23,039 research outputs found
Mileewa digitata Yu & He & Yang 2021, sp. nov.
Mileewa digitata sp. nov. (Figs 1–2) Length: ³4.2–4.6 mm, ♀ 4.6–4.7 mm. Coloration. Head, thorax and forewing black. Upper half of crown brownish (black in some males), bearing three yellowish longitudinal stripes and some irregular cloudy spots. Eyes and ocelli off-white. Scutellum yellow white in distal half, medium of basal half brown yellowish, with a distinct vertical line and two round black spots beside end of vertical line (thorax and scutellum totally black, without stripe in some males). Face, thorax in ventral view and legs yellow white, pretarsus black, abdomen yellowish ventrally, subgenital plates and pygofer dark brown. Forewing with scattered yellow-brown translucent spots in basal half, outer margin with transparent edge, posterior margin with two equirotal transparent markings at medium, distal area with two preapical crescent-shaped transparent markings. External features. Head anterior margin roundly produced in dorsal view, median length of crown equal to interocular width; ocelli located on line between anterior eye angles; lateral frontal suture extending onto crown and attaining ocelli; coronal suture reaching two-third median length of crown. Face with frontoclypeus moderately convex, muscle impressions distinct, anteclypeus longitudinally swollen, clypeal suture obscure. Male genitalia. Male pygofer trapezoidal in lateral view, dorsal margin straight, surface with few microsetae, ventral process slender not reaching posterior margin, curved dorsad. Anal tube appendage absent. Subgenital plates longer than pygofer, surface bearing biserial macrosetae and slender and short setae from basal 1/4 to apex. Connective trapezoidal, longer than wide. Style slender, apex with finger-like process directed dorsad, with few teeth. Aedeagus elongate, preatrium absent, dorsal apodeme developed, aedeagal shaft sinuate in lateral view, with numerous denticule in distal half, apex either with two long processes directed basad, five small asymmetric processes or a few teeth (see Remarks); gonopore preapical on posterior surface. Etymology. The specific epithet is derived from the Latin word digitus meaning finger-like, referring to the finger-like process of the style. Material examined. Holotype ³, China: Tibet, Motuo, Beibeng, 1 August 2020, coll. Xiaofei Yu & Zaihua Yang. Paratypes: 2³³, 18♀♀, the same data as holotype; 23³³, 56♀♀, China: Tibet, Motuo, 27–31 July 2020, coll. Xiaofei Yu & Fangling Xu; 10³³, 7♀♀, China: Tibet, Motuo, Beibeng, 1–3 August 2020, lighted by Xiaofei Yu; 1³, 4♀♀, China: Tibet, Motuo, Beibeng, 13 August 2020, coll. Xianyi Wang; 3³³, 6♀♀, China: Tibet, Motuo, Dexing (1068 m), 15 August 2020, coll. Xianyi Wang. (GUGC). Remarks. This species is very similar to Mileewa octospina Yang, 2010 in external morphological characteristics, but differs from the later in having the ventral pygofer process very slender and apical processes of the aedeagus extended laterad. The variation found in the apex of the aedeagus, i.e., either with two long symmetrical processes, five small asymmetric processes or a few teeth, is treated as intraspecific.Published as part of Yu, Xiao-Fei, He, Hong-Li & Yang, Mao-Fa, 2021, Three new species of Mileewa (Hemiptera: Cicadellidae: Mileewinae) from Tibet China, pp. 435-443 in Zootaxa 4933 (3) on page 436, DOI: 10.11646/zootaxa.4933.3.9, http://zenodo.org/record/455512
Processina chandrai He & Yang & Yu 2018, sp. nov.
