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The influence of the physical environment on invertebrate larval transport, settlement and recruitment with insights on early-stage physiological performance
Get PDFMost marine benthic organisms have a biphasic life cycle whereby the larvae are pelagic, and their distribution, transport and settlement are largely controlled by environmental factors. Among these factors, hydrodynamics have key direct effects. Nearshore water hydrodynamics are strongly influenced by wind. This study investigated whether wind indirectly affects the distribution, settlement and recruitment of invertebrate marine larvae. Additionally, larvae are exposed to a more constant, benign environment than they will experience after settlement, and therefore the possible effects of larval thermal history on the physiological plasticity of settled organisms was also estimated for possible long-term insights into population dynamics. Four sites, two on the western and two on the eastern side of Algoa Bay in Gqeberha (formerly known as Port Elizabeth), South Africa, were chosen for larval collection. Sites at opposite ends of the bay were selected as they were towards either side of the dominant wind directions in this region. At each site, two sampling stations were identified at 300 and 900 metres offshore, with station replicates 300 metres apart. Sampling of marine invertebrate larvae was conducted after two days of westerly or easterly winds at either of the two western or eastern sites. The number of hours of unidirectional wind determined the prevalent wind on the two days prior to sampling. Samples were collected with a plankton pump at three depths; surface, mid-depth and bottom. Simultaneously with plankton sampling, water properties (fluorescence, temperature, zonal and meridional flow, salinity and oxygen) were measured. Settlement/recruitment samples of mussels and barnacles were collected at six sites, two at either edge of the bay and two within the center of the bay. Plastic scouring pads and PVC plastic plates covered with safety-walk, placed at least 30 cm from each other, were used to collect settlers/recruits of mussels and barnacles, respectively. The pads and plates were replaced on a monthly basis for 14 months to quantify monthly settlement/recruitment. At four of the six sites used for quantifying settlement/recruitment, settlers and recruits were collected to measure the physiological responses of these two ontogenetic stages (settlers and recruits) to acute temperature variations which they might experience in the intertidal zone. Three temperature loggers were used to record real time temperature variation at each site over the period (one month) in which the artificial collectors were in position. Factorial ANOVA was used to investigate the effects of wind, depth, site, and distance from the shore on the abundance of larvae in the water column, as well as the effects of month and site on settlement/recruitment. Additionally, multiple regression analyses were used to investigate the effects of physical parameters, including upwelling and dissipation of turbulent kinetic energy, on the abundance of both larvae, settlers and recruits. Distance-based linear models and redundancy analyses were also carried out on the abundances of invertebrate larvae. Correlation analyses were performed to investigate the effect of the number of hours of directional wind on settlement. Lastly, correlation analyses between wind and currents were carried out. The results revealed a link between winds and currents, with wind speed directly proportional to surface current speed, which, as expected, decreased with depth. Current direction was, however, not perfectly aligned with wind direction. Larval distribution, settlement and recruitment were largely associated with the nearshore dissipation of turbulent kinetic energy and upwelling. Thus, overall larval, settler and recruit abundances were indirectly affected by wind. Physiologically, there was no significant difference in oxygen consumption between the juvenile ontogenetic stages (settlers and recruits) of mussels. Recruits exposed to average and maximum temperatures, however, consumed more oxygen than those exposed to low temperatures. Wind and wind-mediated currents as well as temperature have been shown to shape the distribution and shore supply of larvae, and this study contributes to the broad knowledge of population dynamics and replenishment, and the tight relation of abiotic factors affecting biological processes on the shore.Thesis (PhD) -- Faculty of Science, Zoology and Entomology, 202
Case in modern standard Arabic
Get PDFThis thesis provides a novel account of case checking in Modern Standard Arabic (henceforth MSA). It argues against the concept that nominative case in topic and comment structures is default. Default or inherent case is not related to case filter, nor is it assigned by any syntactic method. Some linguists claim that the default case in Arabic language is employed only when no case assigner is available. I argue, in light of the minimalist program (henceforth MP), that default (inherent) case is always a problem for syntactic theory and that the nominative case which the topic and comment sentences bear is valued by a functional head. The topic and comment constructions contain a predicational head (Pred)- a functional head, that is equivalent to vP which occurs in verbal constructions. The head of PredP is accountable for nominative case value on the topic and comment structures. This thesis also provides a novel paradigm regarding the clitics that appear at the end of verbs in MSA. This paradigm proves that these clitics are not agreement, tense, nor mood markers but rather Verbal Case markers and it provides a minimalist program account to explain case checking on verbs in