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Une vie moderne, Zhou Daxin
Daxin Zhou, Naour Françoise. Une vie moderne, Zhou Daxin. In: Perspectives chinoises, n°63, 2001. pp. 72-78
Erymus gilvus Zhou & Zhou, sp. n.
1. Erymus gilvus Zhou & Zhou, sp. n. (Fig. 1 A; 3 A–F) Type material. Holotype: male, CHINA: Hainan: Ledong co.: Jianfeng Ling (E 108.8792, N 18.7211), 808 m, 27.III. 2008, Li Liang collector (IZ-CAS); paratypes: CHINA: Hainan: Ledong co.: Jianfeng Ling, one male, 800 m, 16.VII. 2004, Wu Jie & Chen Yongjie collectors; one female, same data as above, except 24.VII. 2004; Guangxi: Xingan co.: Miaoer Shan (E 110.4943, N 25.8810), one male, one female, 06.VII. 1985, Cui Yunqi collector; one female, same data as above, except 30.VI. 1985; one female, same data as above, except 11.VII. 1985; Guangdong: Zhaoqing city: Dihu Shan (E 112.5533, N 23.1714), one male, 150 m, 21.VIII. 2004, Wu Jie & Chen Yongjie collectors (IZ-CAS). Description. Measurements. BL= 3.72 mm, FL= 1.98 mm, HL= 0.57 mm, HW= 0.48 mm, PL= 0. 65 mm, PW= 0.41 mm, EL= 0.67 mm, EW= 0.58 mm. Body nearly cylindrical, small-sized. Body light brown, except head dark brown; antennae, legs and last abdominal segment yellowish. Head (Fig. 1 A). Subrectangular (HL to HW ratio 1.2), tempora slightly arcuate, occasionally dilated posteriorly, posterior angles rounded. Dorsal integument extensively bearing micropunctures, without any microsculpture; median longitudinal region (of the width ≈ 0.15 mm) smooth, without punctures, other surface (posterior half of head and temporal region) with deep, sparsely-scattered, medium sized punctures, distance between punctures about the diameter of 5–6 punctures; each deflexed portion of tempora with two rows of larger and deeper punctures. Each side of cranium with anterolateral puncture near antennal insertion, midlateral puncture adjacent to dorsal margin of eye. Frontal furrows deep and long, fused with two paired tiny punctures, reaching line connecting mid-length of each eye. Anteocular furrows indistinct. Eyes large, distinctly longer than half length of tempora (eye: tempora = 0.19: 0.28 mm), distinctly protruding laterad. Epistoma relatively broad, not narrowed forwards. Distance between antennal insertions 0.13 mm, longer than distance from antenna to eye (0.08 mm). Antennae. Scape stout, thickened apically, subequal to length of three subsequent antennomeres combined, 0.20 mm; antennomere II dilated subapically, 0.10 mm; III spherical and small, much shorter than II, 0.04 mm; IV and V obviously wider than III, subequal in length of 0.05 mm; last antennomere medium in length, 0.13 mm, subequal to length of two preceding antennomeres combined. Neck. Medium width (0.17 mm), slightly wider than 1 / 3 of head width. Pronotum (Fig. 1 A). Subrectangular, distinctly elongate (PL to PW ratio 1.6), longer but narrower than head. Widened anteriad, lateral margins sinuate, anterior angles rounded, posterior angles broadly rounded, with widest at anterior 1 / 3 and narrowest at posterior 1 / 3. Dorsal surface extensively bearing micropunctures, without any microsculpture. Each side bearing a pair of admedian row of 5 larger punctures, a pair of lateral row of 4 punctures, and also with few irregular punctures distributing near anterior margin. Mesoscutellum. Glossy, without any microsculpture, and with a pair of small punctures at apical 1 / 3. Elytra (Fig. 1 A). Subrectangular, elongate (EL to EW ratio 1.2), the same length of pronotum, but distinctly wider. Humeri well-developed, lateral margins distinctly dilated outwards, hind margin distinctly rounded. Integument glossy, flattened, without microsculpture, each side with 2–3 rows of small punctures along median suture, a row along midwidth, and also with 1–2 rows of tiny punctures on deflexed portion. Abdomen. Cylindrical, broadest at segment VI. Tergites III–VII glossy, each segment covered with shallow transverse microstriae, also with small scattered punctures, interspaces between punctures near 2–3 punctures’ diameter, and punctures denser on tergites VI and VII. Each tergite with a shallow basal impression near anterior margin; surface between two basal transverse carinae glossy, covered with polygonal reticulum, and without punctures. All abdominal sternites glossy, with microstriae and punctures as those on tergites. Male (Fig. 3 A–F). Abdominal segment VIII entirely covered with setiferous punctures, posterior margin of both tergite and sternite subtruncated (Fig. 3 A, B). Tergite IX symmetrical, connected mediobasally. Sternite IX asymmetrical, lateral margins arcuate, with linear-shaped base and rounded apex (Fig. 3 C). Tergite X symmetrical, subrhomboidal, with elongate base and rounded apex (Fig. 3 D). Aedeagus elliptical, anterior portion missing, apical margin slightly emarginated (Fig. 3 E–F), bulbus very small, 0.22 mm long. Parameres symmetrical and quite elongate, 0.27 mm, with paired inner process at basal 1 / 3. Internal sac entirely membranous and variably shaped, without any visible sclerotized structures. Distribution. China (Guangxi, Guangdong, Hainan). Etymology. The specific epithet is derived from the Latin words “ gilv ” (light yellow), and refers to the body coloration. Remarks. Based on the long parameres, the new species is similar to E. gracilis (Fauvel, 1895). But it can be distinguished from the latter by paler body color, more sparsely-scattered punctures on head, a process on each paramere, and by the distinctly narrower tergite X.Published as part of Zhou, Yu-Lingzi & Zhou, Hong-Zhang, 2014, Taxonomy of the genus Erymus Bordoni (Coleoptera: Staphylinidae, Staphylininae, Xantholinini) in China with descriptions of two new species, pp. 521-536 in Zootaxa 3814 (4) on pages 523-527, DOI: 10.11646/zootaxa.3814.4.5, http://zenodo.org/record/22488
Medhiama liupanshanensis Zhou & Zhou, sp. n.