Processina chandrai sp. nov. (Figs. 1–4, 11–12, 22–33) Description. Crown pale orange, with pair of dark brown semilunar markings on anterior margin, small dark brown small spot and line along coronal suture medially, and dark brown longitudinal stripe on each side, extended posterad across pronotum and mesonotum (Figs. 1, 3). Eyes and ocelli pale. Forewing dark brown, translucent, veins yellow or yellowish-white (Figs. 1, 2, 3, 4). Face pale yellow (Figs. 11, 12). Venter of thorax pale, legs pale with apices of tarsi dark brown. Male genitalia. Male pygofer in lateral view (Fig. 22) square-shaped, posterior margin nearly truncate, posterior margin with several short microsetae; pygofer process robust, curving posterodorsally, with brush-like arrangement of spines apically and an acute process on dorsal side (Figs. 22, 23). Subgenital plates (Fig. 26) long, narrow, apices exceeding posterior margin of pygofer, with uniseriate macrosetae medially and some microsetae laterally. Aedeagus (Figs. 24, 25) with shaft short, flattened laterally, constricted and recurved apically, extended into a large spine-like process; basal apodeme short, fan-shaped in lateral view. Connective (Fig. 27) Y-shaped. Style (Fig. 28) slender, apex narrowly rounded with dense fine setae. Female Genitalia. Female seventh sternite (Fig. 29) with posterior margin parabolically rounded; first valvulae (Figs. 30, 31) relatively broad, parallel sided through most of length then abruptly constricted with acuminate apex, dorsal and apical sculpturing sparsely scaly; second valvulae (Figs. 32, 33) weakly sclerotized in basal two thirds, distal part with dorsal margin broadly arched and with few widely spaced short, rounded teeth, abruptly constricted distally with beaklike apex. Measurement. Male: 4.2 mm; female: 4.8 mm. Material examined. Holotype: ♂, THAILAND, Chiang Mai Doi Chiang Dao WS Nature trail, 19°24.278 'N 98 °55.311'E, 491 m, Malaise trap, 21–28. x. 2007, Songkran & Apicharl leg, T3176. Paratype: 1♀, THAILAND, Chiang Mai Doi Chiang Dao WS Nature trail, 19°24.278 'N 98 °55.311'E, 491 m, Malaise trap, 30.ix–7. x. 2007, Songkran & Apicharl leg, T3174. Etymology. The species is named in honor of Professor Chandra Viraktamath in recognition of his many contributions to knowledge of the Oriental leafhopper fauna. Remarks. This new species is readily distinguished from other species of the genus by the peculiar shape of the male pygofer process and the hooked end of the aedeagus. Distribution. Thailand.Published as part of He, Hongli, Yang, Maofa & Yu, Xiaofei, 2018, Three new species of the leafhopper genus Processina (Hemiptera: Cicadellidae: Mileewinae) from Thailand, pp. 279-287 in Zootaxa 4531 (2) on page 281, DOI: 10.11646/zootaxa.4531.2.9, http://zenodo.org/record/261464
Mileewa nigricincta Yu & He & Yang 2021, sp. nov.
<i>Mileewa nigricincta</i> sp. nov. <p>(Figs 5–6)</p> <p> <b>Length:</b> ³ 4.8 mm, ♀ 4.4–4.8 mm.</p> <p> <b>Coloration.</b> Head and thorax bright yellowish dorsally, eyes black. Face, thorax ventrally, legs and abdomen ventrally off-white, face apically, tibia and pygofer yellow. Forewing yellow-white basally, anterior and apical margins black; apically with two oval yellow-pale transparent large spots.</p> <p> <b>External features.</b> Crown distinctly produced, median length slightly greater than interocular width; crown slightly swollen, shallowly concave at ocelli; ocelli located on line between anterior eye angles and end of lateral frontal sutures; coronal suture reaching half median length of crown. Face with frontoclypeus slightly convex, muscle impressions obscure; anteclypeus longitudinally convex, apical margin slightly reflexed, clypeal suture distinct.</p> <p> <b>Male genitalia.</b> Male pygofer in lateral view with dorsal and ventral margins nearly straight, ventral margin with small setae, ventral process long not beyond posterior margin, curved dorsad, apex branched with longer branch darkly pigmented. Anal tube appendage absent. Subgenital plates extending posteriorly beyond pygofer apex, surface with macrosetae on inner margin from basal 1/3 to apex, with long or short microsetae. Connective Y-shaped, longer than wide, stem slender, lateral arms short and broad. Style slender, with small setae near apical one-third, without teeth, apex abruptly twisted laterally. Aedeagus with preatrium nearly quarter length of shaft; dorsal apodeme weakly developed, shaft in lateral view moderately broad, sinuate, apex with pair of robust tapered processes extended ventrad and basad but not divergent from each other in ventral view; gonopore preapical on posterior surface at base of processes.</p> <p> <b>Etymology.</b> The specific name refers to the brown margin of the forewing.</p> <p> <b>Material examined</b>. Holotype ³, China: Tibet, Motuo, Dexing (1068 m), 15 August 2020, coll. Xianyi Wang. Paratypes: 1♀, Tibet, Motuo, 29 July 2020, coll. Xiaofei Yu; 2♀♀, China: Yunnan, Tengchong, Shangyun (1700– 1900 m), 15 July 2002, coll. Maofa Yang (GUGC).</p> <p> <b>Remarks.</b> This new species can be easily distinguished from the other species of the genus by its unusual coloration and shape of the male genitalia.</p>Published as part of <i>Yu, Xiao-Fei, He, Hong-Li & Yang, Mao-Fa, 2021, Three new species of Mileewa (Hemiptera: Cicadellidae: Mileewinae) from Tibet China, pp. 435-443 in Zootaxa 4933 (3)</i> on pages 442-443, DOI: 10.11646/zootaxa.4933.3.9, <a href="http://zenodo.org/record/4555123">http://zenodo.org/record/4555123</a>
Mileewa furcata Yu & He & Yang 2021, sp. nov.
