1,339 research outputs found
Neoperla massevensis Zwick & Zwick 2023, n. sp.
No full text<p> <b> 75. <i>Neoperla massevensis</i> n. sp.</b> </p> <p>(Figs. 429–431)</p> <p> <b>Type material studied.</b> <b>United Republic of Tanzania:</b> ♀ <b>holotype</b> (NEOP334, slide Z19.12), <b>1</b> ♀ <b>paratype</b> (slide Z19.13): Massewe obh. Konde-Land, Anf. November 10, Dr. Fülleborn (MfNB, in alcohol).</p> <p> <b>Habitus</b>. WL 10.6mm (PT; HT damaged). Yellowish grey, no pattern, faded.</p> <p> <b>Male</b>. Unknown.</p> <p> <b>Female</b> (Fig. 429). S8 unpigmented, caudal edge almost hairless, with a very shallow notch between two short lobes. Vagina unmodified, with numerous concentric folds around the attachment of SSt which forms about 1.5 fairly wide rings of uniform width. The basal part with longitudinal folds on the concave and some brown scales on the convex side is almost half as long as the completely scaly distal part.</p> <p> <b>Egg</b> (Figs. 430–431). Size 312*186µm (n = 13). Collar short, with one row of cells, not constraining the large anchor cavity. Anchor mushroom-shaped. Numerous levogyrous striae fade away near the finely punctate rounded top of operculum. Costae are smooth and flat, sulci are exceedingly narrow, with two lines of micropunctures barely visible at 630*. The micropyles are minute holes that can be recognised their curved canal (Fig. 431, arrow), sulci do not widen around micropyles.</p> <p> <b>DNA</b> (Figs. 491–492, 498). Only the female holotype from Tanzania was sequenced with the genome-skimming approach, resulting in 7,155bp of mitochondrial, protein-coding genes. Its sister relationship to <i>N. tangana</i> <b>n. sp.</b> is very strongly supported (99.2/100/100).</p> <p> <b>Notes</b>. <i>Neoperla bareensis</i> <b>n. sp.</b> and <i>N. muyukae</i> <b>n. sp.</b> are similar to <i>N. massevensis</i> but in the former species the opening of the anchor cavity is strongly constrained, the operculum bears cells, and sulci widen around micropyles. <i>Neoperla muyukae</i> <b>n. sp.</b> differs by the prominent pattern on S8, the long SSt, and the widely open funnel-shaped anchor cavity.</p> <p>10 The year is not stated. Dr Fülleborn published on the region in 1908</p> <p> <b>Etymology</b>. The name is derived from the locality, a temporary colonial station founded in 1898, in the region of Langenburg [Tukuyu, -9.26, 33.65] (Fülleborn 1908: 293).</p>Published as part of <i>Zwick, Peter & Zwick, Andreas, 2023, Revision of the African Neoperla Needham, 1905 (Plecoptera: Perlidae: Perlinae) based on morphological and molecular data, pp. 1-194 in Zootaxa 5316 (1)</i> on pages 156-157, DOI: 10.11646/zootaxa.5316.1.1, <a href="http://zenodo.org/record/8154005">http://zenodo.org/record/8154005</a>
Neoperla (Formosita) montivaga Zwick. Not 1977
No full textNeoperla (Formosita) montivaga Zwick, 1977 Material studied: 1 ♀ Assam, Chingsao 13 vi 60 (F. Schmid; slide Z20.33; Mus.Lausanne). The original description records more than 60 straight lines of single punctures on the egg but it is unclear if they are arranged in triplets. The historical preparations are presently not available. The present new female contains no eggs. It is the first record since the original description from Bhutan and extends the range slightly. The caudal process of the vagina (Fig. 56) is shorter than in the other species.Published as part of Zwick, Peter, 2023, A new classification of genus Neoperla and systematic studies of other Perlinae (Plecoptera: Perlidae), pp. 101-131 in Zootaxa 5339 (2) on page 115, DOI: 10.11646/zootaxa.5339.2.1, http://zenodo.org/record/829716
Neoperla crustata Zwick & Zwick 2023, n. sp.
