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Geoplana paranapiacaba Ana Laura Almeida & Fernando P. L. Marques & Fernando Carbayo 2019, SP. NOV.
No full textGEOPLANA PARANAPIACABA ALMEIDA & CARBAYO SP . NOV . Material examined Holotype MZUSP PL 2079 (field number F6660): Cidade Turística de Paranapiacaba, Santo André/ SP, Brazil (23°46′38.9″S, 046°18′0 5.2″W). F. Carbayo et al., coll., 5 June 2015. Cephalic region: horizontal sections on four slides; ovarian region: horizontal sections on nine slides; region behind ovaries: horizontal sections on 14 slides; pre-pharyngeal region: transverse sections on 16 slides; pharynx: sagittal sections on 17 slides; copulatory apparatus: sagittal sections on 20 slides. Paratype MZUSP PL 2087 (field number F6998): Cidade Turística de Paranapiacaba, Santo André/ SP, Brazil. E. M. Froehlich et al., coll., 13 May 1990. Copulatory apparatus: sagittal sections on 39 slides. Distribution Only known from the type locality. Etymology The specific epithet refers to the Tupi-Guarani (indigenous Brazilian tribe) name of the district where the type material was collected. Diagnosis Dorsal colour pattern constituted by a median stripe of orange-ochre, bounded on either side by a black line, externally to which is a broader white line, and in turn this is externally bordered by a thin black line, externally to which is a grey band with dense white mottling, bordered by a thin marginal black line. Diameter of muscular cylinder round ejaculatory duct equivalent to 0.5 times the diameter of this duct. Female-tomale atrial length ratio, 3.5–4. External aspect Preserved animals up to 50 mm long, and 4 mm wide. Body margins nearly parallel; anterior end rounded, posterior end pointed. Dorsum strongly convex, ventral side slightly convex. Dorsal body surface constituted by a median stripe of orange-ochre (onequarter of body width) that passes into the black of cephalic region (one-fifth of body length), bounded on either side by a black line (1/16th) posteriorly merging with each other, externally to which is a broader white line (one-eighth), and this is externally bordered by a black stripe (1/16th), externally to which is a grey stripe (one-eighth) with dense white mottling, then bordered by a marginal thin black line (Fig. 12A). Ventral body surface is whitish with blackish margins. In preserved specimen F6660, dorsal colour slightly faded. In preserved specimen F6998, stripes were varied from faded black grey to pearl orange. Two types of eyes: a conical type contouring only anterior end; and a cup-shaped type, spreading dorsally onto the black stripes. Sensory pits simple invaginations, 40 μm deep, located ventromarginally in a single row from the very anterior tip to at least the end of the anterior fifth of the body. Relative position mouth-to-body length, 61%. Relative position gonopore-to-body length, 74%. Internal morphology Creeping sole 78% of body width. Glandular margin absent. Three typical geoplaninid cutaneous muscle layers present, being as thick as 8% of body height at pre-pharyngeal region. Longitudinal muscle fibres of the innermost layer (35–50 µm thick) arranged into bundles with 17–30 fibres each. Three parenchymal muscle layers present: a dorsal layer of decussate diagonal fibres, a loose supraintestinal layer of transverse and longitudinal fibres, and a loose subintestinal layer with traverse and longitudinal fibres (Fig. 12B, C). Ventral nerve plate present. Mouth situated at a distance from root of the pharynx equivalent to 66% of pharyngeal pocket length. Pharynx cylindrical (Fig. 12D). Oesophagus-topharynx ratio, 26%. Outer pharyngeal musculature consisting of a subepithelial layer (5 µm) of longitudinal muscle, followed by a layer (70 µm) of intermingled circular and longitudinal muscle fibres. Inner pharynx musculature consisting of a thin subepithelial layer of longitudinal muscle, followed by a layer (90 µm) of circular fibres and an innermost layer (15 µm) of longitudinal fibres. Testes dorsally located between supraintestinal