55 research outputs found

    Wiedemannia shamshevi Kustov & Zherebilo

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    Wiedemannia shamshevi Kustov & Zherebilo (Fig. 14) Wiedemannia shamshevi Kustov & Zherebilo, 2014: 166. Material examined. GEORGIA: Lechkhumi ridge (41), 29.iv. 2014, D. Zherebilo (1 ♂, 1 ♀); Khevsureti (47), 19.vii. 2014, D. Zherebilo (32 ♂, 54 ♀); same locality, western slope of Chokismta-Matura (49), 23.vii. 2014, D. Zherebilo (41 ♂, 67 ♀). SOUTH OSSETIA: Gudissky ridge (44), 21.vii. 2013, D. Zherebilo (6 ♂, 6 ♀). Diagnosis. This species is distinguished by the weak ocellar bristles, nearly twice shorter than postpronotal bristles, scutum brown with blue pruinescence on postpronotal lobe and notopleuron, acrostichals extending to third dorsocentral seta, 5 pairs of dorsocentrals without intermediate setae interspersed among primary setae, fore femur without strong bristles apically; terminalia large, clasping cercus long and unilobed, projecting perpendicular to epandrium. Distribution. PALAEARCTIC: Caucasus: Georgia, South Ossetia. Remarks. This high-mountain species is active from April to July, and is common in high-altitude streams and rivers (2500–3000 m a.s.l.), but is known from lower elevations (770 m a.s.l) in early spring.Published as part of Kustov, Semen & Zherebilo, Danil, 2015, New data on the genus Wiedemannia Zetterstedt (Diptera: Empididae) from the Caucasus with description of four new species, pp. 351-369 in Zootaxa 4032 (4) on page 365, DOI: 10.11646/zootaxa.4032.4.1, http://zenodo.org/record/24348

    Wiedemannia ossetiana Kustov & Zherebilo, 2015, sp. nov.