No full text<p> <b> 31. <i>Neoperla crustata</i> n. sp.</b> </p> <p>(Figs. 174–176)</p> <p> <b>Type material:</b> ♁ <b>holotype</b>, <b>Republic of Guinea</b>, piège lumineuse, sans étiquette (slide Z16/66; NEOP073; SMNS, gift J. M.Élouard).</p> <p> <b>Paratypes</b>: <b>Republic of Mali:</b> 3♁: Niger à Bamako [12.39°N, 8.00°W, 487m] 14.10.1984 J.M.Gibon; 2♁, ibidem, 14.9.94, F. Niger (SMNS, slides Z16/53+54).</p> <p> <b>Habitus</b>. WL 8.8–10.0mm. Ochre colour, legs brownish, wings infuscate. A kidney-shaped black ring around each ocellus.</p> <p> <b>Male</b> (Figs. 174–176). Generally similar to <i>N. pilulifera</i> <b>n. sp.</b> but triangular projection of T7 short. T8 flat, front as wide as the T7 process above it, narrowing caudally, with some SB. T9 unmodified, pilose humps not much raised above median furrow. Hemitergal callus tongue-shaped, tip of HT sinuous, short, sharp apex pointing forward. Hind tibiae unmodified.</p> <p>Penis straight, stout, tube distally wide, knee-shaped when endophallus is everted (Figs. 174, 176), distal armature variable. Basal half of endophallus bare, near midlength some large dorsal spines, the inflated bulbous distal third with numerous small spines in regular rows, tip in the form of a short nipple. Sides of endophallus bare, ventrally some spines in a short row. In the holotype, basolateral, dorsolateral, and dorsodistal spine groups on the outside of the penis tube are distinguishable, the other specimens have small plates, each with a projecting short spine, forming a dorsodistal crust (Fig. 175).</p> <p> <b>Female</b>. Unknown.</p> <p> <b>Egg</b>. Unknown.</p> <p> <b>DNA</b> (Figs. 491–492, 494). Only the male holotype from Guinea was sequenced with the genome-skimming approach, resulting in 9,237bp of mitochondrial, protein-coding genes. This species is very strongly supported (56.4/97/98) as sister to <i>N. amoena</i> <b>n. sp.</b> + <i>N. pilulifera</i> <b>n. sp.</b>.</p> <p> <b>Etymology</b>. The Latin adjective <i>crustata</i>, encrusted, refers to the extended dorsodistal spine patches on the penis.</p>Published as part of <i>Zwick, Peter & Zwick, Andreas, 2023, Revision of the African Neoperla Needham, 1905 (Plecoptera: Perlidae: Perlinae) based on morphological and molecular data, pp. 1-194 in Zootaxa 5316 (1)</i> on page 79, DOI: 10.11646/zootaxa.5316.1.1, <a href="http://zenodo.org/record/8154005">http://zenodo.org/record/8154005</a>
Emma Zwick
No full textEmma Zwick models a hat, coat, and dress in 1920. The Bretzman customer card lists her at 3137 Ruckle Street in Indianapolis. Census Records for 1920 show her as being 24 years old and a clerical worker in an insurance office. She was also single at the time and living with her parents, Henry F. and Aretha Zwick, and her grandmother Caroline Zwick. Henry was a traveling salesman who sold Victrolas.This image is a preservation copy made from an unstable original nitrate negative. The image is part of Series III in the Bretzman Collection
Neoperla amoena Zwick & Zwick 2023, n. sp.
No full text29. Neoperla amoena n. sp. (Figs. 161–168) Type material: Holotypus: Republic of Mali, Bafing bei Tinko (N 13°29’ 54’’ / W 10° 45’ 19’), light trap, W. Tobias, 23.9.1993 (NEOP070; SMNS). Paratypes: 1♁, 29.9.1991; 2♁, 1♀; 1♁, 2.9.1992; 6♁, 11.– 12.8.1993; 10♁, 1♀, 23.9.1993; 1♁, 24.9.1993, same locality and collector as holotype. 10♁, 1♀ (NEOP071), Republic of Guinea, 10°28’40’’N, 10°26’42’’W, Faranah, F. Niger, Somorye, 20.06.1996 (Slide Z 16/14) (SMNS, gift R. Fochetti). Republic of South Sudan: 1♁, South Sudan: Musée du Congo Soudan: Rejaf [4.75, 31.583333] 1923 L. Burgeon \ R. dét. 1647 \ Neoperla haugi Nav. P. Navás S. J. det. (NEOP072; MRAC). Additional material studied: Republic of Guinea: 1♁, Milo à Boussoulé 23.10.1984 J.M.Gibon; ♁, Riv. Milo à Kankan (bassin du Niger) 26.10.1984 J.M.Gibon. Republic of Liberia, 8 mi NW Belefuanai, S fork S.Paul R., 11.8.66, Ross & Lorenzen (CASENT 8413124, Slide Z19.32; NEOP069). Republic of Mali: 2♁, 1♀, Niger à Bamako [12.59°N, 8.06°W, 487m] 14.10.1984 J.M.Gibon (slides 16.27 & 16.192). Republic of Senegal: 1♀, Touba [14.87°N, 15.88°W] août 1982 (SMNS, gift Élouard; slide 16.196). Habitus (Fig. 161). WL of males 8.8–10.2mm (n=14), of females 12.7–13.3mm. Sexes differ in colour, male holotype bright yellow, with blackish marks. An arched fascia along occipital suture tapers towards the eyes in the middle it is connected to the