parenchymal muscle layer and intestine; anteriormost testes (at a distance from anterior end equal to 22% of body length) behind ovarian region, posteriormost near root of pharynx. Penis bulb elongated, strongly developed, consisting of tightly packed muscle fibres variously orientated in its anterior section, while posteriorly the fibres are orientated diagonally, extending from 1 mm (holotype) anterior to penis papilla to the level of the gonopore, even more posteriorly in the dorsal body region. Diagonal fibres on the right side of the bulb run obliquely downwards to embrace ventrally the anterior portion of the female atrium; further course of these fibres could not be determined owing to tissue damage. Sperm ducts run dorsolaterally to the ovovitelline ducts. Laterally to penis bulb, sperm ducts bend anteriorly and medially, and then ventrolaterally penetrate the bulb. The ducts subsequently recurve and join to form an unpaired, sinuous sperm duct directed posteriorly (Fig. 13A, B). The unpaired portion continues as an ejaculatory duct inside the penis papilla, which traverses centrally to open at its tip. Sperm present in sperm duct, both along its course outside and in part of anteriorly directed portion inside penis bulb. Epithelium of sperm ducts and unpaired sperm duct pierced by glands producing minute weakly cyanophil granules; surrounded by a 7-µm-thick layer of circular muscles. Ejaculatory duct lined by 15 µm, ciliated epithelium pierced by abundant glands producing erythrophil granules; surrounded by a layer of circular muscle fibres constituting a muscular cylinder with a diameter equivalent to 0.5 times the diameter of this duct. Penis papilla long, with dorsal and ventral insertions at the same transverse plane; it is horizontal and cylindrical, with a length (4 mm) equal to ten times its diameter, even occupying more than half of the female atrium. Papilla covered with a cuboidal, non-ciliated epithelium, pierced by gland cells especially distally, producing erythrophil granules (Figs 12E, 13D). Subepithelial penis musculature consisting of a dense 25- to 105-µm-thick layer of circular muscle, followed by a thin layer (12 µm) of longitudinal muscle. Male atrium not folded (Fig. 13A, B), lined with a low, non-ciliated epithelium, pierced by glands producing an amorphous erythrophil substance and underlain by a 15-µm-thick layer of circular muscles, followed by a thin layer of longitudinal fibres. Ovaries oval shaped, 550 µm in maximal anteroposterior diameter and 250 µm in dorsoventral diameter, and situated at a distance from anterior end equivalent to 16% of body length. Ovovitelline ducts emerge from dorsolateral aspect of ovaries. The oviducts run laterally to the female atrium, then curve medially and join to form a common glandular ovovitelline duct, dorsally to this atrium. Common glandular ovovitelline duct as long as half of the female atrium length, communicates with female genital duct; the latter being a 40-µm-long projection of the dorsoposterior region of the female atrium (Fig. 13A, B, D). Female atrium long, without folds, as long as 3.5–4 times the male atrium length (Fig. 13A, B, D) and lined with a tall epithelium (70 µm). Posterior section of female atrium lined with an epithelium with stratified aspect and some internal gaps; surrounded by a thin layer of longitudinal muscle fibres, followed by 70-µm-thick layer of circular fibres, and a third layer (115 µm) of fibres in criss-cross arrangement. No evident distinction between atrial fibres and those of common muscle coat.Published as part of Ana Laura Almeida, Fernando P. L. Marques & Fernando Carbayo, 2019, ' Endless forms most beautiful': taxonomic revision of the planarian Geoplana vaginuloides (Darwin, 1844) and discovery of numerous congeners (Platyhelminthes: Tricladida), pp. 1-65 in Zoological Journal of the Linnean Society 185 on pages 35-37, DOI: 10.1093/zoolinnean/zly022, http://zenodo.org/record/254655
Geoplana cambara Ana Laura Almeida & Fernando P. L. Marques & Fernando Carbayo 2019, SP. NOV.