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    Wiedemannia ossetiana sp. nov. (Fig. 11) Type material. HOLOTYPE, ♂ labelled [printed in Cyrillic], [SOUTH OSSETIA] “Ju. Ossetia Gudiysskiy hrebet [ridge] / zapadnie skloni g. [western slopes of mount] Mangavtsek / rutchei [stream] / [3000 m a.s.l.] / 24.vii. 2013, D. Zherebilo (ZIN). PARATYPES: same data as holotype (1 ♀, KSU); same locality as holotype, 21.vii. 2013, D. Zherebilo (1 ♂, 8 ♀, KSU); GEORGIA: Khevsureti, south-western slope of the Mountain Tanie, 2850 m a.s.l., 19.vii. 2014, D. Zherebilo (1 ♀, KSU); Khevsureti, the northern slope of Kistanistavi, 2830 m a.s.l., 21.vii. 2014, D. Zherebilo (5 ♂, 2 ♀, ZIN, KSU); same locality, western slope of the Mountain Chokismta-Matura, source of River Argun, 2500 m a.s.l., 23.vii. 2014, D. Zherebilo (1 ♂, KSU). Diagnosis. This species differs by acrostichals divergent around the prescutellar depression, 5 pairs of dorsocentrals with interspersed shorter setae, dark brown scutum with blue pruinescence on prescutellar depression, postpronotal lobe and lower margin of notopleuron; proepisternum and laterotergite with pale setulae, fore femur with 3 anterior setae on apical quarter and unilobed, elongated clasping cercus with straight edge dorsally. Description. Male. Body length 4.2–5.2 mm; wing length 5.2 –6.0 mm. Head brown, face, occiput and gena with blue pruinescence, frons and vertex with olive-brown pruinescence. Gena broad, about 1 / 4 height of eye. Face wide, distinct carina on lower margin. Pair of strong ocellar setae (equal postpronotal setae) and pair of vertical setae, several strong occipitals similarly sized and black, with interspersed shorter postoculars and occipital. Lower postocular setae much finer and paler merging with pale, longer setae on middle and lower occiput. Few dark setulae present on vertex and between ocellar area and eye margin. Antenna dark brown to black, postpedicel and stylus minutely pubescent; scape with setulae dorsally, pedicel slightly shorter than scape, with complete circlet of apical setae; postpedicel triangular, apically pointed, stylus about twice as long as postpedicel, tip blunt. Palpus with numerous dark dorsal setae. Thoracic pleura with blue pruinescence; scutum olive-brown with indistinct darker strips between rows of acrostichals and dorsocentrals; prescutellar depression with blue pruinescence, extending onto concolourous scutellum; postpronotal lobe and lower margin of notopleuron with blue pruinescence. Antepronotum with pair of dark setae and some fine setulae. Proepisternum with pale setulae. Acrostichals small, irregularly biserial, divergent around prescutellar depression; 5 pairs of long dorsocentrals with interspersed numerous shorter setae; 1 postpronotal seta (with 1–2 short setae), 2 notopleural setae (upper stronger), 1 presutural supra-alar seta and several scattered setulae; 1 postalar seta; 1 strong scutellars setae (without weak marginal setae). Laterotergite with patch of fine pale setulae. Legs entirely dark brown to black, coxae with blue pruinescence. All legs with evenly distributed setae; hind tibia with longer dorsal and ventral setae on apical third. Fore femur narrowed apically, with 3 stout anterior setae on apical quarter. All coxae with longer setae anteriorly: hind coxae with pale setae, mid and fore coxae with mixed pale and blackish setae. Wing membrane infuscate, veins brown; pterostigma distinct, elongate, brown; single basal costal seta. Origin of veins M 1 and M 2 widely separated at end of cell dm. R 4 and R 5 straight and divergent. Halter dark brown with blue pruinescence. Abdominal sclerites with blue pruinescence. Terminalia (Fig. 11): Hypandrium subequal in length to epandrium, ventrally bearing with numerous setae on apical part. Epandrium rhomboid, covered with numerous dark long setae especially ventrally and laterally. Clasping cercus unilobed, elongate, evenly arched toward tip ventrally, with straight edge dorsally, clothed with moderately long setulae; inner surface of posterior margin with numerous setae, becoming longer on lower third. Phallus more or less linear, brownish; distiphallus without swelling in middle. Female. Similar to male, except terminalia; cercus short ovate and minutely pilose. Distribution. PALAEARCTIC: South Ossetia, Georgia. Remarks. Wiedemannia ossetiana sp. nov. is very similar to W. (Chamaedipsia) oredonensis Vaillant, 1967 known from high mountain landscapes of France and Spain. The new species differs from the latter species by almost straight and thinner phallus, very indistinct cercal plate, surstylus not prominent and details of the structure of the clasping cercus: straight dorsal edge, almost rectangular apex and distribution of inner setae (distinct row between angle of rectangular apex and posterior base). In W. (C.) oredonensis, the phallus is rather thick and distinctly S-shaped, cercal plate quite developed, surstylus distinct and clothed with moderately long setae, and details of the structure of the clasping cercus (dorsal edge not straight, apex distinctly rounded) and distribution of inner setae differs. The new species is known only from high-altitude streams and rivers of Central Caucasus (2500–3000 m a.s.l.), and is active in July. Etymology. The new species is named in reference to its type locality, South Ossetia.Published as part of Kustov, Semen & Zherebilo, Danil, 2015, New data on the genus Wiedemannia Zetterstedt (Diptera: Empididae) from the Caucasus with description of four new species, pp. 351-369 in Zootaxa 4032 (4) on pages 361-362, DOI: 10.11646/zootaxa.4032.4.1, http://zenodo.org/record/24348

    New data on the genus Wiedemannia Zetterstedt (Diptera: Empididae) from the Caucasus with description of four new species

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    Kustov, Semen, Zherebilo, Danil (2015): New data on the genus Wiedemannia Zetterstedt (Diptera: Empididae) from the Caucasus with description of four new species. Zootaxa 4032 (4): 351-369, DOI: 10.11646/zootaxa.4032.4.

    Wiedemannia pseguashae Kustov & Zherebilo, 2015, sp. nov.

