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    The Interpretations and Structures of Universal Quantifier GUME ‘all’ in Sibe

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    This paper discusses the syntactic properties and interpretations of the universal quantifier GUME ‘all’ in the Sibe language. GUME imparts universality and maximality to various preceding expressions, including noun phrases, pronouns, free-choice items, adverbial temporal phrases, and unconditional clauses. Employing a feature-checking approach, I propose that GUME possesses an uninterpretable universal feature [u∀], which triggers its associates to move to the specifier position of GUME. This feature-checking mechanism not only clarifies the leftness condition but also contributes to the determination of scope interpretations. Furthermore, the analysis highlights similarities as well as distinctions between GUME quantification in Sibe and DOU quantification in Mandarin Chinese, emphasizing the significance of universal quantifiers across languages

    Subject Incorporation and Derivation by the Affricative Prefix e-/o- in Ainu

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    Some Ainu verbs incorporate nouns that function as arguments. This phenomenon is known as “incorporation,” and this study focuses on subject incorporation. Sato (2012) noted that the subject is typically an element that is difficult to incorporate, but it can be included if it occupies only a background position in relation to the event as a whole. I examine the derivation by the prefix e-/o-. This has the same function as the affiliation form. I point out that intransitive verbs formed by subject incorporation have the property of unaccusative verbs and that the backgrounding of the original subject is confirmed in terms of derivation

    趣旨説明

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    Overseas Research Fellowships by the Ministry of Education in Hokkaido Imperial University

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    第3章 光合成研究の基盤となる実験技術・手法

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    State transition in cyanobacteria and Universal Stress Protein

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    シアノバクテリアの光環境応答メカニズムの一つであるステート遷移は,周辺集光装置であるフィコビリソームから光化学系Ⅰと光化学系Ⅱへのエネルギー分配を,秒から分単位で調節する.陸上植物や緑藻の周辺集光装置LHCIIからのエネルギー分配が調節される同様のメカニズムの解明が比較的進んでいるのに対して,シアノバクテリアのステート遷移には未解明の点が多い.本稿では,ステート遷移のメカニズムとそれに関わる因子を概説した後,ごく最近になってステート遷移にかかわることが示されたUniversal Stress Proteinの一種であるSlr0244について紹介する.最後に,Universal Stress Proteinによるシグナル伝達の様々な可能性について考察する.State transitions, one of the light-environment response mechanisms of cyanobacteria, regulate the distribution of energy from the phycobilisome, a peripheral light-harvesting apparatus, to photosystem I and photosystem II on a second-to-minute scale. While the similar regulatory mechanism of energy distribution from the peripheral antenna LHCII to photosystems in land plants and green algae is relatively well understood, the state transition in cyanobacteria is still largely unexplored. In this paper, after outlining the mechanism of state transitions and the factors involved, we introduce Slr0244, a type of Universal Stress Protein that has recently been shown to be involved in state transitions. Finally, the various possibilities of the signal transduction by the Universal Stress Protein is discussed

    Far-red light driven photosynthesis acquired in the evolutionary process of an Antarctic alga, Prasiola crispa : Its mechanism and evolutionary lineage

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    ナンキョクカワノリは南極の陸上環境に大きなコロニーを形成することで知られている.最近の研究から,ナンキョクカワノリは可視光が届きにくいコロニーの内部環境で可視光より長波長の遠赤色光を吸収する光捕集アンテナ蛋白質(Pc-frLHC)を発現し,アップヒル型のエネルギー移動により可視光より低いエネルギーの光で酸素発生型の光合成を行うことが明らかになった.構造解析の結果,PcfrLHCは11量体のリング構造をしており,長波長吸収型の3量体クロロフィルを結合することが分かった.系統解析の結果,Pc-frLHCはトレブクシア藻綱の中で,緑藻の多くが持っている可視光吸収型LHCIのひとつからアミノ酸配列の変化を経て進化したことが示唆された.Prasiola crispa is known to form large colonies in terrestrial environments of Antarctica. In our recent studies, it has been revealed that P. crispa expresses a far-red light-harvesting chlorophyll (Chl)-binding protein complex (Pc-frLHC) in the internal environment of colony where infrared light is predominant, and performs photosynthesis using far-red light, which is lower energy than visible light, through uphill energy transfer. Structural analysis has revealed that Pc-frLHC has a homo 11-mer ring like structure and binds long-wavelength absorbing trimeric chlorophylls. Phylogenetic analysis suggests that Pc-frLHC evolved in Trebouxiophyceae, through changes in amino acid sequence from one of the visible light absorbing LHCI that is a common LHC in green algae

    ウェアラブル機器を用いた概日リズムと睡眠覚醒リズムの評価・調整法の確立とその臨床応用 [全文の要約]

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    この博士論文全文の閲覧方法については、以下のサイトをご参照ください。https://www.lib.hokudai.ac.jp/dissertations/copy-guides

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