2. <i>Medhiama liupanshanensis</i> Zhou & Zhou, sp. n. <p>(Fig. 4 A–H)</p> <p> <b>Type material.</b> Holotype: male, <b>CHINA</b>: <b>Ningxia</b>: <b>JingYuan co.:</b> Liupan Shan: Shanpo Linchang, 2200 m, 06.VII.2008, Zhou Haisheng & Zhao Zongyi collected (IZ-CAS); Paratypes: 1 female, same data as holotype; Xixia Linchang, 1 male, 2100 m, 26.VI.2008, Zhou Haisheng & Zhao Zongyi collected; Erlonghe Linchang, 1 female, 2050 m, 22.VI.2008, Zhou Haisheng & Zhao Zongyi collected (IZ-CAS).</p> <p> <b>Description. Measurement.</b> BL= 5.89 mm, FL= 3.23 mm, HL = 1.08 mm, HW= 0.80 mm, PL= 1.10 mm, PW= 0.70 mm, EL= 0.95 mm, EW= 0.95 mm.</p> <p>Body nearly cylindrical and medium sized. Entirely brown, except legs, apical 1/2 of last antennomere, last segment of maxillary and labial palpi lighter in color.</p> <p> <b>Head</b>. Oblong (HL to HW ratio 1.36), tempora substraight, not widened, posterior angles rounded. Dorsal integument entirely bearing microsculpture composed of shallow polygonal reticulum, and extensively distributed relatively small punctures, puncture intervals not wide, being equal about puncture diameter. Median longitudinal region impunctate, width equals to 4–5 puncture diameters; frontal region with shallow microstriae, and 3 pairs of tiny punctures. Each side of cranium without anterolateral puncture, with midlateral puncture far from dorsal margin of eye (5–6 puncture diameters to eye), temporal puncture rather near lateral margin (at lateral 1/6) and occipital puncture rather near posterior margin (at posterior 1/6). Frontal furrows deep and long, convergent backwards; anteocular furrows indistinct, scarcely observable. Eye quite small and rather flat, diameter not over 1/5 of temporal length (eye: tempora = 0.13: 0.82 mm). Epistoma relatively wide, subrectangular and flat, with a pair of tiny punctures. Distance between antennal insertions 0.24 mm, longer than distance from antenna to eye (0.19 mm).</p> <p> <b>Antennae</b>. Scape stout, thickened apically, much longer than three subsequent antennomeres combined, 0.44 mm; antennomere II elongate, 0.11 mm; III elongate, slightly shorter than II, 0.10 mm; IV and V subequal in length, 0.080 mm; last antennomere of medium length, 0.19 mm, subequal to two preceding antennomeres combined.</p> <p> <b>Mouthparts</b>. Labrum bilobed, with a median longitudinal groove. Maxillary palpus elongate, segment III longest, last segment slender and obconical and shorter than the penultimate. Labial palpus distinctly slender, last segment longest.</p> <p> <b>Neck</b>. Medium width (0.29 mm), nearly 1/3 of head width, with a transverse substraight ridge on anterior 1/3.</p> <p> <b>Pronotum</b>. Relatively shorter (PL to PW ratio 1.57), of same length as head, but narrower. Widest at anterior 1/3, narrowest at posterior 1/3. Anterior angles widely rounded and slightly protruding, lateral margins sinuate from middle, posterior angles rounded. Integument bearing shallow microstriae, and a pair of admedian row of 12–14 punctures, smaller than those on head. Areas outside admedian rows with additional, irregular, sparsely set punctures.</p> <p> <b>Mesoscutellum.</b> Shiny, extensively bearing polygonal reticulum and with a pair of small punctures on apical 1/4.</p> <p> <b>Elytra</b>. Subquadrate (EL to EW ratio 1.0), shorter but distinctly wider than pronotum. Humeri well developed, lateral margins widened posteriorly, hind margin rounded. Integument slightly wrinkled, without microsculpture; each side symmetrically with regular rows of punctures, interspaces between rows 1–2 puncture diameters; deflexed portion of each elytron with 3–4 rows of punctures.</p> <p> <b>Legs</b>. First four segments of protarsi stout, not dilated, those of mesotarsi relatively slender; each last segment as long as the II–IV combined. Protibia with apical ctenidium and subapical ctenidia, meso- and metatibia only with apical ctenidium.</p> <p> <b>Abdomen.</b> Cylindrical, broadest at segment VII. Tergites III–VII shiny, surface entirely covered with a distinct mixture of extensive microstriae and polygonal reticulum; punctures small, sparsely scattered, interspace between them 3–4 puncture diameters, but much denser on tergite VI and VII. Each tergite with median longitudinal impunctate region, width about 3–4 puncture diameters, without distinct basal impression near anterior margin. Surface between two basal transverse carinae of tergites III–VII bearing distinct polygonal reticulum. All abdominal sternites shiny, with microstriae and setiferous punctures as those on tergites.