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    Wiedemannia pseguashae sp. nov. (Fig. 12) Type material. HOLOTYPE, ♂ labelled [printed in Cyrillic], [SOUTH OSSETIA]: “C. Kavkaz, Ju. Ossetia, hr. [ridge] Britauli / C-B skloni g. [north-east slopes of Mountain Britavtsak / lev.bereg r. [left bank of River] Britadon / Vis. [high] 2500 m [a.s.l.] / 42 ° 29 ' 13.4 "N, 044° 10 ' 34.6 " E / 21.vii. 2013, D. Zherebilo” (ZIN). PARATYPES: SOUTH OSSETIA: same data as holotype (7 ♂, 5 ♀, ZIN; 7 ♂, 5 ♀, KSU); Central Caucasus, South Ossetia, Gudiysskiy ridge, source of River Britadon, northern slope of the Mountain Mangavtsek, 2700 m a.s.l., 21.vii. 2013, D. Zherebilo (2 ♂, 5 ♀, KSU). GEORGIA: Khevsureti, south-western slope of Mountain Tanie, 2850 m a.s.l., 19.vii. 2014, D. Zherebilo (8 ♂, 3 ♀, KSU); Khevsureti, the northern slope of Kistanistavi, 2830 m a.s.l., 21.vii. 2014, D. Zherebilo (7 ♀, KSU); same lokality, 2900 m a.s.l., 21.vii. 2013, D. Zherebilo (104 ♀, KSU). Diagnosis. This species is distinguished by acrostichals divergent around the prescutellar depression, 6–7 pairs of dorsocentrals with interspersed numerous shorter setae, dark brown scutum with light brown pruinescence on prescutellar depression; 2 anterior setae on fore femur and unilobed, elongate, narrowed apically clasping cercus with inner numerous setae on posterior margin, becoming longer on lower third. Description. Male. Body length 4.5–5.1 mm; wing length 5.0–6.0 mm. Head brown, face, occiput and gena with blue pruinescence, frons and vertex with olive-brown pruinescence. Gena broad, about 1 / 4 height of eye. Face wide, distinct carina on lower margin. Pair of strong ocellar setae (equal postpronotal setae) and pair of vertical setae, several strong occipitals similarly sized and black, with interspersed shorter postoculars. Lower postocular setae much finer and paler merging with pale, longer setae on middle and lower occiput. Few dark setulae present on vertex and between ocellar area and eye margin. Antenna dark brown to black, postpedicel and stylus minutely pubescent; scape with setulae dorsally, pedicel almost equal in size to scape, with complete circlet of apical setae; postpedicel longer, apically pointed, stylus twice as long as postpedicel, tip blunt. Palpus with numerous dark dorsal setae. Thoracic pleura with blue pruinescence; scutum dark brown with indistinct darker strips between rows of acrostichals and dorsocentrals; prescutellar depression with light brown pruinescence, extending onto concolourous scutellum; postpronotal lobe and lower margin of notopleuron with lighter pruinescence. Antepronotum with pair of dark setae and some fine setulae. Proepisternum with whitish fine setulae. Acrostichals small, irregularly biserial, divergent around prescutellar depression; 6–7 pairs of dorsocentrals with numerous interspersed shorter setae; 1 postpronotal seta (with several short setae), 2 notopleural setae (upper stronger), 1 presutural supra-alar seta and with numerous scattered setulae; 1 postalar seta; 1 strong scutellar setae and 2 weak marginal setae. Laterotergite with patch of fine, dark setulae. Legs entirely dark brown to black, coxae with blue pruinescence. All legs with evenly distributed setae; hind tibia with longer dorsal and ventral setae on apical third. All coxae with longer setae anteriorly. Fore femur narrowed apically, with 2 stout, closely approximated anterior setae on apical third and 1 same seta near tip. Wing membrane infuscate, veins brown; pterostigma distinct, elongate, brown; single basal costal seta. Origin of veins M 1 and M 2 widely separated at end of cell dm (some specimens with veins M 1 and M 2 form additional cell after dm). R 4 and R 5 straight and divergent. Halter dark brown with blue pruinescence. Abdominal sclerites with blue pruinescence. Terminalia (Fig. 12): Hypandrium subequal in length to epandrium, bearing several setae. Epandrium rhomboid, covered with numerous dark long setae especially ventrally and laterally. Clasping cercus unilobed, elongated, narrowed apically clothed with fine setulae; inner surface of posterior margin with numerous setae, becoming longer on lower third. Phallus more or less linear, brownish; distiphallus without swelling in middle. Female. Similar to male, except terminalia; cercus short ovate and minutely pilose. Distribution. PALAEARCTIC: South Ossetia, Georgia. Remarks. Wiedemannia pseguashae sp. nov. is most similar to W. (Chamaedipsia) pyrenaica Vaillant 1967 and W. (Chamaedipsia) beckeri, but differs by the structure of the male terminalia, especially the form of the clasping cercus and distribution of inner setae. Wiedemannia pseguashae sp. nov. can be differentiated from related new species described in this article as outlined in the key. The new species is known only from high-altitude streams and rivers of Central Caucasus (2500–3000 m a.s.l.) and is active in July. Etymology. The epithet refers to “Pseguash” (maid of rivers), a pagan goddess in the mythology of ancient Cherkessian.Published as part of Kustov, Semen & Zherebilo, Danil, 2015, New data on the genus Wiedemannia Zetterstedt (Diptera: Empididae) from the Caucasus with description of four new species, pp. 351-369 in Zootaxa 4032 (4) on pages 362-364, DOI: 10.11646/zootaxa.4032.4.1, http://zenodo.org/record/24348