ocellar mark. Occiput bright yellow, frontoclypeus weakly infuscate. Palpi and scapus yellow, pedicellus and flagellum dark brown. Pronotum light brown, wings chocolate brown, Rs with 2 terminal branches. Base of femora bright yellow, distal third and all tibiae and tarsi dark brown. Female light ochre, the turbid wings and the tibiae infuscate. Male (Figs. 162–166). From the membranous caudal area of T7 rises a dark blunt pyramidal process with some SB near tip and on underside, it projects over front of T8. T8 flat, membranous except for a caudally narrowing median sclerite band with some SB, under the T7 process. T9 with a median furrow behind the divided antecosta. Sclerites extend from the low pilose lateral humps into the furrow where they are connected, resulting in a V-shaped pattern. HT 10 in dorsal view curved, the sharp tip directed forward, mediobasal callus conical, relatively narrow, tips partly concealing the small, Y-shaped epiproct. Penis slender, straight, no spines on tube (Fig. 164). Dorsodistal membranous area with a patch of closely appressed spines directed distad (Fig., 166). Endophallus about as long as penis, wide, gently curved, distally gradually narrower. Basal half bare, distally with scattered spines, mainly dorsally. Largest spines close to base, distal spines smaller, more numerous, on almost entire diameter, some small spines also ventrally, near tip (Fig. 165). Female (Fig. 167). Generally similar to the complex. The vagina is short and wide, with only few transverse basal folds. The SSt is a wide and stout half ring. Egg (Fig. 168). Smooth, ovoid, wide short collar with distinct cells, similar to N. pilulifera. DNA (Figs. 491–492, 494). The male holotype from Mali, a male paratype from South Sudan, and a female paratype from Guinea were sequenced for the COX1 DNA barcode fragment (holotype) and with the genome-skimming approach (paratypes). They cluster together tightly with highest support (100/100/100), and the species is maximally supported (100/100/100) as sister to a specimen of N. sp. aff. amoena (NEOP 069). Together, they are very strongly supported (56.4/97/98) as sister to N. crustata n. sp.. Notes. The wide SSt is characteristic of female N. amoena. Differences in coloration concealed conspecificity of sexes until revealed by DNA. However, not all males are colourful, the less colourful ones are possily faded. Nemoura nigricauda (Cameroon) and N. bella (Lake Kivu area) with similar colouration differ by a raised hump on T8 and the blunt to spatulate hemitergal callus plus penes are very different. Etymology. The Latin adjective amoenus, pleasant, beautiful refers to the attractive appearance of some males.Published as part of Zwick, Peter & Zwick, Andreas, 2023, Revision of the African Neoperla Needham, 1905 (Plecoptera: Perlidae: Perlinae) based on morphological and molecular data, pp. 1-194 in Zootaxa 5316 (1) on pages 74-76, DOI: 10.11646/zootaxa.5316.1.1, http://zenodo.org/record/815400
Neoperla larvata Zwick & Zwick 2023, n. sp.
No full text66. Neoperla larvata n. sp. (Figs. 380–385) Type material: Holotype ♀, Republic of Sierra Leone, Pampana près de Magburaka [8.67°N, 11.96°W], 5.2.89 (slide Z16/76, NEOP 238, J.M. Élouard, SMNS). Republic of Sierra Leone, 1♀ Pampana près de Magburaka [8.67°N, 11.96°W], 5.2.89 (slide Z21/33). Central African Republic: Surroundings of Bayanga, sur plate-forme à 40 m du sol dans un kungu, à 50 m du camp 21– 23.10.2008, 03°01’49.5”N, 16°08’31.7”E, 21– 23.10.2008: 1♁ Paratype (NEOP240), 1♁ Paratype (NEOP 241), 1♀ Paratype (NEOP239). Republic of Guinea: 1♀ Paratype, Republic of Guinea, Kolenté à Simbareya [9.93°, -12.60°], 3.2.89 (NEOP237, slide Z16/59, SMNS). Additional material studied: Democratic Republic of the Congo: 1♀, Musée du Congo Yambata II/ III-1914 De Giorgi \ R. det. M 5891 (pinned slide; MRAC). République de Côte d’Ivoire: 1♀, Coll. Mus. Tervuren Côte d’Ivoire: Danané 26-VII-1966 D. Thys & W. Verheyen (pinned slide; MRAC). Republic of Ghana, 5♁, 4♀, Western Reg., Ankasa Resource Res., Nkwanta Camp 05°16.566’N 002°18342’ 7–8 Jun 2005 K.B.Miller, colr. (BYU). 1♀, Slide GH.01_10 Ghana Western region, Ankasa game prod. Reserve 10–16.12.93 St. 12 J. Kjaerendsen & T. Andersen light trap; NUFU-project ZMBN Ref. No. 3. 1♁, 1♀ (Slides Z21.41, 21.42), Ghana, Volta region, Agumatsa waterfalls, Wli, St. 10 19.11.1993 J. Kjaerendsen light trap; Ref. No. 2. 