</p> <p> <b>Male</b> (Fig. 4 A–F). Abdominal segment VIII entirely covered with setiferous punctures, posterior margins of tergite VIII and sternite VIII both slightly emarginated (Fig.4 A, B). Tergite IX symmetrical, connected mediobasally. Sternite IX, with long linear-shaped base and rounded in right margin (Fig.4 D). Tergite X symmetrical and broadest at anterior 1/4, anterior 1/4 sharply narrowed and with obtusely rounded apex (Fig.4 C). Aedeagus elliptical and large (Fig. 4 E, F), basal bulb 1.16 mm long. Parameres symmetrical and simple, 0.34 mm and slightly shorter than 1/3 of basal bulb length. Internal sac broadly bag-like, gradually widened and with cellshaped structure in median portion (Fig. 4 E).</p> <p> <b>Female</b> (Fig. 4 G, H). Sternite VIII not distinctly oblong, and posterior margin sharply protruding posteriorly (Fig. 4 G). Genital segment with a large pair of subtriangular supplementary sclerites, and a broad sternite with not distinctly protruding anterior margin (Fig. 4 H).</p> <p> <b>Distribution.</b> China (Ningxia).</p> <p> <b>Etymology</b>. The specific epithet is the Latinized adjective derived from the Chinese name (Pin-yin) of the type locality: Liupan Shan.</p> <p> <b>Remarks.</b> This new species can be easily distinguished from its congeners by the following character combination: sparsely punctate head, emarginated posterior margin of male sternite VIII, special form of sternite IX, tergite X, unique shape of inner sac and a broad sternite with slightly protruding anterior margin in female genital segment.</p>Published as part of <i>Zhou, Yu-Lingzi & Zhou, Hong-Zhang, 2012, Taxonomy of the genus Medhiama Bordoni, 2002 (Coleoptera: Staphylinidae, Staphylininae, Xantholinini) with descriptions of three new species, pp. 169-191 in Zootaxa 3478</i> on pages 175-178, DOI: <a href="http://zenodo.org/record/282239">10.5281/zenodo.282239</a>
Medhiama xiaolongmenensis Zhou & Zhou, sp. n.
1. Medhiama xiaolongmenensis Zhou & Zhou, sp. n. (Fig. 3 A–H) Type material. Holotype: male, CHINA: Beijing: Mentougou District: Xiaolongmen, 1600 m, 03.VII. 1998, Zhou Haisheng collected (IZ-CAS); Paratypes: 2 males, 11 females, same data as holotype; Sichuan: Baoxing co.: Longdong town, Ganyang Gou, 1 male, 5 females, 1796 m, 12.VIII. 2003, Wu Jie collected; same locality, 1 female, 1795 m, 12.VIII. 2003, Yu Xiongdong collected; Puji Gou, 1 female, 2180 m, 14.VIII. 2003, Yu Xiaodong & Peng Xuewei collected (IZ-CAS). Description. Measurement. BL= 6.75 mm, FL= 3.52 mm, HL= 1.16 mm, HW= 0.86 mm, PL= 1.20 mm, PW= 0.74 mm, EL= 1.05 mm, EW= 1.05 mm. Body nearly cylindrical and medium sized. Entirely dark brown. Legs brown. Antennae dark brown, apical half of last antennomere lighter in color. Maxillary and labial palpi dark brown. Head. Oblong (HL to HW ratio 1.36), tempora widened posteriorly, posterior angles widely rounded. Dorsal integument entirely bearing microsculpture composed of distinct polygonal reticulum, and extensively distributed deep and medium sized punctures, puncture intervals rather narrow, about 1 / 2 puncture diameter. Median longitudinal region impunctate, width equals to 2–3 puncture diameters; frontal region with distinct transverse microstriae, and 3 pairs of tiny punctures. Each side of cranium without anterolateral puncture, but with midlateral puncture far from dorsal margin of eye (5–6 puncture diameters to eye), temporal puncture rather near lateral margin (at lateral 1 / 6) and occipital puncture rather near posterior margin (at posterior 1 / 6). Frontal furrows deep and long, convergent backwards; anteocular furrows indistinct, scarcely observable. Eye of small size and flat, diameter nearly 1 / 4 of temporal length (eye: tempora = 0.19: 0.76 mm). Epistoma relatively wide, subrectangular and flat, with a pair of tiny punctures. Distance between antennal insertions 0.23 mm, subequal to distance from antenna to eye (0.21 mm). Antennae. Scape stout, thickened apically, much longer than three subsequent antennomeres combined, 0.44 mm; antennomere II slightly elongate or subglobular, 0.10 mm; III elongate, slightly longer than II, 0.11 mm; IV and V subequal in length, 0.09 mm; last antennomere of medium length, 0.19 mm, subequal to two preceding antennomeres combined. Mouthparts. Labrum bilobed and relatively long, with a median longitudinal groove. Maxillary palpus