No full textGEOPLANA CAMBARA ALMEIDA & CARBAYO SP . NOV . Material examined All specimens were collected in Parque Nacional de Saint-Hilaire/Lange, Matinhos/PR, Brazil. Holotype MZUSP PL 2062 (field number F1611): F. Carbayo et al., coll., 10 January 2008. Cephalic region: transverse sections on 21 slides; ovarian region: horizontal sections on 13 slides; pharynx: transverse sections on 18 slides; copulatory apparatus: sagittal sections on 15 slides. Paratype MZUSP PL 2063 (field number F1635) (immature): F. Carbayo et al., coll., 11 January 2008. Cephalic region: horizontal sections on nine slides; pharynx: sagittal sections on five slides; portion behind pharynx: sagittal sections on four slides. Paratype MZUSP PL 1009 (field number F1614): F. Carbayo et al., coll., 1 January 2008 (25°45′51.7″S, 048°37′21.6″W). Preserved in absolute ethanol. Distribution Areas covered with Atlantic forest in Parque Nacional de Saint-Hilaire/Lange, Matinhos, state of Paraná, Brazil. Etymology The specific epithet refers to the Tupi-Guarani (indigenous Brazilian tribe) name of the river flowing nearby the sampling site. Diagnosis Dorsal colour pattern constituted by a median orange stripe, bounded on either side by a thin black stripe, externally to which is a broader white stripe, and this in turn is externally bordered by a black line, externally to which is a grey band with dense white mottling, externally to which is a marginal black line. Longitudinal parenchymal muscle fibres disposed around intestine as a loose muscular tube. Diameter of muscular cylinder surrounding ejaculatory duct equivalent to 12 times the diameter of this duct. External aspect Live animals 43 mm long and 3.5 mm wide. Body elongated, margins nearly parallel; anterior end rounded, posterior pointed. Dorsum strongly convex, ventral side flat. Dorsal colour constituted by a median orange band, bounded on either side by a thin black stripe (one-third of body width) that grades into the black of the cephalic region (1/12th of body length), externally to which is a broader white stripe (one-sixth), and this in turn is externally bordered by a black line, externally to which is an area (one-sixth) coloured with a grey band with dense white mottling, externally to which is a marginal black line (Fig. 18A). Ventral body surface cream, with black margins. The dorsal colours of preserved specimens remained unchanged. Two types of eyes: a conical type, around anterior tip; and a cup-shaped type, which are located from a short distance behind anterior tip to posterior end, extending dorsally onto whitish stripes. Sensory pits simple invaginations, located ventromarginally in a single row from the very anterior end up to at least a distance equivalent to 13% of body length. Relative position mouth-to-body length, 50%. Relative position gonopore-to-body length, 64%. Internal morphology Creeping sole 80% of body width. Glandular margin absent. Three typical geoplaninid cutaneous muscle layers present, being as thick as 6% of body height. Muscle fibres of the longitudinal, innermost layer (6–12 µm thick) arranged into bundles with seven to 20 fibres each. Three parenchymal muscle layers present: dorsal layer of decussate diagonal fibres, supraintestinal layer of transverse and longitudinal fibres, and subintestinal layer with transverse and longitudinal fibres. Longitudinal fibres around intestine present, constituting a loose tube of fibres mixed up with transverse ones. Ventral nerve plate present. Mouth situated at a distance from root of the pharynx equivalent to 47% of pharyngeal pocket length. Pharynx cylindrical (Fig. 18B). Oesophagus-topharynx ratio, 17%. Outer pharyngeal musculature consisting of a subepithelial layer (5 µm) of longitudinal muscle, followed by a layer (65 µm) of intermingled circular and longitudinal muscle fibres. Inner pharynx musculature (Fig. 18C) consisting of a thin subepithelial layer of longitudinal muscle, followed by a layer (45 µm) of circular fibres and an innermost layer (10 µm) of longitudinal fibres. Testes dorsally located between supraintestinal parenchymal muscle layer and intestine; anteriormost testes slightly anterior to ovarian region, posteriormost follicles slightly anterior to root of pharynx. Penis bulb elongated, extending from 1.2 mm anterior to penis papilla to behind level of gonopore; very strongly developed dorsally to male atrium. It consists of tightly packed muscle fibres variously orientated anteriorly. Diagonal fibres on the right side of the bulb run obliquely downwards to embrace ventrally the anterior portion of the female atrium. Those fibres seem to cross the nerve plate and to anchor on ventral epidermis. Sperm ducts run dorsolaterally to the ovovitelline ducts. Laterally to ventral insertion of penis papilla, sperm ducts bend anteriorly and medially and then ventrolaterally to penetrate the bulb. The ducts subsequently join near root of penis papilla to form the ejaculatory duct. This duct traverses the central region of penis papilla to open at its tip (Fig. 