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Annotated type catalogue of the Bulimulidae (Mollusca, Gastropoda, Orthalicoidea) in the Natural History Museum, London
The type status is described of 404 taxa classified within the family Bulimulidae (superfamily Orthalicoidea) and kept in the London museum. Lectotypes are designated for Bulimus aurifluus Pfeiffer, 1857; Otostomus bartletti H. Adams, 1867; Helix cactorum d’Orbigny, 1835; Bulimus caliginosus Reeve, 1849; Bulimus chemnitzioides Forbes, 1850; Bulimus cinereus Reeve, 1849; Helix cora d’Orbigny, 1835; Bulimus fallax Pfeiffer, 1853; Bulimus felix Pfeiffer, 1862; Bulimus fontainii d’Orbigny, 1838; Bulimus fourmiersi d’Orbigny, 1837; Bulimus (Mesembrinus) gealei H. Adams, 1867; Bulimus gruneri Pfeiffer, 1846; Bulimus humboldtii Reeve, 1849; Helix hygrohylaea d’Orbigny, 1835; Bulimus jussieui Pfeiffer, 1846; Bulimulus (Drymaeus) binominis lascellianus E.A. Smith, 1895; Helix lichnorum d’Orbigny, 1835; Bulimulus (Drymaeus) lucidus da Costa, 1898; Bulimus luridus Pfeiffer, 1863; Bulimus meleagris Pfeiffer, 1853; Bulimus monachus Pfeiffer, 1857; Bulimus montagnei d’Orbigny, 1837; Helix montivaga d’Orbigny, 1835; Bulimus muliebris Reeve, 1849; Bulimus nigrofasciatus Pfeiffer in Philippi 1846; Bulimus nitelinus Reeve, 1849; Helix oreades d’Orbigny, 1835; Helix polymorpha d’Orbigny, 1835; Bulimus praetextus Reeve, 1849; Bulinus proteus Broderip, 1832; Bulimus rusticellus Morelet, 1860; Helix sporadica d’Orbigny, 1835; Bulimus sulphureus Pfeiffer, 1857; Helix thamnoica var. marmorata d’Orbigny, 1835; Bulinus translucens Broderip in Broderip and Sowerby I 1832; Helix trichoda d’Orbigny, 1835; Bulinus ustulatus Sowerby I, 1833; Bulimus voithianus Pfeiffer, 1847; Bulimus yungasensis d’Orbigny, 1837.
The type status of the following taxa is changed to lectotype in accordance with Art. 74.6 ICZN: Bulimulus (Drymaeus) caucaensis da Costa, 1898; Drymaeus exoticus da Costa, 1901; Bulimulus (Drymaeus) hidalgoi da Costa, 1898; Bulimulus (Drymaeus) interruptus Preston, 1909; Bulimulus (Drymaeus) inusitatus Fulton, 1900; Bulimulus latecolumellaris Preston, 1909; Bulimus (Otostomus) napo Angas, 1878; Drymaeus notabilis da Costa, 1906; Drymaeus notatus da Costa, 1906; Bulimulus (Drymaeus) nubilus Preston, 1903; Drymaeus obliquistriatus da Costa, 1901; Bulimus (Drymaeus) ochrocheilus E.A. Smith, 1877; Bulimus (Drymaeus) orthostoma E.A. Smith, 1877; Drymaeus expansus perenensis da Costa, 1901; Bulimulus pergracilis Rolle, 1904; Bulimulus (Drymaeus) plicatoliratus da Costa, 1898; Drymaeus prestoni da Costa, 1906; Drymaeus punctatus da Costa, 1907; Bulimus (Leptomerus) sanctaeluciae E.A. Smith, 1889; Bulimulus (Drymaeus) selli Preston, 1909; Drymaeus subventricosus da Costa, 1901; Bulimulus (Drymaeus) tigrinus da Costa, 1898; Drymaeus volsus Fulton, 1907; Drymaeus wintlei Finch, 1929; Bulimus zhorquinensis Angas, 1879; Bulimulus (Drymaeus) ziczac da Costa, 1898.