<i>Mileewa furcata</i> sp. nov. <p>(Figs 3–4)</p> <p> <b>Length:</b> ³3.8 mm, ♀ 3.9 mm.</p> <p> <b>Coloration.</b> Head and thorax black in dorsal view, eyes black. Face and forewing black. Thorax, legs and abdomen ventrally off-white. Subgenital plates and pygofer black.</p> <p> <b>External features.</b> Crown with anterior margin broadly rounded in dorsal view, median length about equal to interocular width; ocelli located on line between anterior eye angles; lateral frontal suture extending onto crown and attaining ocelli; coronal suture almost reaching 1/2 median length of crown. Face with frontoclypeus weakly convex, muscle impressions distinct; anteclypeus longitudinally convex, flat apically, clypeal suture obscure.</p> <p> <b>Male genitalia.</b> Male pygofer nearly triangular in lateral view, ventral margin bearing few microsetae; ventral process well developed, with elongate supbapical spine directed dorsad. Anal tube appendage absent. Subgenital plates narrow, extended to posterior margin of pygofer, surface with uniseriate macrosetae on inner margin, some long microsetae dispersed on distal one-fourth. Connective longer than wide with stem long, lateral arms short and median anterior lobe quadrate. Style slender, with microsetae near middle and small lateral tooth near distal onethird. Aedeagus with preatrium quarter length of shaft, dorsal apodeme short, apex thin with expanded subapical in-curled lateral margins and a pair of branched apical processes; gonopore apical between bases of processes.</p> <p> <b>Etymology.</b> The species name, a Latin adjective, refers to the branched or furcate aedeagal processes.</p> <p> <b>Material examined</b>. Holotype: ³, China: Tibet, Motuo, 1 August 2020, coll. Xiaofei Yu & Zaihua Yang. Paratypes: 1♀, the same data as holotype (GUGC).</p> <p> <b>Remarks.</b> This new species is similar to <i>Mileewa yigongana</i> Yang, 2005 in external morphological characteristics, but can be readily distinguished from the later by the black scutellum apex (scutellum apex yellow-white in <i>M. yigongana</i>), black face with only anteclypeus apex off-white (frontoclypeus yellow-white except base black in <i>M. yigongana</i>), and the shape of the male ventral pygofer process, subgenital plates and aedeagus.</p>Published as part of <i>Yu, Xiao-Fei, He, Hong-Li & Yang, Mao-Fa, 2021, Three new species of Mileewa (Hemiptera: Cicadellidae: Mileewinae) from Tibet China, pp. 435-443 in Zootaxa 4933 (3)</i> on pages 438-440, DOI: 10.11646/zootaxa.4933.3.9, <a href="http://zenodo.org/record/4555123">http://zenodo.org/record/4555123</a>
Pyemotes turkeyensis Yu, Zhang & He, 2010, sp. nov.
