7% to C. faridpaki. It is also supported as an own PTP and mPTP entity.Published as part of Freyhof, Jörg, Bayçelebi, Esra & Geiger, Matthias, 2018, Review of the genus Cobitis in the Middle East, with the description of eight new species (Teleostei: Cobitidae), pp. 1-75 in Zootaxa 4535 (1) on page 10, DOI: 10.11646/zootaxa.4535.1.1, http://zenodo.org/record/261577
Oxynoemacheilus nasreddini Yogurtcuoglu, Kaya & Freyhof 2021
No full textOxynoemacheilus nasreddini Yoğurtçuoğlu, Kaya & Freyhof 2021 (Fig. 12) Oxynoemacheilus nasreddini Yoğurtçuoğlu, Kaya & Freyhof 2021:138, Figs. 2–5 (type locality: Aksu stream at Ayvalı, 6 km north of Sincanlı, 38.8101 30.2560, Turkey) Material examined. FFR 15588, holotype, 54 mm SL; FFR 15589, paratypes, 8, 42–57 mm SL; Turkey: Afyon prov.: Aksu stream at Ayvalı, 6 km north of Sincanlı, 38.8101 30.2560.— FFR 15607, 2, 52–53 mm SL; Turkey: Afyonkarahisar prov.: stream flowing into Seyitler reservoir at İscehisar, 38.8160 30.7870.— FFR 1537, 35, 30–66 mm SL; Turkey: Afyonkarahisar prov.: Kali stream at İnli, 1 km south to Maltepe, 38.6004 30.8934.— FFR 1565, 7, 54–76 mm SL; Turkey: Konya prov.: Deliköyboğazı stream 1 km south to Ilgın 38.2370 31.8834.— FSJF 3209, 8, 58–78 mm SL; Turkey: Konya prov.: stream Ali (Karacaağaç) at Doğanhisar, 38.1356 31.6674.— IUSHM 2021- 1429, 2, 63–67 mm SL;— FSJF 3096, 3, 66–71 mm SL; Turkey: Isparta prov.: Özdere stream at Eğirler, 8 km northeast of Gelendost, 38.1976 31.1074.— FFR 15523, 10, 42–68 mm SL; Turkey: Isparta prov.: Özdere stream at Madenli, 6 km northeast of Gelendost, 38.1813 31.1008.— FSJF 2471, 2, 60–67 mm SL; Turkey: Isparta prov.: lower stream Çayköy at Koysazı bridge, 37.8415 30.8916.— FSJF 2516, 1, 61 mm SL; Turkey: Isparta prov.: middle stream Çayköy above Kemerköprü water regulator, 37.8376 30.9008. Additional distribution records. Yeğen et al. 2007: 38.7894 30.0961, 39.0167 30.4681, 39.0533 30.4839, 39.0919 30.4406; Yoğurtçuoğlu et al. 2021a: 38.4473 31.5175, 38.5420 38.5420, 38.7790 30.7910, 38.7880 30.0290, 38.9600 30.4460, 38.9440 30.4420, 38.8550 30.1630, 30.8080 30.3380. Diagnosis. Oxynoemacheilus nasreddini is distinguished from other species of O. angorae group by a combination of characters, none unique to the species. It is distinguished from O. mediterraneus and O. anatolicus by having a more slender caudal peduncle (caudal-peduncle depth 1.5–2.1 times in length vs. 1.2–1.5 in O. mediterraneus and 1.3–1.6 in O. anatolicus). It is further distinguished from O. mediterraneus by an emarginate caudal fin (middle caudal-fin ray 76–91% of length of longest upper caudal-fin ray vs. deeply emarginate, 65–76), a slowly decreasing body depth between the dorsal- and caudal-fin bases (depth of caudal peduncle 70–86% of body depth at anteriormost dorsal fin base vs. almost uniform, 85–92), and the flank blotches being usually disconnected from the saddles on the back, rarely 1–2 flank blotches connected (vs. usually all connected). Oxynoemacheilus nasreddini is further distinguished from O. anatolicus, O. angorae, O. isauricus and O. eregliensis by having few embedded scales on the belly (vs. belly without scales), and the dorsal part of the head and the upper part of the cheek with a vermiculate or marbled pattern (vs. mottled). It is further distinguished from O. angorae by having irregularly shaped and set, vertically elongated blotches on the flank, rarely a mottled or marbled pattern (vs. a series of dark-brown midlateral blotches usually fused into a wide, irregular shaped midlateral stripe, rarely a mottled pattern), the tip of the pectoral fin usually reaching to or slightly beyond the pelvic-fin origin in the male (vs. not reaching), a longer pre-pelvic length (51–55% SL vs. 48–51), a smaller distance between the pelvic- and anal-fin origins (20–23% SL vs. 23–28), and 8–10 pores in the preoperculo-mandibular canal (vs. 10–13). It is further distinguished from O. isauricus by having a deeper caudal peduncle (caudal-peduncle depth 