    Wiedemannia (Eucelidia) zetterstedti Fallen

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    Wiedemannia (Eucelidia) zetterstedti (Fallén) (Fig. 17) Empis zetterstedti Fallén, 1826: 7. Material examined. RUSSIA: Krasnodarskiy Territory, env. Oktyabrskiy (2), 29.v. 2013, S.Yu.Kustov, V.V. Gladun (2 ♂, 8 ♀); env. Betta (1), 12.vi. 2013, V.V. Gladun (3 ♂, 1 ♀); Nature Reserve “Kamyshanova Polyana” (5), 24.vi. 2013, S. Kustov (12 ♂, 16 ♀). GEORGIA: Mingrelia, env. Doberazeni (28), 2.v. 2014, D. Zherebilo (25 ♂, 20 ♀); Mingrelia, ridge Ofitsari (30), 4.v. 2014, D. Zherebilo (7 ♂, 8 ♀); same data, 6.v. 2014, D. Zherebilo (31 ♂, 18 ♀); env. Chinkadziebi (25), 6.iv. 2015, A.V. Zherebilo (2 ♂); env. Zedachukhuneti (27), 7.iv. 2015, A.V. Zherebilo (1 ♂); env. Chaisubani (26), 8.iv. 2015, A.V. Zherebilo (3 ♂, 1 ♀); env. Zedakinchha (34), 9.iv. 2015, A.V. Zherebilo (1 ♂, 1 ♀); env. Opitara (35), 10.iv. 2015, A.V. Zherebilo (3 ♂, 2 ♀); env. Chkumi (36), 11.iv. 2015, A.V. Zherebilo (1 ♂); env. Namohvani (37), 15.iv. 2015, A.V. Zherebilo (2 ♂, 1 ♀); env. Mukhuri (42), 17.iv. 2015, A.V. Zherebilo (1 ♀). ABKHAZIA: River Pshitsa (9), 11.vii. 2013, D. Zherebilo (3 ♂, 2 ♀); env. Sukhum (18), 27.vi. 2013, D. Zherebilo (12 ♂). Diagnosis. Wiedemannia (Eucelidia) zetterstedti differs by the completely olive-brown scutum, minute acrostichals extending to second pair of dorsocentrals, 5 pairs of dorsocentrals without intermediate setae interspersed among primary setae, fore femur with 2–3 strong bristles at the tip, fore coxae with 1–3 erect projecting setae, longer than surrounding setae and longer than width of coxa; clasping cercus bilobed, mittenshaped. Distribution. PALAEARCTIC: Europe: Austria, Belgium, Bosnia and Herzegovina, Bulgaria, Croatia, Cyclades Is., Cyprus, Czech Republic, European Turkey, Finland, Germany, Greek mainland, Hungary, Italian mainland, Lithuania, Montenegro, North Aegean Is., Poland, Slovakia, Slovenia, Sweden, Switzerland, The Netherlands; Caucasus: Russia (Krasnodarsky Territory), Georgia, Abkhazia; Asia: Turkey (Bolu: Yedigöller National Park). Remarks. This species was noted by Joost (1981) from the Sochi River (Russia, Krasnodarskiy Territory), but in the Catalogue of Palaearctic Diptera it was mistakenly indicated from the Georgian territory. Therefore W. (E.) zetterstedti is noted from the territories of Georgia and Abkhazia for the first time. This common species is known from mountain rivers of the Caucasus from elevations between 288–2100 m a.s.l. and is active from April to July.Published as part of Kustov, Semen & Zherebilo, Danil, 2015, New data on the genus Wiedemannia Zetterstedt (Diptera: Empididae) from the Caucasus with description of four new species, pp. 351-369 in Zootaxa 4032 (4) on pages 366-367, DOI: 10.11646/zootaxa.4032.4.1, http://zenodo.org/record/24348