Pyemotes turkeyensis sp. nov. Figs 25–36 Description of male (Figs 25–34) Size. Idiosoma 188–196 long, 108–121 wide. Gnathosoma. Gnathosomal body 24–25 long, 28–29 wide. Seta ch 1 4–5 long; seta ch 2 4–6 long. Subcapitular setae absent. Palpal femoral seta 10–12 long. Palpal genual seta 5–6 long. Palpal tibiotarsal seta 3 long. Palpal solenidion 3 long. Prodorsum. Prodorsal shield trapezoid. Setae v 1 not observed. Setae v 2 5–8 long. Setae sc 1 11–13 long. Setae sc 2 71–82, thinner than half diameter of d, over 2.3 times as long as sc 2 – sc 2. Distance between insertions of setae: v 2 – v 2 32 –35; v 2 – sc 1 5–6; sc 1 – sc 1 34–36, nearly as wide as sc 2 –sc 2; sc 2 – sc 2 32–35; sc 1 – sc 2 5–7. Hysterosomal dorsum. Plate CD subtrapezoid in shape, with posterior margin concave. Setae c 1 15–17, 0.7– 0.8 times as long as c 2. Setae c 2 20–23 long. Distances c 1 – c 1 43–48, 1.2–1.5 times as wide as sc 2 – sc 2, c 2 – c 2 77–78. Setae d 91–124 long, long and thick, about 1.3 times as long as sc 2, over half as long as body length. Distance d–d 28–30, 0.6–0.8 times as wide as c 1 – c 1. Plate EF about as wide as long. Setae e 8–10 long. Setae f 82–100 long, thick, about 0.8 times as long as d. Distance f–f 12–14 wide, about half as wide as d–d. Setae h 1 4–9 long. Setae h 2 2–3 long, bacilliform, much shorter than h 1. Distance h 1 – h 1 15–17 apart. Setae ps 1 6 long. Setae ps 2 3–4 long, a thickened peg. Setae ps 3 4–5 long. Idiosomal venter. Apodeme 1 strong, just reaching base of trochanter I, the pair forming an angle about 90 ° medially. Apodeme 2 strongly tapered in distal 1 / 4, mediad 3 / 4 as thick as prosternal apodeme. Sejugal apodeme not well developed. Prosternal apodeme strong. Setae 1a 11 – 16 long, close to ap 1, but clearly separated from ap 1 by a short distance about diameter of setal base. Setae 1 b 6–7 long, very close to ap 2. Distance 1 a– 1 a. 14–17 apart. Setae 2a 17 – 29 long, apart from ap 2, extending beyond base of 2 b. Setae 2 b 8–12 long. Distance 2 a– 2 b 12–15 apart, 2 a– 2a 18– 20 apart, wider than 1 a– 1 a. Apodeme 3 strong, not connecting with each other, nor with ap 4. Apodeme 4 strong, not connecting with each other, but nearly connecting with ap 5. Apedeme 5 strong and long. Setae 3 a minute, 2–3 long, less than 1 / 4 as long as 3 b. Distance 3 a– 3a 23– 26 wide, about 1.3 times as wide as 2 a– 2 a. Setae 3 b 9–16 long, not reaching bases of 3 a and 4 a. Distance 3 b– 3 b 43 wide, about 1.8 times as wide as 3 a– 3 a. Setae 3 c 12–20 long, very close to and/or on ap 3. Distance 3 c– 3 c 67–68 wide, about 2.6 times as wide as 3 a– 3 a. Setae 4a 7 – 9 long. Distance 4 a– 4a 34– 35 wide, 1.3–1.5 times as wide as 3 a– 3 a. Setae 4 b 8–9 long. Distance 4 b– 4 b 20 wide, nearly 0.8 times as wide as 3 a– 3 a. Legs (Figs 27–30). Leg I 63 long (Fig. 27). Trochanter I 15 long, 0.8 times as long as wide; seta v' 5 long, not reaching base of segment, about 0.3 times as long as trochanter. Femur I 17 long, 1.3 times as long as wide, with 4 setae, d, l', l", v"; seta d 12 long, extending beyond base of l" Ge and about 0.7 times as long as femur; seta l' 8 long, over half as long as d; seta l'' 10 long; seta v'' 28 long. Genu I 9 long, about 0.8 times as long as wide, with 4 setae, l', l", v', v"; seta l' 38 long, extending beyond distal end of tarsus; seta l'' 23 long, about 0.6 time as long as l'; seta v' 10 long, about 1.1 times as long as genu; seta v'' 12 long. Tibia I 14 long, nearly 1.3 times as long as wide, with 6 setae and 1 solenidion, d, l', l", v', v", k, φ; seta d 67 long, about as long as leg I, the longest in leg I, about 5 times as long as tibia; seta l' 18 long, nearly 1.4 times as long as tibia; nearly 0.3 times as long as d; seta l'' 17 long; seta v' 12 long, about equal length of tibia; seta v'' 15 long, about 1.2 time as long as v'; microseta k 9 long, over half as long as tibia; solenidion Φ 1 4 long, about 0.4 times as long as k; solenidion Φ 2 absent. Tarsus I 14 long, nearly 1.5 times as long as wide, with 12 setae and 1 