elongate, segment III longest, last segment slender and obconical and shorter than the penultimate. Labial palpus distinctly slender, last segment longest. Neck. Medium width (0.30 mm), nearly 1 / 3 of head width, with a transverse substraight ridge on anterior 1 / 3. Pronotum. Distinctly elongate (PL to PW ratio 1.62), of same length as head, but narrower. Widest at anterior 1 / 3, narrowest at posterior 1 / 3. Anterior angles widely rounded, not protruding, lateral margins sinuate from middle, posterior angles rounded. Integument bearing distinct transverse microstriae, and a pair of admedian row of 12–14 punctures, smaller than those on head. Areas outside admedian rows with additional, irregular, sparsely set punctures. Mesoscutellum. Shiny, extensively bearing polygonal reticulum and with a pair of small punctures on apical 1 / 4. Elytra. Subquadrate (EL to EW ratio 1.0), shorter but distinctly wider than pronotum. Humeri well developed, lateral margins widened posteriorly, hind margin rounded. Integument slightly wrinkled, without microsculpture; each side symmetrically with regular rows of punctures, interspaces between rows 1–2 puncture diameters; deflexed portion of each elytron with 3–4 rows of punctures. Legs. First four segments stout but not dilated in protarsi and metatarsi, whereas relatively slender in mesotarsi; each last segment as long as the II–IV combined. Protibia with apical ctenidium and subapical ctenidia, meso- and metatibia only with apical ctenidium. Abdomen. Cylindrical, broadest at segment VII. Tergites III–VII shiny, surface entirely covered with observable transverse microstriae, also with denser polygonal reticulum near anterior margin; punctures small, sparsely scattered, interspace between them approximately 3–4 puncture diameters, but much denser on tergites VII and VIII. Each tergite without distinct basal impression near anterior margin. Surface between two basal transverse carinae of tergites III–VII bearing distinct polygonal reticulum. All abdominal sternites shiny, with microstriae and setiferous punctures as those on tergites. Male (Fig. 3 A–F). Abdominal segment VIII entirely covered with setiferous punctures, posterior margin of tergite VIII subtruncated, that of sternite VIII sharply protruding posteriorly (Fig. 3 A, B). Tergite IX symmetrical, connected mediobasally. Sternite IX, with long linear-shaped base and sharp apex (Fig. 3 D). Tergite X symmetrical, broadest at anterior 1 / 3, lateral margins quite expanded outwards (Fig. 3 C). Aedeagus of irregular shape and small in size (Fig. 3 E, F), basal bulb 0.48 mm long. Parameres asymmetrical and foot-shaped, right one obviously larger than left one in dorsal view (Fig. 3 F), base distinctly darker; large sized, 0.40 mm long and subequal to 4 / 5 of basal bulb length. Internal sac located outside basal bulb, brown and soft, without distinct spines (Fig. 3 E). Female (Fig. 3 G, H). Sternite VIII not distinctly oblong, and posterior margin protruding medially (Fig. 3 G). Genital segment with pair of subtriangular supplementary sclerites and broad sternite with distinctly protruding anterior margin (Fig. 3 H). Distribution. China (Beijing, Sichuan). Etymology. The specific epithet is derived from the Chinese name (Pin-yin) of the type locality, Xiaolongmen. Remarks. This new species could be distinguished from its congeners by the following character combination: in male by widened anterior angles of pronotum, protruding posterior margin of male sternite VIII, distinctive shape of sternite IX, tergite X, asymmetrical parameres, unique form of inner sac, and in female by broad sternite of genital segment with a distinctly protruding anterior margin. This is the second species of the genus with asymmetrical parameres (the other being M. puetzi Bordoni, 2003), and the two could be distinguished from each other by different shapes of sternite IX, tergite X and aedeagi. The new species also shared similar foot-shaped parameres with M. schawalleri Bordoni, but the pronotum widened distinctly near anterior angles, punctures on abdomen more densely distributed, and tergite X more outwardly expanded.Published as part of Zhou, Yu-Lingzi & Zhou, Hong-Zhang, 2012, Taxonomy of the genus Medhiama Bordoni, 2002 (Coleoptera: Staphylinidae, Staphylininae, Xantholinini) with descriptions of three new species, pp. 169-191 in Zootaxa 3478 on pages 173-175, DOI: 10.5281/zenodo.28223
Medhiama xiaolongmenensis Zhou & Zhou, sp. n.