1.5–2.1 times in length vs. 2.2–2.6), a more posteriorly positioned pelvic fin (pre-pelvic length 51–55% SL vs. 48–51), and usually a series of short bars on caudal peduncle (vs. a series of midlateral blotches on flank). It is further distinguished from O. eregliensis by having a series of vertically elongated blotches along the lateral midline or slightly below, rarely a mottled pattern (vs. irregularly set and shaped, often round blotches, usually forming a marbled pattern), anal and pelvic fins without brown blotches (vs. present in individuals larger than 50 mm SL), anal-fin origin and anal-fin base without pigmentation (vs. usually with a brown blotch at anal-fin origin, often with brown anal-fin base), a smaller distance between the pelvic- and anal-fin origins (20–23% SL vs. 23–26), and a longer anal fin (anal fin length 16–19% SL vs. 14–16). Oxynoemacheilus nasreddini is differentiated from O. germencicus by having the tip of the pelvic fin usually not reaching to the anus (vs. reaching), and a prominent inner axial stripe (vs. absent). Distribution. Oxynoemacheilus nasreddini is found in tributaries of the endorheic Lake Akşehir, Eber, Eğirdir and Ilgın basins in Central Anatolia.Published as part of Yoğurtçuoğlu, Baran, Kaya, Cüneyt & Freyhof, Jörg, 2022, Revision of the Oxynoemacheilus angorae group with the description of two new species (Teleostei: Nemacheilidae), pp. 451-485 in Zootaxa 5133 (4) on page 470, DOI: 10.11646/zootaxa.5133.4.1, http://zenodo.org/record/653088
Egirdira Freyhof 2022, new genus
No full text<i>Egirdira</i>, new genus <p>Fig. 1–4</p> <p> <b>Material examined.</b> FSJF 2337, 25, 23–53 mm SL; FSJF 3638, 36, 25–74 mm SL; Turkey: Isparta prov.: spring Karaot at shore of Lake Eğirdir, about 4 km north of Yenice, 38.1349 30.9074.— FSJF 3097, 2, 56–61 mm SL; Turkey: Isparta prov.: Özdere stream at Eğirler, 8 km northeast of Gelendost, 38.1976 31.1074.— FSJF 2475, 1, 46 mm SL; Turkey: Isparta prov.: lower stream Çayköy at Koysazý bridge, southeast of Eğirdir, 37.8415 30.8916.</p> <p> <b>Diagnosis.</b> <i>Egirdira</i> is distinguished from all genera of Leuciscidae except <i>Delminichthys</i> and <i>Pelasgus</i> by the male having the pectoral-fin rays 2–3 slightly slanted upward, with a conspicuous downward angle at the distal end of the thickened part (vs. pectoral fin of male not modified, identical in shape to that of the female, often longer in the male). It is distinguished from <i>Pachychilon</i> by lacking a median lobe in the lower lip (vs. present) and having the upper lip not covered by the overhanging rostral cap (vs. covered). <i>Egirdira</i> is distinguished from <i>Tropidophoxinellus</i> by lacking a midventral keel in front of the anus (vs. present), having only 0–4 pored lateral-line scales (vs. lateral line complete), and 6½ branched anal-fin rays (vs. 8–11½). <i>Egirdira</i> is distinguished from <i>Delminichthys</i> by having an incomplete lateral line (vs. complete) with the genital papilla of the female not protruding (vs. protruding). <i>Egirdira</i> is superficially similar to <i>Pelasgus</i> and distinguished from all species of this genus by having isolated (or patches of) silvery scales on the flank (vs. all scales equally silvery, brown or yellowish). No other unique character could be found. The following combination of characters allow <i>Egirdira</i> to be distinguished from the various species in <i>Pelasgus</i>: Body considerably compressed (vs. body thick with an ovoid section in all <i>Pelasgus</i> except <i>P. thesproticus</i>), 41–52+2 scales in midlateral row (36–41+ 2 in <i>P. marathonicus</i>, 38–43 + 2–3 in <i>P. thesproticus</i>, 58–73 + 2 in <i>P. prespensis</i>, 53–58 + 2 in P. <i>epiroticus</i>); scales overlapping, not deeply embedded (vs. very small, not overlapping, deeply embedded in skin in <i>P. prespensis</i>), 0–4 pored lateral line scales (vs. 16–23 in <i>P. epiroticus</i>, 7– 14 <i>P.