    Wiedemannia (Philolutra) pseudovaillanti Joost

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    Wiedemannia (Philolutra) pseudovaillanti Joost (Fig. 13) Wiedemannia (Philolutra) pseudovaillanti Joost, 1981: 188. Material examined. RUSSIA: Karachay-Cherkess Republic, env. Rozhkao (13), 8.vi. 2013, D. Zherebilo (2 ♂, 11 ♀); Krasnodarskiy Territory, Nature Reserve “Kamyshanova Polyana” (5), 24.06. 2013, S.Yu. Kustov (3 ♂, 9 ♀); same data, 3.v. 2015, S.Yu. Kustov, D. Zherebilo (2 ♀); Nature Reserve “Kamyshanova Polyana” (7), 2.v. 2014, S.Yu. Kustov (2 ♂); Nature Reserve “Kamyshanova Polyana” (8), 4.v. 2014, S.Yu. Kustov (3 ♀); same data, 17.vii. 2014, V.V. Gladun (2 ♂, 3 ♀); same data, 19.vii. 2014, V.V. Gladun (1 ♂, 2 ♀); same data, 6.ix. 2014, S.Yu. Kustov (3 ♂, 2 ♀); same data, 2.v. 2015, S.Yu. Kustov, D. Zherebilo (6 ♀); Nature Reserve “Kamyshanova Polyana” (6), 1.vi. 2014, S.Yu. Kustov (1 ♂); same data, 17.vii. 2014, Е. Pavlova, (1 ♀); env. Guamka (3), 30.04. 2015, D. Zherebilo (2 ♂, 1 ♀). GEORGIA: Samtskhe-Javakheti (29), 20.iv. 2014, D. Zherebilo (1 ♂); Shavsheti ridge (24), 4.iv. 2015, A.V. Zherebilo (2 ♂); env. Hihadziri (33), 06.iv. 2015, A.V. Zherebilo (5 ♂, 10 ♀); env. Chinkadziebi, 360 m a.s.l., 06.iv. 2015, A.V. Zherebilo (1 ♀); env. Zedachukhuneti (27), 07.iv. 2015, A.V. Zherebilo (2 ♂, 1 ♀); env. Chaisubani (26), 08.iv. 2015, A.V. Zherebilo (3 ♂, 5 ♀); env. Zedakinchha (34), 09.iv. 2015, A.V. Zherebilo (1 ♂, 1 ♀); env. Opitara (35), 10.iv. 2015, A.V. Zherebilo (1 ♂); env. Chkumi (36), 11.iv. 2015, A.V. Zherebilo (2 ♂, 1 ♀). ABKHAZIA: River Kelasuri (21), 12.iv. 2014, D. Zherebilo (1 ♂). Diagnosis. This species is distinguished by the completely dark brown scutum, acrostichals divergent around the prescutellar depression, 5–6 pairs of dorsocentrals with few intermediate setae, scutellum with several marginal setulae, fore femur apically without strong anterior setae; clasping cercus with anterior lobe short, pointed, bearing cluster of apical setae; dorsal process subtriangular, inner margin clothed in dense setae. Distribution. PALAEARCTIC: Caucasus: Russia (Krasnodarsky Territory, Karachay-Cherkess Republic), Georgia, Abkhazia. Remarks. Joost (1981) reported this species from the territory of Russian Caucasus—Dombai and Teberda, but Chvála & Wagner (1989) in the Catalogue of Palaearctic Diptera mistakenly indicated the species only from the Georgian territory. Therefore the species is noted for Georgia and Abkhazia for the first time. This species is commonly known from mountain rivers at elevations from 288–1500 m a.s.l. and is active from April to July.Published as part of Kustov, Semen & Zherebilo, Danil, 2015, New data on the genus Wiedemannia Zetterstedt (Diptera: Empididae) from the Caucasus with description of four new species, pp. 351-369 in Zootaxa 4032 (4) on pages 364-365, DOI: 10.11646/zootaxa.4032.4.1, http://zenodo.org/record/24348