5 ♁ Ghana Volta Region, Wli Falls, Afegame, St. 10, 08.03.1993 J.S.Amakye & J. Kjaerendsen light trap; NUFU-project ZMBN Sample no. 1; 3♁, 1♀, Ghana Volta Region, Wli Falls, Afegame, St. 12, 07.03.1993 J.S.Amakye & J. Kjaerendsen light trap; NUFU-project ZMBN Sample no. 1; 1♁, 2♀ Ghana Volta region, River Menu at Mensu 18.11.1993 J. Kjaerendsen light trap. Republic of Guinea. 1♀, Guinée, Konkouré à Linsan [11.62°N, 12.66°W] 29.01.1987 (slide Z16/152); 5♀, Niandan, Bambaya, 25.10.1984 (slide Z86/12); 1♀ Guinea, Samou, Grde Chute 13.2.1986, J.M.Gibon (all SMNS, gift Élouard). République de Côte d’Ivoire: 2♀: 15 km N Lakota, 20–21 Oct.1971, black light trap, J.A.Gruwell (USNM). 2♁, Férédougouba 04.03.85 PL Baterte; 1♀, same locality, 30.3.85; 1♀, Cavally à Gu’é, Région de Toulépleu [6°35N, 8°25E] 01.02.1988 (all ex coll. Élouard, SMNS). 1♁, Elfenbeinküste, Boa ca 10 km obh. Mündung in den Sassandra, B.Statzner, 19.4.1977 (SMNS, Slide 86/24). 1♁: N'Zi Fluss oberhalb Strassenbrücke von Katiola [8.124, -5.100] nach Dabakala [8.23, -4.571], nahe Dorf Tinbé, B.Statzner, 12.1.1977 (Z21.38); 1♀: N'Zi-River, ca. 10 km N Bocauda 25.01.1977 B.Statzner; 1♀: Cavally R. (blackwater) nr Taȉ [~ 5.87°N, 7.45°W] 02.07.1977 B.Statzner (all SMNS). Republic of Liberia. 1♁, 8♀, Liberia, 8 mi NW Belefuanai [7.26N, 9.43W], S fork S.Paul R., 11.8.66, Ross & Lorenzen (CASENT 8413055); 1♀ Liberia, 36 mi S Voinjama [8.42N, 9.75S], 13.8.66, Ross & Lorenzen (CASENT 8413052). 1♁: Liberia, Suakoko, Feb. 20, 1952 [USNM, in ETOH, from photographs by W.H. Li]; Suakoko, light trap, C.C.Blickenstaff: 1♁, 19.12.51; 3♁, 31.1.52; 2♀, 14.3.52; 1♁, 1♀, 10–20.3.52; 2♁, 3♀, 3–9.5.52; 1♁, 7.11.52 (all USNM, pinned). 1♁, Liberia, Mt. Coffee, II-1897 Mrs. Sharp; 1♁, Liberia, Mt. Coffee, 27-II-1897 R.P.Currie (both in ETOH, USNM). Federal Republic of Nigeria. 1♀: 21 mi W Kaduna [10°33‘N, 7°27‘E], 650 m, 11.9.66, Ross & Lorenzen (CASENT 8413050). 1♁ (Z21.34), 1♁: Ikom [6°5’N, 8°37’E], Ogoja Prov., Nigeria. 6–7.I.1949 at light B.Malkin (Z21.34; ZMBerlin). 12♁, 1♀, NIGERIA: Benin 1-IV-75 JTMedler (BYU). Republic of Sierra Leone. 1♀, Kaba à Gotamba, Kilims Nat. Pk. 04.02.1988; 1♀, F. Rokel à Makaba [8.40652, -11.84601] 08.02.1985; 5♀, Pampana près de Magburaka [8.67°N, 11.96°W], 5.2.89 (slide Z16/76; all SMNS, ex coll. Élouard); 1♀, Northern Province, Kondembaia, 22.11.1984 W. Rossi (gift R.Fochetti; slide SMNS 98/3). Togolese Republic: 1 ♁ (Z21.40) 2 ♀ (Z21.39), Sio à Kati [6.88, 0.85], 24.11.1985; 3 ♀, Tététou [7.00, 1.50, on] (Mono R.), 25.11.1985; 2 ♀, Amoutchou, 28.11.1985; 2 ♀, R. Mono, Ngonlutu, 29.11.1985; 5 ♀, Kpassi (Mono R.), 30.11.1985; 1 ♀, R. Mono, 01.12.1985; 4 ♀, Tchamba [9.03N, 1.42E], 02.12.1985 (all V. Landa, SMNS). Habitus (Fig. 380). Male endophallus and eggs same as N. heideae n. sp., species are externally indistinguishable. Female (Figs. 381–383). Similar to N. heideae n. sp. but differs by a long and narrow tail-like section of the SSt with scattered tiny hooklets instead of normal scales like in the basal part of the SSt. The nail on S8 is conical (Fig. 381) and especially in some specimens from Ghana and Togo narrow and sharply pointed (Figs. 382–383). Variation. The relation between the length of the wide scaly part of the SSt and the narrow section with sparse spicules varies but could not be quantified. The nail is always conical but its width and also the width of the base vary. A single female from Guinea has slender eggs (NEOP237, Fig. 384) but the chorion structure is the same as in stout eggs of N. heideae n. sp. DNA (Figs. 492, 497). The female holotype from Sierra Leone, two female paratypes, and two male paratypes from Guinea and the Central African Republic were sequenced for the COX1 DNA barcode fragment, representing the wide geographic spread of this species. The monophyly of the species is very strongly supported (94.1/100/100) as is its sister relationship to N. heideae n. sp. (99.2/100/100). Etymology. The name alludes to the circumstance that the quasi-identity of males with N. heideae n. sp. for some time masked (Latin larva, a mask) the existence of the present species.Published as part of Zwick, Peter & Zwick, Andreas, 2023, Revision of the African Neoperla Needham, 1905 (Plecoptera: Perlidae: Perlinae) based on morphological and molecular data, pp. 1-194 in Zootaxa 5316 (1) on pages 143-144, DOI: 10.11646/zootaxa.5316.1.1, http://zenodo.org/record/815400
Neoperla usambara Zwick & Zwick 2023, n. sp.