1. <i>Medhiama xiaolongmenensis</i> Zhou & Zhou, sp. n. <p>(Fig. 3 A–H)</p> <p> <b>Type material.</b> Holotype: male, <b>CHINA: Beijing:</b> Mentougou District: Xiaolongmen, 1600 m, 03.VII.1998, Zhou Haisheng collected (IZ-CAS); Paratypes: 2 males, 11 females, same data as holotype; <b>Sichuan: Baoxing co.</b>: Longdong town, Ganyang Gou, 1 male, 5 females, 1796 m, 12.VIII.2003, Wu Jie collected; same locality, 1 female, 1795 m, 12.VIII.2003, Yu Xiongdong collected; Puji Gou, 1 female, 2180 m, 14.VIII.2003, Yu Xiaodong & Peng Xuewei collected (IZ-CAS).</p> <p> <b>Description. Measurement.</b> BL= 6.75 mm, FL= 3.52 mm, HL= 1.16 mm, HW= 0.86 mm, PL= 1.20 mm, PW= 0.74 mm, EL= 1.05 mm, EW= 1.05 mm.</p> <p>Body nearly cylindrical and medium sized. Entirely dark brown. Legs brown. Antennae dark brown, apical half of last antennomere lighter in color. Maxillary and labial palpi dark brown.</p> <p> <b>Head</b>. Oblong (HL to HW ratio 1.36), tempora widened posteriorly, posterior angles widely rounded. Dorsal integument entirely bearing microsculpture composed of distinct polygonal reticulum, and extensively distributed deep and medium sized punctures, puncture intervals rather narrow, about 1/2 puncture diameter. Median longitudinal region impunctate, width equals to 2–3 puncture diameters; frontal region with distinct transverse microstriae, and 3 pairs of tiny punctures. Each side of cranium without anterolateral puncture, but with midlateral puncture far from dorsal margin of eye (5–6 puncture diameters to eye), temporal puncture rather near lateral margin (at lateral 1/6) and occipital puncture rather near posterior margin (at posterior 1/6). Frontal furrows deep and long, convergent backwards; anteocular furrows indistinct, scarcely observable. Eye of small size and flat, diameter nearly 1/4 of temporal length (eye: tempora = 0.19: 0.76 mm). Epistoma relatively wide, subrectangular and flat, with a pair of tiny punctures. Distance between antennal insertions 0.23 mm, subequal to distance from antenna to eye (0.21 mm).</p> <p> <b>Antennae</b>. Scape stout, thickened apically, much longer than three subsequent antennomeres combined, 0.44 mm; antennomere II slightly elongate or subglobular, 0.10 mm; III elongate, slightly longer than II, 0.11 mm; IV and V subequal in length, 0.09 mm; last antennomere of medium length, 0.19 mm, subequal to two preceding antennomeres combined.</p> <p> <b>Mouthparts</b>. Labrum bilobed and relatively long, with a median longitudinal groove. Maxillary palpus elongate, segment III longest, last segment slender and obconical and shorter than the penultimate. Labial palpus distinctly slender, last segment longest.</p> <p> <b>Neck</b>. Medium width (0.30 mm), nearly 1/3 of head width, with a transverse substraight ridge on anterior 1/3.</p> <p> <b>Pronotum</b>. Distinctly elongate (PL to PW ratio 1.62), of same length as head, but narrower. Widest at anterior 1/3, narrowest at posterior 1/3. Anterior angles widely rounded, not protruding, lateral margins sinuate from middle, posterior angles rounded. Integument bearing distinct transverse microstriae, and a pair of admedian row of 12–14 punctures, smaller than those on head. Areas outside admedian rows with additional, irregular, sparsely set punctures.</p> <p> <b>Mesoscutellum.</b> Shiny, extensively bearing polygonal reticulum and with a pair of small punctures on apical 1/4.</p> <p> <b>Elytra</b>. Subquadrate (EL to EW ratio 1.0), shorter but distinctly wider than pronotum. Humeri well developed, lateral margins widened posteriorly, hind margin rounded. Integument slightly wrinkled, without microsculpture; each side symmetrically with regular rows of punctures, interspaces between rows 1–2 puncture diameters; deflexed portion of each elytron with 3–4 rows of punctures.</p> <p> <b>Legs</b>. First four segments stout but not dilated in protarsi and metatarsi, whereas relatively slender in mesotarsi; each last segment as long as the II–IV combined. Protibia with apical ctenidium and subapical ctenidia, meso- and metatibia only with apical ctenidium.</p> <p> <b>Abdomen.</b> Cylindrical, broadest at segment VII. Tergites III–VII shiny, surface entirely covered with observable transverse microstriae, also with denser polygonal reticulum near anterior margin; punctures small, sparsely scattered, interspace between them approximately 3–4 puncture diameters, but much denser on tergites VII and VIII. Each tergite without distinct basal impression near anterior margin. Surface between two basal transverse carinae of tergites III–VII bearing distinct polygonal reticulum. All abdominal sternites shiny, with microstriae and setiferous punctures as those on tergites.