MSA. Regarding sentence initial DP, and contra to the proposal that in SVO sentence initial DP is a subject, I have scrutinised the nature of this DP and provided enough evidence that it is a topic. Additionally, I have investigated the relationship between agreement asymmetry and the position of the subject in MSA from the minimalist feature inheritance account viewpoint. Mainly, I reviewed null pro hypothesis which proved to be redundant to account for the derivation of VSO or SVO word orders. I have also proved what were considered agreement markers are not agreement markers but resumptive pronouns. Finally, I proved that there is not agreement asymmetry in MSA. Agreement pattern in MSA results from the agree operation and it is achieved under Probe-Goal alignment; regardless of whether the Goal is raised up to Spec-TP or remains in situ (Spec-VP), the agreement is not supposed to change. This conclusion is in harmony with the principles of MP Agree Theory.Thesis (PhD) -- Humanities, Linguistics and Applied Language Studies, 202
It's all in your head
Get PDFMy thesis is a collection of short stories that delve into the multifaceted nature of death, with a specific emphasis on murder, violence, death and their profound aftermath, including themes of loss, grief, and trauma. These stories offer diverse perspectives, narrated by perpetrators, voiceless victims, and those left to pick up the shattered pieces. Additionally, some stories are seen through the eyes of those tasked with handling the bodies. Often set in unnamed locations, they delve into imaginary and fantastical worlds, while remaining grounded in recognisable situations and spaces. I draw inspiration from popular and genre fiction such as horror, crime fiction and true crime stories, but approach my writing from a psychological lens, employing stylistic experimentation to challenge readers' expectations. The power of silence is a recurring motif. Rather than focusing on unearthing facts or revealing the "truth" like crime fiction often does, my narratives delve into what isn't or at times cannot be told — the unsayable. Some of my stories explore silence inherent in violence, grief, and the inability to articulate one's experience in the face of a violent act or a life prematurely ended. Others delve into the silence of untold stories and the dark secrets of the perpetrators. By exploring these contrasting perspectives, I aim to offer a nuanced exploration of death and its aftermath. The writing styles of Stephen Graham Jones, Lydia Davis, and Kuzhali Manickavel influence my work. Lydia Davis, known for her mastery of very short, flash fiction, ranging from single sentences to a paragraph or two, inspires me with her precise observations of the human condition. Her minimalist prose, carefully selecting and arranging words and sentences, encapsulates the power of less-is-more storytelling. Manickavel creates surreal yet tangible worlds, combining idiosyncratic, intense and eerie elements with unfiltered expression. Drawing from her ability to blur the lines between the surreal and the real, I infuse my stories with a sense of disquieting authenticity. Stephen Graham Jones stands out for his ability to explore morbid themes in a compelling and unconventional manner. His experimentation with horror fiction tropes, the visceral realism of his prose, and his complex characters inspire me to capture the unsettling feeling that something dreadful has occurred without explicitly detailing the facts and intricacies.Thesis (MA) -- Faculty of Humanities, School of Languages and Literatures, 202
The effect of Gracilaria gracilis (Rhodophyta) on growth and behaviour of farmed abalone Haliotis midae when included in the abalone diet
Get PDFAbalone feed naturally on algae, but commercial abalone farms rely on formulated feed. The inclusion of algae with the formulated feed in the diet improves growth, health and feed conversion ratio (FCR). However, the kelp that is currently included in the feed is wild harvested and this resource is limited and under increasing pressure. Thus, there is a need to develop an alternative sustainable source of algae. The aim was to use Gracilaria gracilis algae (that was produced with mussel in an integrated multitrophic aquaculture system; IMTA) which was then included into abalone feeds and its influence on abalone growth, FCR and behaviour was evaluated. Sea-based G. gracilis was harvested off an existing IMTA system, manufactured into Abfeed® and fed to abalone to determine growth efficiency. Abalone (30 - 40 g abalone-1) originated from the same cohort at Whale Rock Abalone Farm (Pty) Ltd., Hermanus. Abalone were fed either: Abfeed® S34 only with no algae (G0, the control); Abfeed® with sea-based IMTA G. gracilis included in the formulated feed at 0.75 % (G0.75); 1.50 % (G1.50); 3.00 % (G3.00); 6.00 % (G6.00); and 12.00 % (G12.00) inclusion; and Abfeed® S34 with fresh-live G. gracilis (cultured at Whale Rock Abalone Farm (Pty) Ltd.; Gfresh + S34s). Abalone shell length, whole body mass and FCR were measured at a four-months interval over eight months (two growth cycles). Differences in behaviour (i.e., level of activity and feeding activity) were also monitored and comparing between treatments. There was significant difference in mean whole-body mass between treatments (RMANOVA, F(6, 29) = 3.71, p = 0.007). Abalone fed the diet with the highest inclusion of dry G. gracilis (G12.00) had a lower mean whole-body mass value (74.13 ± 2.94 g abalone-1) than abalone from all the other diets after eight months. There was a negative relationship between the condition factor and dry G. gracilis inclusion rate for the first growth cycle (y = - 0.0044x + 1.222, R2 = 0.24, p=0.009) and second-growth cycle (y = - 0.0096x + 1.2233, R2 = 0.51, p = 0.00002). Abalone receiving fresh G. gracilis were less quiescent and more alert on the first hours after food distribution (p < 0.05). The same proportion of abalone with G0, G0.75, and G6.00 G. gracilis inclusion were feeding all along the 9 hours post-feeding. This study will potentially reduce the dependence of abalone farms on wild-harvested kelp in formulated feed. This will improve their contribution to a more environmentally sustainable global production of sea food products, making them more competitive on international markets.Thesis (MSc) -- Faculty of Science, Ichthyology and Fisheries Science, 202