No full text50. Neoperla usambara n. sp. (Figs. 278–284, 289–291) Types and aditional material taken with them: United Republic of Tanzania: Tanga region W. Usambara Mts., Mazumbai, ZMB’s Tanzania Exp, (after Andersen & Johansen 1992): Kaputu stream on Kwagoroto Hill, from 1770m down to 1400m (see Notes): Site A: 1♀, 3.– 4.11.1990; 1♀ paratype, 4.– 9.11.1990; 3♁ paratypes, 4.– 13.2.1990; 4♀, 4.– 12.2.1991. Site B: 6♀, 2.– 3.11.1990; 1♁ paratype, 1♀, 3.– 4.11.1990, ♁ holotype, 4.– 6.11.1990, 1♁, 1♀, 6.– 9.11.1990. Site bW, below/beyond waterfall, 4♁, 4♀ paratypes, 6. 12.11.1990; 1♀, 6.– 12.11.1990. Site F, 1♀ 2.– 6.11.1990. Site G, 1♁ paratype, 1♀, 30.10.– 6.11.1990. Sites I, J, L, M: 1♁, 4♀, 4.11.–54.12.1990. Site N: 1♀ paratype, 3 ♀, 20.– 26.11.1990. 1♁ paratype, United Republic of Tanzania: Tanga region W. Usambara Mts., Mpoude forest 30.11.– 6.12.1990 Malaise trap. Holotype in ZMBN, paratypes and additional material in ZMBN, some also in SMNS. Aditional material studied. United Republic of Tanzania: 1♀: Tanganjika, Usambara-Berge, Sakatani 1500m lg. Lindemann und Pavlitzki, 10.xi.1952 (ZSM, pinned slide of genitalia, no egg). Habitus. WL of males 11.7–14.2mm (n=6), of females 15.2–16.7mm. Yellowish, a dark heart-shaped macula over the ocelli. Flagellum and outer edge of tibiae dark. Wings turbid, yellowish. Male (Figs. 278–284). T7 with small erect caudal process. A broad swelling with SB on T8 opposite the vertical posterior face of T7. T8 caudally bare, weakly sclerotised, flat. T9 unmodified. HT10 short, straight, mediobasal callus rounded (Figs. 278–279). The stout penis (Figs. 280–284) has a rhomboid apex covered with small blunt teeth, the lateral corners of the rhombus project, each with some spines directed towards base. More distally the endophallus resembles a garden hose of about 25% the width of the penis tube and approximately three times as long. Endophallus with a single row of slender spines and two rows of much smaller spines. End of endophallus coiled, from its tip a narrow terminal tube (tt in Figs. 280–281) leads back into the penis, exits through the basal penis opening, and is attached to the ejaculatory duct (ed in Fig. 284). Female (Fig. 289). S8 largely brown, with a pale anchor-pattern near the caudal edge. Vagina slender, with faint basolateral sclerites and anterodorsally with a forward-directed cone formed by many concentric folds to which the SSt attaches from in front. SSt very long and narrow, entirely scaly, with two slender bands of spines projecting into the cone (arrow in Fig. 289). Egg (Figs. 290–291).Average size 365*208µm, ovoid, widest near midlength. Many straight striae with smooth costae twice as wide as the sulci.Along each side of the narrow sulcus runs one line of punctures but in the widenings around micropyles (arrow in Fig. 291) the punctation is irregular. The operculum is parabolic, only a small apical area has some cells. The sessile collar is wide and short, with one ring of cells. Anchor mushroom-shaped, the stem rises from a moderately deep bowl-shaped cavity (Fig. 290). DNA. No data. Notes. The penis of the holotype was naturally fully everted, the entire specimen was slide mounted, to preserve this condition. Neoperla usambara n. sp. is known only from the Usambara Mountains where it co-occurs with N. sambarua n. sp. The two species are similar in male and female genitalia, but eggs are very different. An expedition report (Andersen & Johansen 1992) marked 11 Malaise trap sites along Kaputu stream on Kwagoroto Hill by numbers, from 1770m down to 1400m where the stream enters a small lake. However, the Plecoptera locality labels distinguish 13 sites by letters. Most specimens of N. usambara n. sp. and N. sambarua n. sp. were collected at sites A, B, and ‘ beyond \ below waterfall ’, presumably the uppermost sites. Information to match the different codes is no longer available (T. Andersen, in a letter). Phenology of the two species agreed. Operation of emergence traps began in early November, numerous individuals were taken until mid-November, much fewer until mid-December. In January and February, after the hottest season, numbers increased again. Etymology. The species is named after the Usambara mountains, the name is a noun in apposition.Published as part of Zwick, Peter & Zwick, Andreas, 2023, Revision of the African Neoperla Needham, 1905 (Plecoptera: Perlidae: Perlinae) based on morphological and molecular data, pp. 1-194 in Zootaxa 5316 (1) on pages 112-115, DOI: 10.11646/zootaxa.5316.1.1, http://zenodo.org/record/815400
Neoperla panafricana Zwick & Zwick 2023, n. sp.