</i> thesproticus), 6½ branched anal-fin rays (vs. 7–7½ in <i>P. minutus</i>, <i>P. marathonicus</i>, <i>P. stymphalicus</i> and <i>P. thesproticus</i>), ½11½ scale rows on caudal peduncle (vs. ½7–8½ in <i>P. marathonicus</i>, ½8–9½ in <i>P. stymphalicus</i> and <i>P. thesproticus</i>), scales pockets not covering parts of exposed scales (vs. covering in <i>P. laconicus</i>), a prominent midlateral stripe (vs. absent or indistinct in <i>P. minutus</i>, <i>P. marathonicus</i>, <i>P. prespensis</i>, <i>P. stymphalicus</i> and <i>P. thesproticus</i>), no isolated or irregularly shaped patches of dark-brown scales along lateral midline (vs. present in <i>P. laconicus</i>).</p> <p> <b>Type species.</b> <i>Pararhodeus niger</i> Kosswig & Geldiay 1952</p> <p> <b>Included species.</b> <i>Egirdira nigra</i> (Kosswig & Geldiay, 1952)</p> <p> <b>Etymology.</b> The name is derived from Lake Eğirdir (female)</p> <p> <b>Remarks.</b> It was the aim of this study to find a unique morphological character state distinguishing <i>Egirdira</i> from all species of <i>Pelasgus</i>. Over a period of more than 10 years the available material was taken several times from the collection and fishes were examined but no unique character shared by all species of <i>Pelasgus</i> and distinguishing them from <i>Egirdira</i> could be detected. The only exception is the isolated (or patches of) silvery scales on the flank in <i>Egirdira</i>, absent in <i>Pelasgus</i>. The inclusion of <i>Egirdira</i> in <i>Pelasgus</i> is not supported by any of the molecular studies published so far. Perea <i>et al</i>. (2010) and Geiger <i>et al</i>. (2014) presented molecular phylogenetic analyses justifying the generic status of <i>Egirdira</i>, despite the absence of a unique diagnostic morphological character state. Indeed, this is not an uncommon situation in Leuciscidae: genera such as <i>Alburnus</i>, <i>Alburnoides</i>, <i>Acanthobrama</i>, <i>Petroleuciscus</i>, <i>Ladigesocypris</i>, <i>Tropidophoxinellus</i>, and perhaps others lack unique morphological characters, or these might be internal characters not studied so far.</p> <p> <b> <i>Phoxinellus egridiri</i>, a junior synonym of <i>E. nigra.</i></b> Kosswig & Geldiay (1952: 12) described <i>Pararhodeus niger</i> in a small booklet published in Turkish language about the fishes of Lake Eğirdir. The description is not very informative but Kosswig & Geldiay (1952) provide an identification key (page 14) and show a photograph of the species at the end of the publication, which clearly represents the fish usually identified as <i>P. egridiri</i>. The description (Kosswig & Geldiay 1952: 12) translates as: “Another small fish species is <i>Pararhodeus niger</i>. The name “ <i>niger</i> ” of this fish, which is nearly 8–10 cm and one of the relatives of large cyprinid species, is originated from the colour particularly exhibited by males in breeding season. Out of the breeding season, the dorsum is dark-grey, and flank is dark-silvery to velvet-black, and partly with silvery blotches. Its feeding habits is similar to Yosun Balığı [<i>Anatolichthys</i>], and the diet is mostly composed of small aquatic animals and mosquito larvae. The mouth is suitable for foraging by being located at the tip of the head.” This information is sufficient for a species description to suit the criteria of availability and there is no doubt, that <i>P. niger</i> is an available species name. <i>Pararhodeus</i> is masculine and the species name “ <i>niger</i> ” is an adjective. When associated with <i>Egridira</i> (feminine), it must be declined to <i>nigra</i>.</p> <p> Karaman (1972) was obviously not aware of this publication when he described <i>Phoxinellus egridiri</i>. This is surprising, as Kosswig went in 1955 from Istanbul to Hamburg, where he lived in retirement since 1969. Küçük <i>et al.</i> (2009:282) too, were aware that Karaman (1972) had described <i>P. niger</i> again: “It was first recorded as <i>Pararhodeus niger</i> by Kosswig and Geldiay (1952). Later, Karaman (1972) defined it as a new species in genus <i>Phoxinellus</i>.” Küçük <i>et al.