The following junior subjective synonyms are established: Bulimus antioquensis Pfeiffer, 1855 = Bulimus baranguillanus Pfeiffer, 1853; Drymaeus bellus da Costa, 1906 = Drymaeus blandi Pilsbry, 1897; Bulimus hachensis Reeve 1850 = Bulimus gruneri Pfeiffer, 1846 = Bulimus columbianus Lea, 1838; Bulimus (Otostomus) lamas Higgins 1868 = Bulimus trujillensis Philippi, 1867; Bulimulus (Drymaeus) binominis lascellianus E.A. Smith, 1895 = Bulimulus (Drymaeus) binominis E.A. Smith, 1895; Drymaeus multispira da Costa, 1904 = Helix torallyi d’Orbigny, 1835; Bulimulus (Drymaeus) plicatoliratus Da Costa, 1898 = Bulimus convexus Pfeiffer, 1855; Bulimus sugillatus Pfeiffer, 1857 = Bulimus rivasii d’Orbigny, 1837; Bulimus meridionalis Reeve 1848 [June] = Bulimus voithianus Pfeiffer, 1847.
New combinations are: Bostryx montagnei (d’Orbigny, 1837); Bostryx obliquiportus (da Costa, 1901); Bulimulus heloicus (d’Orbigny, 1835); Drymaeus (Drymaeus) lusorius (Pfeiffer, 1855); Drymaeus (Drymaeus) trigonostomus (Jonas, 1844); Drymaeus (Drymaeus) wintlei Finch, 1929; Drymaeus (Mesembrinus) conicus da Costa, 1907; Kuschelenia (Kuschelenia) culminea culminea (d’Orbigny, 1835); Kuschelenia (Kuschelenia) culmineus edwardsi (Morelet, 1863); Kuschelenia (K.) gayi (Pfeiffer, 1857); Kuschelenia (Kuschelenia) tupacii (d’Orbigny, 1835); Kuschelenia (Vermiculatus) anthisanensis (Pfeiffer, 1853); Kuschelenia (Vermiculatus) aquilus (Reeve, 1848); Kuschelenia (Vermiculatus) bicolor (Sowerby I, 1835); Kuschelenia (Vermiculatus) caliginosus (Reeve, 1849); Kuschelenia (Vermiculatus) cotopaxiensis (Pfeiffer, 1853); Kuschelenia (Vermiculatus) filaris (Pfeiffer, 1853); Kuschelenia (Vermiculatus) ochracea (Morelet, 1863); Kuschelenia (Vermiculatus) petiti (Pfeiffer, 1846); Kuschelenia (Vermiculatus) purpuratus (Reeve, 1849); Kuschelenia (Vermiculatus) quechuarum (Crawford, 1939); Naesiotus cinereus (Reeve, 1849); Naesiotus dentritis (Morelet, 1863); Naesiotus fontainii (d’Orbigny, 1838); Naesiotus orbignyi (Pfeiffer, 1846); Protoglyptus pilosus (Guppy, 1871); Protoglyptus sanctaeluciae (E.A. Smith, 1889).