No full text<p> <b> 24. <i>Neoperla panafricana</i> n. sp.</b> </p> <p>(Figs. 133–142)</p> <p> <b>Type material</b>: <b>Republic of South Africa: Holotype</b> ♁ (NEOP158, penis slide), <b>paratype</b> ♀ (NEOP156, situs & egg slides: RSA/15 E Cape 60–90m - 31.4465S / 29.7372E Mbotyi Forest, Mkozi River, beating 14– 15.11.2013 (SMNS, gift R.Ruta). 2♁, p <b>aratypes,</b> RSA, 15.12.95, Legalameetse [24.20, 30.31; 1100m], leg. F.Koch (MfN; NEOP154, NEOP155); 13♁, 5♀, <b>paratypes,</b> Prov. Limpopo, Waterberge, Mapote Farm, 22.–24.Feb.2012, 24°07’51.41”S 28°38’40.88”E D. Bartsch (NEOP136, NEOP137, NEOP138, NEOP139, NEOP140, NEOP141, NEOP142, NEOP143, NEOP144, NEOP145, NEOP146, NEOP147, NEOP148, NEOP149, NEOP150, NEOP151, NEOP152, NEOP153; SMNS). <b>Central African Republic</b>: 1♀ <b>paratype</b> (NEOP129): 06-III-2012, nuit, chasse de nuit (18h00-00h30): banc de sable au milieu de la Sangha, M’boki (sud village Likembé, bord Sangha), [ca 2,471 964 N 16,081 238 E]. <b>Republic of Namibia</b>: Kunene R., Epupa Falls [17.00S, 13.25E], 1♁, 1♀ (NEOP133), 20.04.1994, W.Mey. <b>Republic of Senegal</b>, 1♁ <b>paratype</b> (NEOP131, Z16.89), 1♀ <b>paratype</b> (NEOP130, Z16.159), Touba [14.87°N, 15.88°W] août 1982 (gift J.M.Élouard, SMNS).</p> <p> <b>Additional material: Central African Republic</b>, near Bayanga: 1♀ (NEOP 135): 15.10.2008 18h–5h 03°03’58.3” N 16°08’59.6” E 528 m Sur plate-forme à 54 m du sol dans un Ayous (<i>Triplochiton scleroxylon</i>, Sterculiaceae) à 50 m du camp. 2♀, near Bayanga, sur plate-forme à 40 m du sol dans un kungu, à 50 m du camp 23.10.2008 18h–6h 03°01’49.5” N 16°08’31.7” E 567 m (NEOP 134). <b>Republic of Angola:</b> Angola 20.XI.13, Malanje Prov.. 2km N of Socqueco, 994m, P. Schuele leg. -16- (NEOP 132). <b>République de Côte d’Ivoire:</b> 15 ♁, riv. Férédougouba (bassin Sassandra [4.9433, -6.0850]) 12.08.1982 J.M.Gibon (SMNS). <b>Republic of Guinea</b>: 4 ♁, Guinea, Light trap, no other collecting details (SMNS, Slide 18.104, NEOP 335, gift Élouard). <b>Republic of Namibia:</b> Kunene R., Epupa Falls [17.00S, 13.25E]: 4♀, 20.02.1994; 2♀, 21.– 23.2.2008; 2♁, Namibia, Kunene River, Swaartboisdrift, 28.XI.2000 (all leg. W.Mey; MfN & SMNS). 1♁, CSIR Water Research AML218 \ Epupa falls Kaokoveld, S.S. Africa, 26.4.1970, Brown taken at light (gift J.Illies, slide Z16/005); 1♁: Kunbu-[illegible] Kavango 2.1.50 \ leg. F.Gaerdes (SMNS). <b>Republic of Senegal</b>, 3♀, Touba [14.87°N, 15.88°W] août 1982 (gift J.M.Élouard, SMNS). <b>Republic of South Africa</b>: 3♀: RSA, Transvaal, Pullen Farm, 23.–24.XI.2004, W.Mey [near Nelspruit, 25.48°S, 30.97°E] (MfNB & SMNS). 1♀ Kwa Zulu-Natal, Mooi R. [-29.21, 30.00, 1383m] (coll. Brinck). 2♁: R.S.Africa: Cape, 8 miles W Humansdorp [-34.04, 24.74], 1.3.1950, Brinck & Rudebeck (slides Z21.12 + 13). <b>Republic of Zimbabwe</b>: 2♀, Glendale [-17.362, 31.062] s.r. 26/12/56 \ C.N.Smithers [print]; 1♁: C. N. Smithers \ Beatrice [-18.27, 30.87] S. R. 26/12/56 (SMNS). <b>United Republic of Tanzania</b>: Nyassa-See, Langenburg [Lake Malawi, Tukuyu], Fülleborn S. 1♁ 16.VII.-8.VIII.98, 1♀: 20.VIII.- IX.[18]98 (MfNB).</p> <p> <b>Habitus</b>. Moderate to large size, WL 11.7–17.7mm, mean 12.5mm in males, 15.3mm in females. Similar to other members of the complex, body light ochre.</p> <p> <b>Male</b> (Figs. 135–142). Similar to <i>N. africana</i>, but process of T7 shorter, tip less sharp. HT10 short, curved little, moderately pointed, mediobasal callus tongue-shaped (Figs. 135, 136). Penis slightly sinuous, apex curving ventrad and truncate, the dorsodistal notch with small spinules is U-shaped and well delimited. The everted endophallus about as long as the penis (Figs. 137, 139), a regularly curved tapering tube, base wide. Convex face and side with many strong hooks, laterobasal hooks and those near tip smaller and stouter, concave ventral side of endophallus bare (e.g., Figs. 138, 141), or with occasional single spines (Fig. 140).