</i> (2009) lists materials of <i>P. niger</i> in the Inland Water Fish Collection of the Fisheries Faculty Museum of the Ege University (ESFM-PISI) in Izmir (ESFM-PISI/1950-007, 1, 49 mm SL; Turkey: Lake Eğirdir. ESFM-PISI/1951- 003, 28, 20- 44 mm SL; Turkey: Lake Eğirdir). These were collected in 1950 and 1951 when Remzi Geldiay worked at ESFM-PISI. It is likely that these specimens are syntypes of <i>P. niger</i>. It is clear from the text (and the context) of Kosswig & Geldiay (1952) that they described the species based on their field collections. As they do not specify the number of syntypes, all specimens collected and “seen” by them are syntypes. Kosswig and Geldiay collaborated widely in zoological research after the World War II, including in ichthyology, until Kosswig relocated to Germany and settled in Hamburg in 1955. Küçük <i>et al.</i> (2009) list the syntypes under <i>Pseudophoxinus egridiri</i> not mentioning that <i>P. niger</i> must have priority over <i>P. egridiri</i>. Here we treat <i>P. egridiri</i> as a junior synonym of <i>P. niger</i>.</p>Published as part of <i>Freyhof, Jörg, 2022, Egirdira, a new generic name for Pararhodeus niger Kosswig & Geldiay, 1952 (Teleostei: Leuciscidae), pp. 586-592 in Zootaxa 5104 (4)</i> on pages 587-590, DOI: 10.11646/zootaxa.5104.4.8, <a href="http://zenodo.org/record/6332245">http://zenodo.org/record/6332245</a>
Oxynoemacheilus karunensis, a new species from the Persian Gulf basin (Teleostei: Nemacheilidae)
No full textFreyhof, Jörg (2016): Oxynoemacheilus karunensis, a new species from the Persian Gulf basin (Teleostei: Nemacheilidae). Zootaxa 4175 (1): 94-100, DOI: http://doi.org/10.11646/zootaxa.4175.1.
Glyptothorax strabonis Ng & Freyhof, 2008, sp. nov.
No full text<i>Glyptothorax strabonis</i> sp. nov. <p>(Fig. 4)</p> <p> <b>Type material.</b> Holotype: UMMZ 245669, 80.1 mm SL; Vietnam: Quang Binh province, market in Phong Nha; J. Freyhof, F. Herder & D. Serov, 7 April 2000.</p> <p>Paratypes. ZFMK 35285–35286, (2), 84.0, 95.5 mm SL; data as for holotype.</p> <p> <b>Diagnosis.</b> <i>Glyptothorax strabonis</i> can be distinguished from all Indochinese congeners in having a smaller eye (6.2–6.5% SL vs. 8.6–12.0). It is further distinguished from congeners in central Vietnam by its thoracic adhesive apparatus, which consists of striae arranged in a broad oval field (Fig. 2) and by the unique combination of the following characters: length of caudal peduncle 20.5–21.2% SL, depth of caudal peduncle 7.5–7.6% SL, body depth at anus 13.7–20.9% SL, head width 17.9-18.8% SL, presence of 14–16 serrae on posterior margin of pectoral spine, and coloration consisting of a dark-brown body with indistinct pale midlateral and mid-dorsal stripes.</p> <p> <b>Description.</b> Biometric data as in Table 2. Head depressed, body subcylindrical. Dorsal profile rising evenly from tip of snout to origin of dorsal fin, sloping gently ventrally from origin of dorsal fin to end of caudal peduncle. Ventral profile flat to anal-fin base, sloping gently dorsally from anal-fin base to end of caudal peduncle. Anus and urogenital openings located at vertical through middle of adpressed pelvic fin. Skin tuberculate, with small tubercles on sides of body. Lateral line complete, mid-lateral. Vertebrae 17+21=38 (3).</p> <p>Head depressed, broad; triangular in lateral view. Snout prominent. Anterior and posterior nares large, separated only by base of nasal barbel. Gill openings broad, extending from immediately ventral to post-temporal to isthmus. Bony elements of dorsal surface of head covered with thick, tuberculate skin. Eye ovoid, its horizontal axis longest; located entirely in dorsal half of head.