    Wiedemannia (Chamaedipsia) beckeri Mik

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    Wiedemannia (Chamaedipsia) beckeri (Mik) (Fig. 1) Chamaedipsia beckeri Mik, 1889: 71. Material examined. RUSSIA: Krasnodarskiy Territory, Nature Reserve “Kamyshanova Polyana” (5), 24.vi. 2013, S.Yu. Kustov (1 ♀); same data, 24.vi. 2013, S. Kustov (1 ♂); same data, 3.v. 2015, S.Yu. Kustov, D. Zherebilo (1 ♂, 1 ♀). GEORGIA: Khevsureti (48), 21.vii. 2014, D. Zherebilo (1 ♂); Samtskhe-Javakheti (29), 20.iv. 2014, D. Zherebilo (2 ♂ 2 ♀); Lechkhumi Ridge (41), 29.iv. 2014, D. Zherebilo (1 ♂); env. Lahepa (23), 29.iv. 2014, D. Zherebilo (1 ♂, 2 ♀); Mingrelia (28), 2.v. 2014, D. Zherebilo (5 ♂, 3 ♀); ridge Ofitsari (32), 5.v. 2014, D. Zherebilo (1 ♂); Shavsheti ridge (24), 4.iv. 2015, A.V. Zherebilo (1 ♂, 1 ♀); env. Chinkadziebi (25), 6.iv. 2015, A.V. Zherebilo (1 ♂); env. Zedakinchha (34), 9.iv. 2015, A.V. Zherebilo (1 ♂, 1 ♀); env. Chkumi (36), 11.iv. 2015, A.V. Zherebilo (1 ♀). ABKHAZIA: env. Sukhum (18), 27.vi. 2013, D. Zherebilo (1 ♂); River Bzyb (11), 11.vii. 2013, D. Zherebilo (3 ♂, 1 ♀); River Pshitsa (9), 11.vii. 2013, D. Zherebilo (1 ♂). Diagnosis. This species differs by acrostichals divergent around the prescutellar depression, 5 pairs of dorsocentrals with interspersed shorter setae, olive-brown scutum (postpronotal lobe and lower margin of notopleuron slightly lighter) with triangular patch of blue pruinescence on prescutellar depression, fore femur apically without strong anterior setae; clasping cercus unilobed, elongated, broader basally and narrowed to apex. Distribution. PALAEARCTIC: Europe: Austria, Bosnia and Herzegovina, French mainland, Germany, Hungary, Italian mainland, Poland, Romania, Slovakia, Slovenia, Switzerland; Caucasus: Russia (Krasnodarskiy Territory), Abkhazia, Georgia. Remarks. Joost (1981) first reported this species from the Caucasus, collected the stream Yusengi (Russia: Kabardino-Balkaria Republic). In the Catalogue of Palaearctic Diptera (Chvála & Wagner 1989), this species was mistakenly indicated only from the territory of Georgia. Therefore W. (Chamaedipsia) beckeri is noted from the territories of Georgia and Abkhazia for the first time. Wiedemannia beckeri is common from mountain rivers of the Caucasus from elevations between 360–2830 m a.s.l. and is active from April to July.Published as part of Kustov, Semen & Zherebilo, Danil, 2015, New data on the genus Wiedemannia Zetterstedt (Diptera: Empididae) from the Caucasus with description of four new species, pp. 351-369 in Zootaxa 4032 (4) on page 354, DOI: 10.11646/zootaxa.4032.4.1, http://zenodo.org/record/24348