Type material of the following taxa is figured herein for the first time: Bulimus cinereus Reeve, 1849; Bulimus coriaceus Pfeiffer, 1857; Bulimulus laxostylus Rolle, 1904; Bulimus pliculatus Pfeiffer, 1857; Bulimus simpliculus Pfeiffer, 1855.This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.NHM Repositor
XIX International Botanical Congress, Shenzhen: report of the Nomenclature Section, 17th to 21st July 2017
This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.NHM Repositor
Multidisciplinary investigation of two Egyptian child mummies curated at the University of Tartu Art Museum, Estonia (Late/Graeco-Roman Periods)
Two ancient Egyptian child mummies at the University of Tartu Art Museum (Estonia) were, according to museum records, brought to Estonia by the young Baltic-German scholar Otto Friedrich von Richter, who had travelled in Egypt during the early 19th century. Although some studies of the mummies were conducted, a thorough investigation has never been made. Thus, an interdisciplinary team of experts studied the remains using the most recent analytical methods in order to provide an exhaustive analysis of the remains. The bodies were submitted for osteological and archaeothanatological study, radiological investigation, AMS radiocarbon dating, chemical and textile analyses, 3D modelling, entomological as well as aDNA investigation. Here we synthesize the results of one of the most extensive multidisciplinary analyses of ancient Egyptian child mummies, adding significantly to our knowledge of such examples of ancient funerary practices.© 2020 Oras et al. This is an open access article distributed under the terms of the Creative Commons Attribution License, which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.NHM Repositor
262 Voyages Beneath the Sea: a global assessment of macro- and megafaunal biodiversity and research effort at deep-sea hydrothermal vents
For over 40 years, hydrothermal vents and the communities that thrive on them have been a source of profound discovery for deep-sea ecologists. These ecosystems are found throughout the world on active plate margins as well as other geologically active features. In addition to their ecologic interest, hydrothermal vent fields are comprised of metallic ores, sparking a nascent industry that aims to mine these metal-rich deposits for their mineral wealth. Here, we provide the first systematic assessment of macrofaunal and megafaunal biodiversity at hydrothermal vents normalized against research effort. Cruise reports from scientific expeditions as well as other literature were used to characterize the extent of exploration, determine the relative biodiversity of different biogeographic provinces, identify knowledge gaps related to the distribution of research effort, and prioritize targets for additional sampling to establish biodiversity baselines ahead of potential commercial exploitation. The Northwest Pacific, Southwest Pacific, and Southern Ocean biogeographic provinces were identified as high biodiversity using rarefaction of family-level incidence data, whereas the North East Pacific Rise, Northern East Pacific, Mid-Atlantic Ridge, and Indian Ocean provinces had medium biodiversity, and the Mid-Cayman Spreading Center was identified as a province of relatively low biodiversity. A North/South divide in the extent of biological research and the targets of hydrothermal vent mining prospects was also identified. Finally, we provide an estimate of sampling completeness for each province to inform scientific and stewardship priorities.This is an open access article under the terms of the Creative Commons Attribution License, which permits use, distribution and reproduction in any medium, provided the original work is properly cited.NHM Repositor
Mariana serpentinite mud volcanism exhumes subducted seamount materials: implications for the origin of life
The subduction of seamounts and ridge features at convergent plate boundaries plays an
important role in the deformation of the overriding plate and influences geochemical cycling
and associated biological processes. Active serpentinization of forearc mantle and serpentinite
mud volcanism on the Mariana forearc (between the trench and active volcanic arc) provides
windows on subduction processes. Here, we present (1) the first observation of an extensive
exposure of an undeformed Cretaceous seamount currently being subducted at the Mariana
Trench inner slope; (2) vertical deformation of the forearc region related to subduction of Pacific
Plate seamounts and thickened crust; (3) recovered Ocean Drilling Program and International
Ocean Discovery Program cores of serpentinite mudflows that confirm exhumation of various
Pacific Plate lithologies, including subducted reef limestone; (4) petrologic, geochemical and
paleontological data from the cores that show that Pacific Plate seamount exhumation covers
greater spatial and temporal extents; (5) the inference that microbial communities associated
with serpentinite mud volcanism may also be exhumed from the subducted plate seafloor
and/or seamounts; and (6) the implications for effects of these processes with regard to