Computational design for genome reduction of Lactococcus lactis towards microbial chassis development
Get PDFAccess restricted. Expected release date 2026.Thesis (MSc) -- Faculty of Science, Biotechnology Innovation Centre, 202
Developing a community of practice to promote the use of biological control in the integrated management of Prosopis in South Africa
Get PDFProsopis spp. are non-native species present in the arid parts of southern Africa. These trees originated in the Americas and were first introduced as a source of fodder, shade, and wood, but have become invasive, negatively impacting local biodiversity, and disrupting the delivery of ecosystem services. Some species of Prosopis hybridise freely, complicating identification and subsequent control. The control of Prosopis in the Northern Cape Province of South Africa is still contentious, because of the perceived benefits to some landowners. The effectiveness of biological control agents to control the spread of Prosopis has been quantified, more damaging agents have been considered in recent years because of the continued increase in Prosopis density. Research into the establishment of a Community of Practice (CoP) to address the differences in perceptions regarding the control of Prosopis in the Northern Cape Province is considered an appropriate starting point to conceptualise the challenges to the successful integrated management of Prosopis. As farmers are the main stakeholders involved with Prosopis control, understanding the role of farmers and the functioning of farming enterprises in the Northern Cape is imperative for the development of both a CoP as well as the appropriate management of Prosopis. To foster engagement with as many stakeholders as possible, workshops dealing with biological control as part on an integrated approach to the control of Prosopis were held in towns in the Northern Cape Province: Groblershoop, Brandvlei, Kenhardt, Upington, Prieska and Williston. Some of the main concerns of the landusers include the host specificity of the released biocontrol agents, as well as increased transparency and communication. This is especially in regards to the research on present and future biological control agents that will possibly improve the perceptions of stakeholders. The mechanical harvesting and use of biomass as a control method was supported, especially where farmers were removing Prosopis from their properties, leaving large amounts of biomass in the veld. Continued research into Prosopis in the Northern Cape needs to consider the possibility of secondary invasions, especially in areas where native bush encroachment is already a concern. The impact of clearing in terms of natural grazing and animal improvement needs to be quantified, to conceptualise the importance of maintaining better veld quality versus using Prosopis for fodder. Biological control targeting not only the reproductive output of the trees, but also the standing biomass has support from the majority of the land users in the Northern Cape Province following these workshops and this study shows the importance of a socio-ecological approach to the control of landscape scale invasion.Thesis (MSc) -- Faculty of Science, Zoology and Entomology, 202
Functional significance of mixed-species groups for zebra (Equus quagga) in savanna habitats
Get PDFGlobally, herbivore populations are facing decline, which threatens their crucial role within the ecosystems they inhabit. Herbivores are essential to ecosystem structure and function because they act as prey for higher trophic levels, cycle soil nutrients, and alter vegetation structure. Understanding the drivers that shape herbivore habitat use and selection are therefore critical for making informed conservation and management decisions. Like all prey species, herbivores must balance a trade-off between acquiring resources and avoiding predation which is influenced by a number of species-specific intrinsic traits and various ecological drivers. Herbivores can also track risk over space and time, and certain habitat structures and landscape features can influence risk perception; this is termed the landscape of fear. To mitigate the risk-reward trade-off, herbivores employ a suite of behavioural responses. Group formation is likely the most well-documented of these responses. However, grouping results in the information-competition trade-off, wherein group members may obtain valuable information regarding the location of resources and/or the risk of predation, but may suffer from increased competition, especially among larger group sizes. Mixed-species groups—an assemblage of two or more species in close spatial association—have the potential to mitigate some of the competitive costs of larger conspecific groups and enhance the anti-predator and/or foraging benefits of grouping. African savannas provide a particularly useful natural laboratory to study mixed-species groups, due to the high diversity of prey species and habitat heterogeneity present. Previous studies have investigated the mixed-species effect within these systems, but have yet to quantify some of the identity-dependent costs and benefits associated with mixed-species herding. Thus, the overall aim of my study was to quantify the functional significance of mixed-species groups for zebra (Equus quagga) herding with two of their most common herding partners, impala (Aepyceros melampus) and wildebeest (Connochaetes taurinus). My study was conducted at multiple spatial scales in the southern portion of the Kruger National Park, South Africa between 2021 and 2022. I found that mixed-species herding altered zebra landscape use, resulting in a quantifiable intensification of habitat use compared to when they herd with conspecifics. I did not observe significant habitat expansion for zebra in mixed-species herds compared to single-species herds, likely because the anti-predator benefits of mixed-species herding convey enhanced foraging opportunities in already-safe habitats. Zebra foraging with impala achieved greater foraging success and experienced lower levels of competition, compared to zebra foraging with wildebeest. However, compared to foraging with conspecifics, the levelled landscape of fear zebra experienced when foraging with wildebeest resulted in increased foraging success across the landscape. My findings highlight the context-dependency of the costs and benefits associated with different herding partners for zebra, and suggest that in mixed-species herds with wildebeest, zebra experience increased potential fitness benefits compared to herding with impala or conspecifics. Overall, the findings of my study suggest that mixed-species herding is an important tool in the suite of anti-predator behaviours employed by prey species, and one with significant implications for survivorship, habitat use, and ultimately, community level interactions.Thesis (MSc) -- Faculty of Science, Zoology and Entomology, 202
Acacia dealbata invasion and clearing in Upper Tsitsana communal areas of South Africa: human perceptions and ecological impacts
Get PDFThe number of invasive alien plants in South Africa and globally continue to increase mostly due to increased human movement and climate change. It is estimated that approximately 10 000 hectares of South Africa’s terrestrial land is invaded by invasive alien species. Recent studies have reported that invasion by invasive alien plants cost South Africa billions of Rands every year through loss of water and ecosystem services. Although much has been done to understanding invasion trends, patterns, and processes in South Africa, little is knowns regarding human benefits linked to alien plant invasion as well as human perceptions regarding management interventions, particularly clearing by the national Working for Water programme. This thesis took a socio-ecological approach to assess (i) local people perceptions and knowledge of Acacia dealbata invasion and clearing, and (ii) extent of grazing grass diversity and composition recovery following A. dealbata clearing. To answer the above-mentioned objectives, both qualitative (face-to-face interviews) and quantitative (vegetation surveys) methods were used in grasslands located in upper Tsitsana catchment, South Africa. Results from the 165 interviews conducted in six villages show that most villagers are aware of A. dealbata in the area, but do not know that it is an invasive alien plant. Respondents accrue several benefits from the plant that include firewood (98% across all villages), construction pools (34% across all villages), and shade (3% across all villages). Respondents identified several costs associated with A. dealbata invasion, such as roots damage to houses (37% across all villages), hindering growth of crops (15% across all villages), and takes up yard space (29% across all villages). Most of the villages ranked the effects of A. dealbata on grazing as highly severe. Villages indicated that the Working for Water clearing programme presents them with socio-economic benefits such as employment and money. Some villagers noted that the clearing of A. dealbata in the area restores grazing grass (62% across all villages) and water (32% across all villages). Based on these interview survey results, the study concludes that A. dealbata has both costs and benefits that support local people’s livelihoods; therefore, its management should consider views for local users. Grass surveys conducted in 1 m2 quadrats that were replicated 15 times across four paired A. dealbata cleared and grassland sites showed that the removal of A. dealbata facilitates the recovery of native grasses. Of the 14 different grazable grass species that were identified across all quadrats only one (Bulbostylis sp.) is not palatable to livestock. Although grass cover was significantly (p < 0.05) higher in the grasslands than the cleared patches, grass richness was significantly (p < 0.05) higher in the cleared patches an indication that recovery of different grasses is taking place. Based on these vegetation survey results; the study concludes that recovery of grazing grasses after A. dealbata clearing four years ago is following a positive trajectory as the cleared sites are looking like the grassland sites. Overall, the study recommends that management of A. dealbata in Tsitsana landscape requires a socio-ecological approach to understand user benefits whilst at the same time developing measure to control the plant. The study has provided evidence that A. dealbata clearing facilitates passive restoration of grazing grass and local villages are aware of these restoration benefits.Thesis (MSc) -- Faculty of Science, Environmental Science, 202
The influence of electronic-consumer experience (e-CX) on consumer post-purchase outcomes of South African e-commerce retail organisations
No full textAccess restricted. Expected release in 2026.Thesis (PhD) -- Faculty of Commerce, Management, 202
A critical analysis of the factors influencing knowledge sharing in Indian family owned businesses in South Africa
Get PDFAccess restricted. The expected release date is 2025.Thesis (PhD) -- Faculty of Commerce, Management, 202