    Wiedemannia (Philolutra) vaillanti Joost

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    Wiedemannia (Philolutra) vaillanti Joost (Fig. 16) Wiedemannia (Philolutra) vaillanti Joost, 1981: 189. Material examined. RUSSIA: Krasnodarskiy Territory, Nature Reserve “Kamyshanova Polyana” (7), 2.v. 2014, S.Yu. Kustov (2 ♂, 1 ♀); Nature Reserve “Kamyshanova Polyana” (8), 4.v. 2014, S.Yu. Kustov (11 ♂, 4 ♀); Nature Reserve “Kamyshanova Polyana” (6), 13.vi. 2014, S.Yu. Kustov (4 ♂, 3 ♀). Karachay-Cherkess Republic, River Phiya (16), 10.vi. 2013, D. Zherebilo (59 ♂, 47 ♀); env. Rozhkao (13), 8.vi. 2013, D. Zherebilo (1 ♀); watershed of River Bolshaya Laba (12), 9.vi. 2013, D. Zherebilo (2 ♂, 7 ♀). GEORGIA: Lechkhumi ridge (41), 28.iv. 2014, D. Zherebilo (2 ♂, 1 ♀); Mingrelia, ridge Ofitsari (31), 3.v. 2014, D. Zherebilo (2 ♂, 1 ♀); ridge Ofitsari (30), 4.v. 2014, D. Zherebilo (1 ♂); same data, 6.v. 2014, D. Zherebilo (4 ♂, 2 ♀); ridge Ofitsari (32), 5.v. 2014, D. Zherebilo (1 ♂). ABKHAZIA: River Bzyb (20), 30.vi. 2013, D. Zherebilo (3 ♂, 2 ♀); same data, 1.vii. 2013, D. Zherebilo (41 ♂, 32 ♀). Diagnosis. Wiedemannia (Philolutra) vaillanti differs by the completely dark brown scutum, acrostichals divergent around prescutellar depression, 5 pairs of dorsocentrals without interspersed shorter setae, fore femur apically without strong anterior setae; clasping cercus with anterior lobe prolonged, digitiform and slender, without cluster of apical setae; dorsal process anvil-shaped, inner margin without dense setae. Distribution. PALAEARCTIC: Caucasus: Russia (Krasnodarsky Territory, Karachay-Cherkess Republic), Georgia, Abkhazia; Asia: Turkey (Bolu: Yedigöller National Park). Remarks. This species was described by Joost (1981) from Tscheget, Terskol and Dombai (Russia: Kabardino-Balkaria Republic and Karachay-Cherkessia Republic), but Chvála & Wagner (1989) in the Catalogue of Palaearctic Diptera mistakenly indicated the species from the Georgian territory. Therefore W. vaillanti is noted from the territories of Georgia and Abkhazia for the first time. This common species is known from mountain rivers of the Caucasus from elevations between 780–2100 m a.s.l. and is active from April to July.Published as part of Kustov, Semen & Zherebilo, Danil, 2015, New data on the genus Wiedemannia Zetterstedt (Diptera: Empididae) from the Caucasus with description of four new species, pp. 351-369 in Zootaxa 4032 (4) on page 366, DOI: 10.11646/zootaxa.4032.4.1, http://zenodo.org/record/24348

    Wiedemannia nartshukae Kustov & Zherebilo, 2015, sp. nov.