</p> <p> <b>Male</b> (Fig. 3 A–F). Abdominal segment VIII entirely covered with setiferous punctures, posterior margin of tergite VIII subtruncated, that of sternite VIII sharply protruding posteriorly (Fig. 3 A, B). Tergite IX symmetrical, connected mediobasally. Sternite IX, with long linear-shaped base and sharp apex (Fig. 3 D). Tergite X symmetrical, broadest at anterior 1/3, lateral margins quite expanded outwards (Fig. 3 C). Aedeagus of irregular shape and small in size (Fig. 3 E, F), basal bulb 0.48 mm long. Parameres asymmetrical and foot-shaped, right one obviously larger than left one in dorsal view (Fig. 3 F), base distinctly darker; large sized, 0.40 mm long and subequal to 4/5 of basal bulb length. Internal sac located outside basal bulb, brown and soft, without distinct spines (Fig. 3 E).</p> <p> <b>Female</b> (Fig. 3 G, H). Sternite VIII not distinctly oblong, and posterior margin protruding medially (Fig. 3 G). Genital segment with pair of subtriangular supplementary sclerites and broad sternite with distinctly protruding anterior margin (Fig. 3 H).</p> <p> <b>Distribution.</b> China (Beijing, Sichuan).</p> <p> <b>Etymology</b>. The specific epithet is derived from the Chinese name (Pin-yin) of the type locality, Xiaolongmen.</p> <p> <b>Remarks.</b> This new species could be distinguished from its congeners by the following character combination: in male by widened anterior angles of pronotum, protruding posterior margin of male sternite VIII, distinctive shape of sternite IX, tergite X, asymmetrical parameres, unique form of inner sac, and in female by broad sternite of genital segment with a distinctly protruding anterior margin. This is the second species of the genus with asymmetrical parameres (the other being <i>M. puetzi</i> Bordoni, 2003), and the two could be distinguished from each other by different shapes of sternite IX, tergite X and aedeagi. The new species also shared similar foot-shaped parameres with <i>M. schawalleri</i> Bordoni, but the pronotum widened distinctly near anterior angles, punctures on abdomen more densely distributed, and tergite X more outwardly expanded.</p>Published as part of <i>Zhou, Yu-Lingzi & Zhou, Hong-Zhang, 2012, Taxonomy of the genus Medhiama Bordoni, 2002 (Coleoptera: Staphylinidae, Staphylininae, Xantholinini) with descriptions of three new species, pp. 169-191 in Zootaxa 3478</i> on pages 173-175, DOI: <a href="http://zenodo.org/record/282239">10.5281/zenodo.282239</a>
Gilliesia pulchra Zhou 2004
<i>Gilliesia pulchra</i> Zhou, 2004 <p> <b>Material examined:</b> 2 male imagoes and 2 female imagoes (reared from nymphs), 2 female subimagoes, 21 nymphs, Bang-Gun-Jian Mountain, De-Hong City, Yunnan Province, China, 97°50′52″E, 24°23′13″N, 1800 m alt., 28-VI-2022, leg. Xuhongyi Zheng, Pengxu Mu; 2 male imagoes and 1female imago, Nan-Zhu forestry center, Chi-Shui City, Guizhou Province, China, 105.42°E, 28.34°N, 9-VI-1995, leg. Changhai Sun, Beixin Wang.</p> <p>Holotype: male imago, Jing-Fu-Shan Mountain, Huang-Cao-Ping, Chongqing municipality, China, 107.10°E, 29.00°N, 1200 m alt., 3-IX-2000, leg. Chuangren Li, Changfa Zhou.</p>Published as part of <i>Gong, De-Wen & Zhou, Chang-Fa, 2023, The imaginal and nymphal morphology of Gilliesia pulchra Zhou, 2004 and its contribution to understanding generic phylogeny (Ephemeroptera: Leptophlebiidae), pp. 531-540 in Zootaxa 5263 (4)</i> on page 532, DOI: 10.11646/zootaxa.5263.4.4, <a href="http://zenodo.org/record/7835942">http://zenodo.org/record/7835942</a>
Xanthophius unicidentatus Zhou & Zhou, 2013, sp. n.
2. <i>Xanthophius unicidentatus</i> sp. n. <p>(Fig. 5 A–H; Fig. 6 A–G)</p> <p> <b>Type material.</b> Holotype: male, <b>CHINA</b>: <b>Yunnan</b>: Mengla co.: Longmen: Pinghe village, 1014 m, 07.X.2010, Zhou Yulingzi collected (IZ-CAS); Paratypes: 1 female, 2 male, same locality and collector as holotype, 945 m, 06.X.2010; Jingdong Yizu Autonomous Co.: Wenlong village, 1 female, 1268 m, 18.VII.2010, Liu Wangang collected; <b>Zhejiang:</b> Shunxi Co.: Qingliangfeng Reserve Zone, 1 female, 400 m, 09.VIII.2008, Zhou Yulingzi; <b>Guangxi:</b> Daxin Co.: Fulong: 1 male, 220 m, 20.IV.1998, Zhou Haisheng collected; Longzhou Co.: Nonggang Reserve Zone, 1 female, 330 m, 15.VI. 2000, Chen Jun collected; <b>Hainan:</b> Qiongzhong Li and Miao Autonomous Co., Limuling, 1 male, 600 m, 15.V.2007, Zhao Zongyi collected (IZ-CAS).</p> <p> <b>Description.</b></p> <p> <b>Measurement</b> (N=3). BL= 3.52–4.27 mm, FL= 2.10–2.37 mm, HL= 0.63–0.70 mm, HW= 0.50–0.60 mm, PL= 0.71–0.77 mm, PW= 0.45–0.53 mm, EL= 0.70–0.80 mm, EW= 0.65–0.70 mm.