</p> <p> <b>Female</b> (Figs. 124, 133–134). S8 structurally unmodified, with three brown spots or with a pale center in a heart-shaped brown macula. The vagina is supported by sclerotised lateral folds: near the gonopore the folds are straight, parallel and stand close together, in front they curve outward and surround a wider area, the ensemble resembles a racket or a calyx (Fig. 124). SSt short and forming only a ring fragment, the modified basal section with longitudinal folds and scales only on the convex side is long. The scale cover widens caudally, often appears a projecting angle (ap in Fig. 134).</p> <p> <b>Egg</b> (Figs. 133–134). Slender ovoid, size varies, 370–445 * 190–240µm, on average 410*218µm and 1.9 times longer than wide. Nine to 11 straight low ridges blend into the base of the operculum. The operculum is a blunt cone or almost rounded. The well-developed collar bears cells. Chorion punctures are as fine as in <i>N. camerunensis</i>.</p> <p> <b>DNA</b> (Figs. 491–492, 495).A total of 30 specimens from Senegal, the Central African Republic, Namibia, Angola and the Republic of South Africa were sequenced, representing most of the geographic spread of this species and providing very strong support (98.2/100/97) for a species complex that also includes <i>N. camerunensis</i> (Enderlein). While sequence variation is distinct and multiple lineages are indicated, specimens from different localities within the Republic of South Africa differ little in their DNA sequences and cluster with similarly looking specimens from Namibia, Angola, and Senegal, but also <i>N. camerunensis</i> (Enderlein). Both sexes are well represented.</p> <p> This species and <i>N. camerunensis</i> (Enderlein) form a complex that cannot be resolved with current molecular characters and should be investigated further with a much broader range of nuclear markers than just the mitochondrial COX1. The male holotype from the strikingly homogeneous South African population was sequenced with the genome-skimming approach and obtained 11,001bp of mitochondrial, protein-coding genes. This sequence is clearly different from the sequences of specimens collected in Cameroon, and the disagreement between molecular and morphological data is unlikely to affect the validity of both species but will probably require the description of further species in future.</p> <p> <b>Notes and variation</b>. The characteristically shaped vaginal sclerites are an apomorphy for <i>N. panafricana</i>. The range is disjunct and there is no evidence of contact between the northwestern and the southern populations. <i>Neoperla panafricana</i> is the only South African representative of the <i>africana</i> -complex, Picker’s figures of eggs suggest his <i>N.</i> spec. C (Picker 1980, his figs. 13, 16) is <i>N. panafricana</i>. The species is uniform across the huge, divided range, only the 2♁ from the extreme SW (Humansdorp) differ in that there are only very few tiny spinules in the dorsodistal notch of the penis tube.</p> <p> <b>Etymology</b>. The Latin adjective <i>panafricana</i> refers to the wide distribution of the species over much of the Ethiopian Region.</p>Published as part of <i>Zwick, Peter & Zwick, Andreas, 2023, Revision of the African Neoperla Needham, 1905 (Plecoptera: Perlidae: Perlinae) based on morphological and molecular data, pp. 1-194 in Zootaxa 5316 (1)</i> on pages 64-67, DOI: 10.11646/zootaxa.5316.1.1, <a href="http://zenodo.org/record/8154005">http://zenodo.org/record/8154005</a>
Prionocyphon papuanus Zwick 2015, nov.sp.