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    Wiedemannia nartshukae sp. nov. (Fig. 10) Type material. HOLOTYPE, ♂ labelled [printed in Cyrillic], [SOUTH OSSETIA]: “Ju. Ossetia, Gudiysskiy hrebet [ridge] / zapadnie skloni g. [western slopes of mount] Mangavtsek/ rutchei [stream] / [3000 m a.s.l.] / 24.vii. 2013, D. Zherebilo” (ZIN). PARATYPES: SOUTH OSSETIA: same data as holotype, 24.vii. 2013, D. Zherebilo (1 ♂, 1 f, KSU); Central Caucasus, Gudiysskiy ridge, source of River Britadon, northern slope of the Mountain Mangavtsek, 2700 m a.s.l., 21.vii. 2013, (1 ♀, KSU). Diagnosis. This species is distinguished by acrostichals divergent around prescutellar depression, 5 pairs of dorsocentrals without intermediate setae, dark brown scutum with light blue pruinescence on prescutellar depression, postpronotal lobe and lower margin of notopleuron; proepisternum with brownish setae, laterotergite with dark setulae; 2–3 stout anterior setae on apical third of fore femur and male clasping cercus unilobed, elongated, evenly narrowed apically, posteriorly with distinct convexity basally. Description. Male. Body length 4.1–4.6 mm; wing length 5.8–6.1 mm. Head brown, face, occiput and gena with blue pruinescence, frons and vertex with olive-brown pruinescence. Gena broad, about 1 / 4 height of eye. Face wide, distinct carina on lower margin. Pair of strong ocellar setae (equal postpronotal setae) and pair of vertical setae, several strong occipitals similarly sized and black, with interspersed shorter postoculars. Lower postocular and occipital setae finer and brownish. Few dark setulae present on vertex and between ocellar area and eye margin. Antenna dark brown to black, postpedicel and stylus minutely pubescent; scape with setulae dorsally, pedicel 1.5 X shorter than scape, with complete circlet of apical setae; postpedicel triangular, apically pointed, stylus about twice as long as postpedicel, tip blunt. Palpus with numerous dark dorsal setae. Thoracic pleura with blue pruinescence; scutum olive-brown; prescutellar depression with light blue pruinescence, extending onto concolourous scutellum; postpronotal lobe and lower margin of notopleuron with light blue pruinescence. Antepronotum with pair of dark setae and some fine setulae. Proepisternum with brownish fine setae, longer on lower part. Acrostichals small, biserial, divergent around prescutellar depression; 5 pairs of long dorsocentrals (without intermediate setae); 1 postpronotal seta (with 2–3 shorter setae), 2 notopleurals (upper stronger), 1 presutural supra-alar seta and several scattered setulae; 1 postalar seta; 1 strong pair of scutellars setae (without weak setae). Laterotergite with, fine dark (some specimens with mixed pale and brownish) setulae. Legs entirely dark brown to black, coxae with blue pruinescence. All legs with evenly distributed setae; hind tibia with longer dorsal and ventral setae on apical third. Fore femur narrowed apically, with 2–3 stout anterior setae on apical third. All coxae with longer setae anteriorly: hind coxae with pale setae, mid and fore coxae with mixed pale and blackish setae. Wing membrane infuscate, veins brown; pterostigma distinctly brown, elongate and very narrow, single basal costal seta. Origin of veins M 1 and M 2 widely separated at end of cell dm. R 4 and R 5 straight and divergent. Halter dark brown with blue pruinescence. Abdominal sclerites with blue pruinescence. Terminalia (Fig. 10): Hypandrium subequal in length to epandrium, with few setae on apical part. Epandrium rhomboid, covered with numerous dark, long setae. Clasping cercus unilobed, elongated, evenly narrowed apically, posteriorly with distinct convexity basally, clothed with setulae of different length. Inner surface of posterior margin basally with numerous long setae. Phallus slightly evenly arched, brownish. Female. Similar to male, except terminalia; cercus short ovate and minutely pilose. Distribution. PALAEARCTIC: South Ossetia. Remarks. The new species is distinguished from related species by features outlined in the key. Wiedemannia nartshukae sp. nov. is most similar to W. (Chamaedipsia) longicornis (Mik, 1887), but differs from the latter by 5 pair of dorsocentrals and structure of male clasping cercus. Among species described in this article, W. nartshukae sp. nov. closely resembles W. ossetiana sp. nov., but can be distinguished by such features as brownish setae on proepisternum, dark or mixed setulae on laterotergite, light blue pruinescence on prescutellar depression, dorsocentrals without any short setae between them, and structure of the male terminalia, especially by the form of the clasping cercus. The new species is known only from high-altitude streams and rivers of Central Caucasus (2500–3000 m a.s.l.) and is active in July. Etymology. The new species is named in honor of Russian dipterologist, Emilia P. Nartshuk (St. Petersburg).Published as part of Kustov, Semen & Zherebilo, Danil, 2015, New data on the genus Wiedemannia Zetterstedt (Diptera: Empididae) from the Caucasus with description of four new species, pp. 351-369 in Zootaxa 4032 (4) on pages 360-361, DOI: 10.11646/zootaxa.4032.4.1, http://zenodo.org/record/24348

    Wiedemannia (Philolutra) chvali Joost

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    Wiedemannia (Philolutra) chvali Joost (Fig. 4) Wiedemannia (Philolutra) chvali Joost, 1981: 186. Material examined. RUSSIA: Karachay-Cherkess Republic, env. Phiya (15), 11.viii. 2013, D. Zherebilo (1 ♂); River Kislaya (14), 12.viii. 2013, V. Gladun (1 ♂, 2 ♀). Diagnosis. This species differs by acrostichals extending only to the third dorsocentral seta, 5 pairs of dorsocentrals without intermediate setae interspersed among primary setae, brown scutum with blue pruinescence on notopleuron, fore femur apically without strong anterior setae; clasping cercus broad, mitten-shaped, lobes not slender. Distribution. Caucasus: Russia (Karachay-Cherkess Republic, Kabardino-Balkaria Republic). Remarks. In the original description of W. (Philolutra) chvali Joost (1981) the author indicated that acrostichals were absent. However the study of additional specimens shows that they all bear distinct short acrostichal setae, extending to third dorsocentral seta. This species was described from the Karachay-Cherkess and Kabardino-Balkaria Republics of Russia. In the Catalogue of Palaearctic Diptera (Chvála & Wagner 1989) the species was erroneously indicated from Georgia and we know of no records from this territory. This uncommonly collected species is known from mountain rivers from altitude 1300–2500 m a.s.l. active from June to August.Published as part of Kustov, Semen & Zherebilo, Danil, 2015, New data on the genus Wiedemannia Zetterstedt (Diptera: Empididae) from the Caucasus with description of four new species, pp. 351-369 in Zootaxa 4032 (4) on pages 356-357, DOI: 10.11646/zootaxa.4032.4.1, http://zenodo.org/record/24348
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