</p> <p>Body nearly cylindrical and of small size. Head distinctly dark brown; elytra brown with paler apical 1/5; pronotum, abdomen, legs and labial and maxillary palpi yellowish brown; antennae brown except for the apical 1/2 of last segment yellowish.</p> <p> <b>Head</b> (Fig. 6 A). Subrectangular (HL to HW ratio 1.25), tempora not distinctly widened posteriorly, posterior angles narrowly rounded. Dorsal integument without microsculpture; but with numerous medium-sized punctures, puncture intervals equal to 2–3 punctures’ diameter. Median longitudinal region impunctate; deflexed portion of tempora without any punctures. Each side of cranium with a puncture on epistoma, 3–4 punctures situated in frontal furrow, an anterolateral puncture near antennal insertion, a midlateral puncture about a distance of 2 punctures’ diameter from dorsal margin of eye, temporal puncture and occipital puncture not apparent. Frontal furrows long, but not longer than eye’ length; anteocular furrows distinct and short, with 3–4 medium sized punctures. Eye medium-sized, equal to the length of tempora (eye: tempora = 0.20: 0.40 mm). Epistoma relatively broad, slightly narrowed apically. Distance between antennal insertions (0.17 mm) being larger than that from antenna to eye (0.08 mm).</p> <p> <b>Antennae.</b> Scape stout, thickened apically, much longer than three subsequent antennomeres combined, 0.25 mm long; antennomere II elongate, 0.08 mm long; III slightly elongate, shorter than II, 0.06 mm long; IV and V subequal in length and 0.04 mm long; last antennomere relatively long, 0.15 mm, equal to length of three preceding antennomeres combined.</p> <p> <b>Neck.</b> Wide (0.20 mm), narrower than half the width of head.</p> <p> <b>Pronotum</b> (Fig. 6 B). Subrectangular, elongate (PL to PW ratio 1.55), longer and slightly narrower than head. Anterior angle well-defined and protruding laterally, lateral margins sinuate, widest at anterior 1/3 and narrowest at posterior 1/3, posterior angles broadly rounded. Dorsal integument without microsculpture, but bearing a pair of admedian rows of 9–10 punctures, a pair of lateral rows of 8–9 punctures, and also with few irregular punctures near anterior and lateral margins of pronotum.</p> <p> <b>Elytra</b> (Fig. 6 C). Subrectangular (EL to EW ratio 1.08), slightly shorter and wider than pronotum. Humeri well developed, lateral margins divergent posteriorly, hind margin rounded. Dorsal integument bearing 4–5 rows of punctures on each side, interspaces between rows of ca. two punctures’ diameter; deflexed portion of each elytron with 2–3 rows of denser punctures.</p> <p> <b>Legs.</b> First four segments of protarsi stout but not dilated, and those of meso- and metatarsi not slender. Last segment slightly shorter than the length of II–IV combined, respectively.</p> <p> <b>Abdomen.</b> Cylindrical, broadest at segment VII. Tergites III–VII slightly shiny, each segment without any microsculpture, and also distributed with small, sparse punctures, interspaces between punctures about 4–5 punctures’ diameter. Each tergite without a basal impression near anterior margin. All abdominal sternites shiny, with microstriae and punctures as those on tergites.</p> <p> <b>Male</b> (Fig. 5 A–G; Fig. 6 D–F). Abdominal segment VIII entirely covered with punctures, posterior margin of tergite slightly rounded, and that of sternite subtruncate (Fig. 5 A, B). Tergite IX symmetrical, connected mediobasally. Sternite IX asymmetrical, with subparalleled lateral margins, very narrow base, and rounded apex (Fig. 5 D). Tergite X symmetrical, broadest at apical 1/4, sharply narrowed at apical 1/5, with narrow and subtruncate base, and concaved posterior margin (Fig. 5 C). Aedeagus small (Fig. 5 E; Fig. 6 D–F), basal bulb subspherical, 0.42 mm long. Parameres symmetrical, not attached to basal bulb, distinctly elongate, 0.30 mm long, and extended to dorsal side (Fig. 5 F), with a small denticle in inner side. Internal sac as in Fig. 5 G, bearing numerous, dark brown, hard processes on apical and middle parts, basal part light brown.</p> <p> <b>Distribution.</b> China (Yunnan, Guangxi, Zhejiang, Hainan).</p> <p> <b>Etymology.</b> The specific epithet is derived from the combination of Latin words “ <i>unicus</i> ” (single) and “ <i>dentatus</i> ” (with denticle) and refers to the single denticle in the inner side of each paramere.</p> <p> <b>Remarks.