No full textPrionocyphon papuanus nov.sp. (Figs 10-12) H o l o t y p e: Irian Jaya: Jayawijaya Langda 27.- 28.8.1992 leg. A.Riedel 2100-2300m (SMNS). Habitus. BL 2.2mm, BL/BW ~1.5, HCW corresponds to 51% of BW. Regularly oval, fairly flat. Dorsal surface light brown, legs and antennomeres 1-3 yellowish, antennomeres 4 and 5 infuscate, more distal segments missing. Normal punctures on entire surface, those on head and pronotum equal, finer than on elytra. Pilosity semi-erect, a bit shaggy, yellowish. The front of clypeus is straight in the middle above the labrum. On each side of the labrum the clypeus forms a rounded plate extending forward next to the flat scape which is distinctly larger. Scape flat, with sharp front edge. Pedicel small, inserted under the edge of the scape. Antennomere 3 minute, 4 (which tends to be the most slender antennal segment) only about twice as long as wide, antennomere 5 shorter and stouter, both with round cross section, neither flattened nor serrate. SAR ending where it meets the eye. Lower face of head and thorax typical of the genus, head with small but distinct buttonhole-configuration. Prosternal process slender, about drop-shaped. Receiving mesoventral groove wide, U-shaped. Pilosity differs between abdominal sternites. It is sparse and mainly composed of small sensilla with large insertion ring on S3. S4 bears a mixture of normal setae and sensilla. Numbers change successively on posterior sternites, S7 is densely covered by only setae. T7 is caudally wider than S7 and has two short apodemes. It bears the spiracle and is largely covered with hair-like microtrichia directed mediad on the sides, caudad along a middle strip. Near the caudal edge, much finer microtrichia are directed backward, forming a dense fringe along the edge. Very small true hairs with insertion rings are interspersed in this caudal region. M a l e. T8 with short nearly straight stout apodemes connected by a strong basal sclerite, plate transverse, short. Caudal half and rear edge covered with microtrichia and minute socketed setae, a few longer setae along edge. S8 not developed. Apodemes of T9 more slender, converging but not connected. Plate entirely membranous, pale, hairless (Fig. 10). S9 (half missing, Fig. 11) with slender base supported by paired sclerites, caudal half tongue-shaped, medially divided, densely covered with setae. Tegmen and pala connected, forming a single compact ovoid structure with re-enforced rounded front edge (Fig. 12). Parameres and styli originate laterally in the basal fourth at the level where trigonium and parameroids also originate. The parameres are slender sinuous rods with some apical setae, the styli are weak, long and straight processes with apical microtrichia-like frazzles. The contour of the large parameroids determines the ovoid overall shape of the genitalia. The parameroids wrap around the base of trigonium, caudally they lie along its sides. The trigonium is a flat bottle-shaped structure. The truncate narrow apex is divided, each lobe slightly extended sideways and beset with single large conical teeth. The endophallus shines through the base of the trigonium. On each side appears a pack of dense slightly divergent folds, possibly colourless slender teeth. More distally the endophallus is exposed on either side of the narrow part of trigonium. It shows numerous very delicate parallel folds (Fig. 12 is diagrammatic in this respect!) which curve at the top where they are oriented centripetally, apparently around the terminal opening. F e m a l e. Unknown. Notes. The frontoclypeal lobes are shared with Australian Prionocyphon species (for illustrations, see WATTS 2010, ZWICK in prep.) but also with other genera, e.g., Mescirtes. Several of the Australian species also exhibit an intimate connection between tegmen and penis, and an armed endophallus with visible caudal opening (ZWICK, in prep.). The pattern of abdominal pilosity is close to the heterogenous pattern of many Australian species. In Table 1 of ZWICK (in prep.) the new New Guinean species would stand a few lines beneath P. storeyi WATTS, 2010. E t y m o l o g y. The species name is an adjective describing the origin of the beetle.Published as part of Zwick, Peter, 2015, Three new Marsh Beetles (Col.: Scirtidae) from New Guinea and Java, pp. 1885-1895 in Linzer biologische Beiträge 47 (2) on pages 1889-1891, DOI: 10.5281/zenodo.528626
FIGURES 50–56 in A new classification of genus Neoperla and systematic studies of other Perlinae (Plecoptera: Perlidae)
No full textFIGURES 50–56. Neoperla (Formosita) spp. of the fasciata-group. Holotype of N. (F.) fasciata new species: 50, head and pronotum; 51–52, vagina in dorsal and lateral views. Neoperla (F.) sp., cf. N. (F.) bimaculata Li et al. 2021: 53, egg; 54, head and pronotum; 55, vagina, dorsal view; 56, N. (F.) montivaga Zwick. Not to scale. od, common oviduct; sa, spermatheca; sd, spermathecal duct; v, vagina. 56 from Zwick (1977), the others are originals.Published as part of Zwick, Peter, 2023, A new classification of genus Neoperla and systematic studies of other Perlinae (Plecoptera: Perlidae), pp. 101-131 in Zootaxa 5339 (2) on page 113, DOI: 10.11646/zootaxa.5339.2.1, http://zenodo.org/record/829716
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