evolution of life.Copyright 2020 The Authors. Published by the Royal Society under the terms of the
Creative Commons Attribution License http://creativecommons.org/licenses/
by/4.0/, which permits unrestricted use, provided the original author and
source are credited.NHM Repositor
Adakite-like granitoids of Songkultau: A relic of juvenile Cambrian arc in Kyrgyz Tien Shan
The early Paleozoic Terskey Suture zone, located in the southern part of the Northern Tien Shan domain in Kyrgyzstan, comprises tectonic slivers of dismembered ophiolites and associated primitive volcanics and deep-marine sediments. In the Lake Songkul area, early-middle Cambrian pillow basalts are crosscut by the Songkultau intrusion of coarse-grained gneissose quartz diorites and tonalites with geochemical characteristics typical for high-SiO2 adakites (SiO2 > 56 wt.%, Al2O3 > 15 wt.%, Na2O > 3.5 wt.% and high Sr/Y and La/Yb ratios). The Songkultau granitoids have positive initial εNd (+3.8 to +6.4) and εHf (+12.3 to +13.5) values indicating derivation from sources with MORB-like isotopic signature. Volcanic formations, surrounding the Songkultau intrusion, have geochemical affinities varying from ocean floor to island arc series. This rock assemblage is interpreted as a relic of an early-middle Cambrian primitive arc where the adakite-like granitoids were derived from partial melting of young and hot subducted oceanic crust. An age of 505 Ma, obtained for the Songkultau intrusion, shows that hot subduction under the Northern Tien Shan continued until middle Cambrian. The primitive arc complexes were obducted onto the Northern Tien Shan domain, where the Andean type continental magmatic arc developed in Cambrian and Ordovician. Formation of the Andean type arc was accompanied by uplift, erosion and deposition of coarse clastic sediments. A depositional age of ca. 470 Ma, obtained for the gravellites in the Lake Songkul area, is in agreement with the timing of deposition for lower Ordovician conglomerates elsewhere in the Northern Tien Shan, and corresponds to the main phase of the Andean type magmatism. The Songkultau adakites in association with surrounding ocean floor and island arc formations constitute a relic of a primitive Cambrian arc and represent a juvenile domain of substantial size identified so far within the predominantly crustal-derived terranes of Tien Shan. On a regional scale this primitive arc can be compared with juvenile Cambrian arcs of Kazakhstan, Gorny Altai and Mongolia.©2020 China University of Geosciences (Beijing) and Peking University. Production and hosting by Elsevier B.V. This is an open access article under the CC BY-NC-ND license (http://creativecommons.org/licenses/by-nc-nd/4.0/).NHM Repositor
Zircon-hosted apatite inclusions: A powerful tool for reconstruction of Cl contents in melts
Chlorine in the exsolved volatile phase plays an important role in complexing with metals in the extraction and concentration of metals in magmatic-hydrothermal ore deposits. Therefore, tracking the concentration and evolution of Cl in the parent melt is of particular importance in understanding how such deposits form. In theory, the incorporation of Cl into apatite could be used to track the volatile content of melts; however, low closure temperatures and the rapid diffusion of halogens in apatite make it susceptible to sub-solidus re-equilibration by later thermal events and hydrothermal fluids. This susceptibility compromises its ability to retain the primary halogen signature. However, the common occurrence of apatite as an inclusion phase in zircon crystals, together with the refractory nature of zircon, open up the possibility that such inclusions may preserve primary Clmelt compositions [1]. The Rio-Blanco-Los Bronces porphyry copper district is located in central Chile and hosts several world class porphyry copper deposits as well as barren intrusions [2]. This makes it an excellent area for an investigation of the role of Clmelt in the formation of porphyry copper deposits, as well as the effect of sub-solidus re-equilibration of Cl in apatite. For this study we analysed apatite crystals that occur both in the groundmass and as inclusions in zircons in four samples from the Los Bronces porphyry copper district using EPMA for halogen and major elements and LA-ICP-MS for trace elements. These samples include a barren intrusion unrelated to mineralisation that precedes mineralisation by around 10 Ma, and pre-, syn- and post-mineralisation porphyries. Apatite inclusions hosted in zircon crystals typically exhibit a large range in Cl concentrations (<0.5 –2.5 wt.% Cl), with all inclusion data exhibiting polymodal distributions of Cl concentrations. By contrast, groundmass apatites from all samples are characterised by uniformly low Cl concentrations (<0.5 wt.% Cl). These results are consistent with the apatite crystals in the groundmass having experienced sub-solidus re-equilibration related to the pervasive hydrothermal alteration in the district. The wide range in Cl concentrations recorded by the apatite inclusions is interpreted to reflect changing Clmelt for the duration of apatite and zircon crystallisation, perhaps linked to volatile saturation and preferential partitioning of Cl into the aqueous phase. Additionally, the apatites hosted in zircon crystals show significant inter-sample variations, evolving from low Cl concentration (<0.5 wt. % Cl) in the barren intrusion, to higher Cl concentrations (0.5 – 2.5 wt.