18D, E). Sperm present in distal portion of sperm ducts, along their course both outside and inside penis bulb. Sperm ducts lined with a cuboidal, ciliated epithelium, pierced by glands producing fine weakly erythrophil granules; sperm ducts and anterior portion of ejaculatory duct surrounded by a circular muscle. Distal portion of ejaculatory duct surrounded by a layer of circular-to-decussate muscles. Thickness of muscles surrounding the ejaculatory duct increases from 50 µm proximally to a maximum of 150 µm in mid-papilla, and constituting a muscular cylinder with a diameter equivalent to 12 times the diameter of this duct. Penis papilla long, horizontal and cylindrical, with dorsal and ventral insertions at the same transverse plane; tip pointed (Fig. 18D, E). It occupies the entire male atrium and most of the female atrium. Total length of papilla equal to seven times its greatest diameter. Papilla covered with a cuboidal epithelium, pierced by two types of gland cells, producing erythrophil and cyanophil granules, respectively. Space immediately beneath mid-dorsal epithelium of papilla packed by necks of latter type of gland cells (Fig. 19A, B). Subepithelial musculature consisting of dense layer of circular muscle, tending to decussate distally, 185 µm thick dorsally, 145 µm ventrally, followed by a 15-µm-thick layer of longitudinal muscle. Distally, abundant decussate fibres in stroma. Male atrium ample, not folded, lined with a squamous epithelium, and underlain by a 5-µm-thick layer of circular muscles, followed by an inconspicuous layer of longitudinal fibres. Ovaries oval shaped, 425 µm in diameter in longitudinal axis, and situated at a distance from anterior end equivalent to 17% of body length. Ovovitelline ducts emerge from dorsolateral wall of ovaries. Laterally to female atrium, the oviducts curve medially and join to form common glandular ovovitelline duct. Common glandular ovovitelline duct as long as a quarter of female atrium length, after a knee-shaped bend communicating with a 30-µm-long female genital duct lined with cuboidal, ciliated epithelium, the latter opening into the female atrium. Female atrium long, not folded (Figs 18D, 19C, D) and as long as two times the male atrium. Posterior, curved section lined with 20-µm-tall stratified-like, non-ciliated epithelium and surrounded by a 30-µmthick mass of very thin, seemingly circular muscle fibres, mixed with apparently minute erythrophil granules (Fig. 19D). Anterior section of female atrium lined with cuboidal epithelium and surrounded by a 10- to 40-µm-thick layer of circular and longitudinal muscle fibres. A 10- to 110-µm-thick layer of longitudinal muscles embraces entire female atrium.Published as part of Ana Laura Almeida, Fernando P. L. Marques & Fernando Carbayo, 2019, ' Endless forms most beautiful': taxonomic revision of the planarian Geoplana vaginuloides (Darwin, 1844) and discovery of numerous congeners (Platyhelminthes: Tricladida), pp. 1-65 in Zoological Journal of the Linnean Society 185 on pages 46-48, DOI: 10.1093/zoolinnean/zly022, http://zenodo.org/record/254655
Cephaloflexa Carbayo & Leal-Zanchet 2003, comb. n.
No full textGenus Cephaloflexa Carbayo & Leal-Zanchet, 2003 Cephaloflexa nataliae (Froehlich, 1959) comb. n.Published as part of Fernando Carbayo, Marta Álvarez-Presas, Cĺaudia T. Olivares, Fernando P. L. Marques, Eud Óxia M. Froehlich & Marta Riutort, 2013, Molecular phylogeny of Geoplaninae (Platyhelminthes) challenges current classification: proposal of taxonomic actions, pp. 508-528 in Zoologica Scripta 42 on page 521, DOI: 10.5281/zenodo.39981
Luteostriata Carbayo 2010
No full textGenus Luteostriata Carbayo, 2010 Luteostriata arturi (Lemos & Leal-Zanchet, 2008) comb. n. Luteostriata pseudoceciliae (Lemos & Leal-Zanchet, 2008) comb. n.Published as part of Fernando Carbayo, Marta Álvarez-Presas, Cĺaudia T. Olivares, Fernando P. L. Marques, Eud Óxia M. Froehlich & Marta Riutort, 2013, Molecular phylogeny of Geoplaninae (Platyhelminthes) challenges current classification: proposal of taxonomic actions, pp. 508-528 in Zoologica Scripta 42 on page 522, DOI: 10.5281/zenodo.39981
Figure 25 in 'Endless forms most beautiful': taxonomic revision of the planarian Geoplana vaginuloides (Darwin, 1844) and discovery of numerous congeners (Platyhelminthes: Tricladida)
No full textFigure 25. Geoplana boraceia Almeida & Carbayo sp. nov. A, dorsal view of the living specimen MZUSP PL 1086 (field number F4358), ~40 mm in length. B, photomicrograph of a sagittal section of the pharynx of specimen MZUSP PL 2077. C, photomicrograph of a sagittal section of penis papilla and female atrium of specimen MZUSP PL 2070. D, micro-computed tomography-based image of a transverse section of the penis papilla of specimen MZUSP PL 2095. Arrows point to muscle detached from the penis bulb running towards the ventral epidermis. Right body side to the right. E, micro-computed tomography-based image of a parasagittal section of the right side of penis bulb of specimen MZUSP PL 2095. Arrows point to muscles detached from the penis bulb running towards the ventral epidermis.Published as part of Ana Laura Almeida, Fernando P. L. Marques & Fernando Carbayo, 2019, 'Endless forms most beautiful': taxonomic revision of the planarian Geoplana vaginuloides (Darwin, 1844) and discovery of numerous congeners (Platyhelminthes: Tricladida), pp. 1-65 in Zoological Journal of the Linnean Society 185 on page 58, DOI: 10.1093/zoolinnean/zly022, http://zenodo.org/record/254655