</b> This new species can be distinguished from its congeners by the following characters: elytra brown with paler apical 1/5, tempora not distinctly widened posteriorly, anteocular furrows short, paramere with a small denticle in inner side, and internal sac as illustrated (Fig. 5 G), bearing numerous, dark brown, hard processes on apical and middle parts. Some conspecific variations were found: 1) female specimens from the type locality had head of same length, but wider, and more dilated posteriorly (HL to HW ratio 1.18); pronotum slightly sinuate; abdominal puncture intervals of ca. 3 punctures’ diameter; 2) specimen from Zhejiang: anteocular furrow slightly longer, abdominal puncture intervals of 2–3 punctures’ diameter; 3) specimen from Hainan: head and pronotum longer, eye smaller, abdominal puncture intervals of 2–3 punctures’ diameter; 4) male specimen from Guangxi: head distinctly dilated posteriorly; pronotum shorter, with distinctly sinuate lateral margin, a pair of admedian row of 11–12 punctures, abdominal puncture intervals of 2–3 punctures’ diameter; in female, abdominal puncture intervals of 2–3 punctures’ diameter. This species is similar externally to <i>X. filum</i> (Kraatz), but can be distinguished by the following characters combination: puncture intervals on head equal to 2–3 punctures’ diameter, interspaces between abdominal punctures about 4–5 punctures’ diameter, and paramere with a small denticle in inner side.</p>Published as part of <i>Zhou, Yu-Lingzi & Zhou, Hong-Zhang, 2013, Two new species of Xanthophius Motschulsky (Coleoptera: Staphylinidae, Staphylininae, Xantholinini) from China with notes on X. filum (Kraatz), pp. 363-380 in Zootaxa 3626 (3)</i> on pages 370-373, DOI: 10.11646/zootaxa.3626.3.5, <a href="http://zenodo.org/record/221592">http://zenodo.org/record/221592</a>
Stenus (Hemistenus) pullidistortus Zhao & Zhou 2005
2. <i>Stenus</i> (<i>Hemistenus</i>) <i>pullidistortus</i> Zhao & Zhou, 2005 (Fig. 1) <p> Zhao & Zhou 2005: 103 (subgenus <i>Hemistenus</i>; type locality: China: Sichuan).</p> <p> <b>Material examined.</b> <b>China:</b> Sichuan: 2 males, 3 females, Bao­xing, Feng­tong­zhai (102.84°E, 30.36°N), 1500 m, 4 Sep. 2005, Hong­Zhang Zhou collected; 1 female, Baoxing, Feng­tong­zhai (102.84°E, 30.36°N), 1500 m, 3 Sep. 2005, Hong­Zhang Zhou collected; 1 female, Bao­xing, Feng­tong­zhai (102.84°E, 30.36°N), 1500 m, 2 Sep. 2005, Hong­Zhang Zhou and Zhao Cai­Yun collected; 2 males, 2 females, Bao­xing, Feng­tongzhai, Da­shui­gou (102.84°E, 30.36°N), 1500 m, 4 Sep. 2005, Cai­Yun Zhao collected; 2 males, 14 ex., Bao­xing, Feng­tong­zhai, Da­shui­gou (102.84°E, 30.36°N), 1500 m, 3 Sep. 2005, Xiao­Yan Li collected; 1 female, Bao­xing, Feng­tong­zhai, Da­shui­gou (102.84°E, 30.36°N), 1500 m, 3 Sep. 2005, Ye­Jun Zhang collected; 3 males, Emei mountain (103.5°E, 29.62°N), 28 Aug. 2005, Cai­Yun Zhao collected; 1 female, Emei mountain (103.5°E, 29.62°N), 29 Aug. 2005, Xiao­Yan Li collected.</p> <p> <b>Remarks.</b> Among the specimens collected since we had described this species we found female specimens. The spermatheca is illustrated in Fig. 1. After careful examination, we found the paraglossae of this species are coniform.</p>Published as part of <i>Zhao, Cai-Yun & Zhou, Hong-Zhang, 2006, A new species of Stenus (Hemistenus) with new records and a key to Chinese species of the viridanus group (Coleoptera, Staphylinidae), pp. 21-29 in Zootaxa 1304 (1)</i> on page 24, DOI: 10.11646/zootaxa.1304.1.2, <a href="http://zenodo.org/record/5070970">http://zenodo.org/record/5070970</a>
Stenus (Hypostenus) yiae , Zhao & Zhou 2020, sp. nov.
2. Stenus (Hypostenus) yiae Zhao & Zhou, sp. nov. Type Material. Holotype: male, CHINA, Beijing, Chaoyang, Datunxiang (116°24′E, 39°59′N), 21.V. 1996, Hong-zhang Zhou collected [Deposited in Institute of Zoology, Chinese Academy of Sciences (IZ-CAS)]; Paratypes: 1 female, same data as holotype; 1 male, 1 female, CHINA, Beijing (116°28′E, 40°13′N), 1942; 1 male, 2 females, CHINA, Fujian, Shaxian (117°46′E, 26°25′N), 15.VII.1977, Xiao-nan Luo collected; 2 males, 3 females, CHINA, Shanghai (121°29′E, 31°13′N), A. SAVIO collected; 14 ex., CHINA, Shanghai (121°29′E, 31°13′N), O. PIEL collected. [Deposited in Institute of Zoology, Chinese Academy of Sciences (IZ-CAS)] Diagnosis. This new species belongs to the cicindeloides group. It can be easily distinguished from S. (H.) verticalis Benick and S. (H.) primivenatus Zhao & Zhou sp. nov. by the absence of spots on elytra. It is similar to S. (H.) cicindeloides (Schaller), but differs from the latter by having a broad apical part of median lobe and a different spermatheca structure. Detailed description and illustrations of the species are provided by Zhao & Zhou (2008).Published as part of Zhao, Cai-Yun & Zhou, Hong-Zhang, 2020, Validation of Stenus (Hypostenus) primivenatus and Stenus (Hypostenus) yiae (Coleoptera, Staphylinidae, Steninae), pp. 591-592 in Zootaxa 4881 (3) on page 591, DOI: 10.11646/zootaxa.4881.3.11, http://zenodo.org/record/428387
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