</p> <p>Barbels in four pairs. Maxillary barbel long, slender; extending to middle of pectoral-fin base. Nasal barbel slender, extending to four-fifths of distance between its base and anterior orbital margin. Inner mandibular-barbel origin close to midline, extending to midway between its base and that of pectoral spine. Outer mandibular barbel originating posterolateral of inner mandibular barbel, extending to four-fifths of distance between its base and that of pectoral spine.</p> <p>Mouth inferior, premaxillary tooth band partially exposed with mouth closed. Oral teeth small, villiform; in irregular rows on all tooth-bearing surfaces. Premaxillary teeth in a single broad semilunate band. Dentary teeth in two narrow crescentic bands separated at midline.</p> <p>Dorsal fin above anterior third of body, with I,6 (3) rays; fin margin convex; spine short, straight; with 3– 4 very small dentations on posterior margin. Adipose fin with anterior margin straight or very slightly concave, posterior margin angular. Caudal fin strongly forked, with lower lobe very slightly longer than upper lobe and i,7,8,i (3) principal rays. Procurrent rays symmetrical, extending only slightly anterior to fin base. Vertical through anal-fin origin ventral to that through adipose-fin origin. Anal fin with straight anterior margin, straight or slightly concave posterior margin; with iv,10 (3) rays. Pelvic-fin origin at vertical through posterior end of dorsal-fin base. Pelvic fin with slightly convex margin and i,5 (3) rays; tip of adpressed fin not reaching anal-fin origin. Pectoral fin with I,9 (3) rays; posterior fin margin slightly concave; anterior spine margin smooth, posterior margin with 14–16 serrae.</p> <p>Thoracic adhesive apparatus present, consisting of ridges of skin (striae) in elongate oval field extending from isthmus to just posterior to level of last pectoral-fin base. Median depression present on posterior third of adhesive apparatus. Striae orientated anterodistally, radiating from median depression. Striae uninterrupted except for posteriormost lateral ones, which are dissociated into irregular small, unculiferous patches.</p> <p> <b>Coloration.</b> In 70% ethanol: Dorsal and lateral surfaces of head, and body dark brown, fading to beige on ventral surfaces. Nuchal plate elements beige in some individuals. A thin, beige mid-dorsal stripe extending from base of last dorsal-fin ray to origin of adipose fin present in some individuals. Laterosensory pores along lateral line rimmed in beige, imparting appearance of a diffuse beige mid-lateral line in some individuals. Dorsal and pelvic fins with brown fin rays, diffuse melanophores on fin membranes and hyaline distal margin. Pectoral fin with brown on bases of fin rays and brown band along middle third. Anal fin with brown base and brown melanophores on fin rays forming an irregular subdistal band; rest of fin hyaline. Adipose fin brown, with hyaline distal margin. Caudal fin brown, with beige procurrent rays and tips of lobes hyaline. Maxillary and nasal barbels brown dorsally, beige ventrally.</p> <p> <b>Etymology.</b> From the Latin <i>strabonis</i>, meaning one who squints, in reference to the relatively small eye of this species. A noun.</p> <p> <b>Distribution.</b> Known from the Giang River (Song Giang) drainage in central Vietnam (Fig. 3).</p>Published as part of <i>Ng, Heok Hee & Freyhof, Jörg, 2008, Two new species of Glyptothorax (Teleostei: Sisoridae) from central Vietnam, pp. 11-25 in Zootaxa 1873</i> on pages 16-20, DOI: <a href="http://zenodo.org/record/184036">10.5281/zenodo.184036</a>
Oreoglanis infulatus Ng & Freyhof 2001
No full text<p>Oreoglanis infulatus Ng & Freyhof 2001</p> <p>Oreoglanis infulatus Ng & Freyhof 2001: 1165, figs. 1-3. Type locality: Stream at Son Kim, a trib. of Song Lam, 18°24'25"N, 105°11'10"E, Ha Tinh Prov., Viet Nam. Holotype: ZFMK 35719. Paratypes: UMMZ 238025 (6); ZFMK 35720-25 (6).</p> <p>Distribution: Lam River drainage, central Vietnam (Ng & Freyhof, 2001).</p>Published as part of <i>Alfred W. Thomson & Lawrence M. Page, 2006, Genera of the Asian Catfish Families Sisoridae and Erethistidae (Teleostei: Siluriformes)., pp. 1-96 in Zootaxa 1345</i> on page 7
- …