% Cl) in the samples closely temporally associated with porphyry Cu mineralisation. These data suggest that Clmelt was significantly higher (0.05 – 0.40 wt.% Clmelt) in the melts associated with porphyry copper mineralisation compared with the precursor barren magmatism (~0.04 wt.% Clmelt) [3]. We conclude that due to the rapid diffusion of halogens in apatite in the presence of melt or hydrothermal fluid, the study of apatite inclusions hosted in zircon crystals is required to reconstruct primary melt compositions and to track the evolution of Cl concentrations in porphyry-forming magmas. This study reveals high Clmelt concentrations in the magmas related to mineralisation in the Los Bronces district, a property that would have facilitated the efficient extraction and concentration of metals. References: [1] Brugge, E. et al. (2019). Proc. 15th SGA Biennial Meeting, Vol. 2, 983-986. [2] Toro, J.C. et al. (2012). SEG Special Publication 16:105-126. [3] Li, H. and Hermann, J. (2017) Am. Mineral. 102:580-594.Abstracts and programme freely available.NHM Repositor
Ahead of the curve: three approaches to mass digitisation of vials with a focus on label data capture
There has been little research on novel approaches to digitising liquid-preserved natural history specimens stored in jars or vials. This paper discusses and analyses three different prototypes for high-throughput digitisation using cheap, readily available components. This paper has been written for other digitisation teams or curators who want to trial or improve upon these new digitisation approaches in liquid preserved collections.This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
The attached file is the published pdf.NHM Repositor
Species‐level image classification with convolutional neural network enables insect identification from habitus images
1. Changes in insect biomass, abundance, and diversity are challenging to track at
sufficient spatial, temporal, and taxonomic resolution. Camera traps can capture
habitus images of ground-dwelling insects. However, currently sampling involves
manually detecting and identifying specimens. Here, we test whether a convolutional neural network (CNN) can classify habitus images of ground beetles to
species level, and estimate how correct classification relates to body size, number
of species inside genera, and species identity.
2. We created an image database of 65,841 museum specimens comprising 361
carabid beetle species from the British Isles and fine-tuned the parameters of a
pretrained CNN from a training dataset. By summing up class confidence values
within genus, tribe, and subfamily and setting a confidence threshold, we trade-off
between classification accuracy, precision, and recall and taxonomic resolution.
3. The CNN classified 51.9% of 19,164 test images correctly to species level and
74.9% to genus level. Average classification recall on species level was 50.7%.
Applying a threshold of 0.5 increased the average classification recall to 74.6% at
the expense of taxonomic resolution. Higher top value from the output layer and
larger sized species were more often classified correctly, as were images of species in genera with few species.
4. Fine-tuning enabled us to classify images with a high mean recall for the whole
test dataset to species or higher taxonomic levels, however, with high variability.
This indicates that some species are more difficult to identify because of properties such as their body size or the number of related species.
5. Together, species-level image classification of arthropods from museum collections
and ecological monitoring can substantially increase the amount of occurrence
data that can feasibly be collected. These tools thus provide new opportunities in
understanding and predicting ecological responses to environmental change.This is an open access article under the terms of the Creative Commons Attribution License, which permits use, distribution and reproduction in any medium,
provided the original work is properly cited.
© 2019 The Authors. Ecology and Evolution published by John Wiley & Sons Ltd.
The attached file is the published pdf.NHM Repositor
Metabarcoding unsorted kick‐samples facilitates macroinvertebrate‐based biomonitoring with increased taxonomic resolution, while outperforming environmental DNA
Pereira‐da‐Conceicoa, L, Elbrecht, V, Hall, A, Briscoe, A, Barber‐James, H, Price, B. Metabarcoding unsorted kick‐samples facilitates macroinvertebrate‐based biomonitoring with increased taxonomic resolution, while outperforming environmental DNA. Environmental DNA. 2020; 00: 1– 19. https://doi.org/10.1002/edn3.116© 2020 The Authors. Environmental DNA published by John Wiley & Sons Ltd
This is an open access article under the terms of the Creative Commons Attribution License, which permits use, distribution and reproduction in any medium, provided the original work is properly cited.
The attached file is the published pdf.NHM Repositor