Going Beyond Counting First Authors in Author Co-citation Analysis
Get PDFThe present study examines one of the fundamental aspects of author co-citation analysis (ACA) - the way co-citation
counts are defined. Co-citation counting provides the data on which all subsequent statistical analyses and mappings
are based, and we compare ACA results based on two different types of co-citation counting - the traditional type that
only counts the first one among a cited work's authors on the one hand and a non-traditional type that takes into
account the first 5 authors of a cited work on the other hand. Results indicate that the picture produced through this non-traditional author co-citation counting contains more coherent author groups and is therefore considerably clearer. However, this picture represents fewer specialties in the research field being studied than that produced through the traditional first-author co-citation counting when the same number of top-ranked authors is selected and analyzed. Reasons for these effects are discussed
Variations on the Author
Get PDF“Variations on the Author” discusses two of Eduardo Coutinho’s recent films (Um Dia na Vida, from 2010, and Últimas Conversas, posthumously released in 2015) and their contribution to the general question of documentary authorship. The director’s filmography is characterized by a consistent yet self-effacing form of authorial self-inscription: Coutinho often features as an interviewer that rather than express opinions propels discourses; an interviewer that is good at listening. This mode of self-inscription characterizes him as an author who is not expressive but who is nonetheless markedly present on the screen. In Um Dia na Vida, however, Coutinho is completely absent form the image, while Últimas Conversas, on the contrary, includes a confessional prologue that moves the director from the margins to the center of his films. This article examines the ways in which these works stand out in the filmography of a director who offers new insights into the notion of cinematic authorship
Appropriate Similarity Measures for Author Cocitation Analysis
Get PDFWe provide a number of new insights into the methodological discussion about author cocitation analysis. We first argue that the use of the Pearson correlation for measuring the similarity between authors’ cocitation profiles is not very satisfactory. We then discuss what kind of similarity measures may be used as an alternative to the Pearson correlation. We consider three similarity measures in particular. One is the well-known cosine. The other two similarity measures have not been used before in the bibliometric literature. Finally, we show by means of an example that our findings have a high practical relevance.information science;Pearson correlation;cosine;similarity measure;author cocitation analysis
Matuxia Carbayo, Álvarez-Presas, Olivares, Marques, Froehlich & Riutort, 2013, gen. n
No full textGenus Matuxia gen. n Etymology. Matuxia is a free association of the epithetic names matuta and tuxaua and the first name of Dr. Eudóxia Maria Froehlich, who described the two species of the new genus. The gender is female. Diagnosis. Geoplaninae with medium-sized body, 45– 120 mm in length; body slender, margins nearly parallel, dorsum and ventral side slightly convex; eyes monolobulat- ed, marginally arranged along the body; pharynx cylindrical; prostatic vesicle intrabulbar, bifurcated proximally and broadly communicated with an ejaculatory cavity inside penis papilla; penis papilla apparent, with dorsal insertion posterior to the ventral; male atrium not folded, separate from the female by a fold; ascending portion of the ovovitelline ducts lateral to the gonopore canal or to the female atrium, joining each other above it; genital canal dorsoanteriorly flexed, arising from the dorsal region of the female atrium; female atrium rounded, clothed with an epithelium with multilayered aspect. Distribution. States of Rio de Janeiro and Stao Paulo, in Brazil. Type species. Geoplana tuxaua E. M. Froehlich, 1955 b Species of Matuxia. Matuxia tuxaua (E. M. Froehlich, 1955 b) * Matuxia matuta (E. M. Froehlich, 1955 b) *Published as part of Fernando Carbayo, Marta Álvarez-Presas, Cĺaudia T. Olivares, Fernando P. L. Marques, Eud Óxia M. Froehlich & Marta Riutort, 2013, Molecular phylogeny of Geoplaninae (Platyhelminthes) challenges current classification: proposal of taxonomic actions, pp. 508-528 in Zoologica Scripta 42 on page 522, DOI: 10.5281/zenodo.39981
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