solenidion, pl', pl", pv', pv", ft', ft", tc', tc", p', p", s, u', ω; seta pl' 31 long, about 2.3 times as long as tarsus; seta pl'' 30 long, nearly as long as pl'; seta pv' 8 long, nearly 0.6 times as long as tarsus; seta pv'' 7 long, about 0.9 times as long as pv'; seta ft' and seta ft'' 11 long, eupathidial, about 0.8 times as long as tarsus; seta tc' 9 long, eupathidial, nearly 0.8 times as long as ft'; seta tc'' 12 long, eupathidial, about equal length of ft'; seta p' 10 long, eupathidial, about as long as tc'; seta p'' 9 long, eupathidial; seta s 8 long, as long as pv'; seta u' and seta u'' 3 long; solenidion ω 7 long, expanded in mid length and then tapered distally. Claw a small hook. Leg II 60 long (Fig. 28). Trochanter II 15 long, about 0.6 times as long as wide; with 1 seta, v' 6 long, not reaching base of trochanter. Femur II 19 long, over 1.2 times as long as wide; with 3 setae, d, l', v"; seta d 26 long, extending beyond base of tarsus; seta l' 9 long, about 1 / 3 times as long as d; seta v'' 11 long, about 0.4 times as long as d. Genu II 10 long, nearly 0.8 times as long as wide; with 3 setae, l', l", v'; seta l' 21 long, nearly 2.2 times as long as genu; seta l'' 14 long, about 1.4 times as long as genu, shorter than l'; seta v' 13 long, about as long as l". Tibia II 12 long, over 1.1 times as long as wide; with 4 setae, d, l', v', v"; seta d 47 long, nearly 0.8 times as long as leg II, the longest in leg II; seta l' 18 long, nearly 0.4 times as long as d; seta v' 23 long, about 1.3 times as long as l'; seta v'' 26 long, about 1.5 times as long as l'; solenidion Φ absent. Tarsus II 17 long, about 1.7 times as long as wide; with 7 setae and 1 solenidion, tc", tc', pl", pv', pv", u', u", ω; seta tc' 20 long, nearly 0.7 times as long as tc"; seta tc'' 29, nearly 1.8 times as long as tarsus; seta pl'' 11 long, about half as long as tc'; seta pv' 11 long, about equal length of pl"; seta pv'' 10 long; seta u' 14 long, nearly half as long as tc"; seta u'' 10 long; solenidion ω 7 long, as long as Ta I and not expanded in mid length. Claws strongly (relatively thick). Leg III 117 long (Fig. 29). Trichanter III 32 long, nearly 1.3 times as long as wide; seta v' 12 long, shorter than ½ Tr. Femur III 37 long, about 2.3 times as long as wide; with 2 setae, d, v'; seta d 95 long, about 0.8 times as long as leg III, extending beyond tip of leg III (including claw), slightly shorter than d Ti; seta v' 15 long, about 0.4 times as long as femur. Genu III 15, nearly 1.2 times as long as wide; with 3 setae, l', l", v'; seta l' 39 long, about 2.6 times as long as genu, reaching mid-length of tarsus; seta l'' 26 long, nearly 0.7 times as long as l', over 1.7 times as long as genu; seta v' 22 long, nearly 1.5 times as long as genu. Tibia III 21.5 long, over 1.9 times as long as wide; with 4 setae, d, l', v', v"; seta d 100 long, nearly 0.9 times as long as leg III, long and thick, thickest in leg III, as long as d Fe; seta l' 24 long, over 0.2 times as long as d; seta v' 29 long, nearly 1.2 time as long as l'; seta v'' 34 long, longer than v" and nearly 1.2 times as long as l'. Tarsus III 24 long, about 2.4 times as long as wide; with 7 setae, tc", tc', pl", pv', pv", u', u"; seta tc' 23 long, about 0.4 times as long as tc"; seta tc'' 53 long, about 2.2 times as long as tarsus; seta pl'' 12 long, half as long as tarsus; seta pv' 17 long, about 1.4 times as long as pl"; seta pv'' 14 long, over 0.8 times as long as pv'; seta u' 23 long, as long as tc'; seta u'' 15 long, about as long as pv". Claws strong. Leg IV 90 long (Fig. 30). Trochanter IV 19 long, about 0.7 times as long as wide; with 1 seta, v' 9 long. Femur IV 33 long, as long as wide; ventral margin concave; with 2 setae, d, v'; seta d 87 long, the thickest and the longest in leg IV, nearly as long as leg IV, nearly 2.7 times as long as femur; seta v' 10 long, about 0.3 times as long as femur. Genu IV 15 long, about 1.2 times as long as wide; with 2 setae, l", v'; seta l'' 21 long, about 1.4 times as long as genu; v' 15 long. Tibia IV 21 long, 1.9 times as long as wide; with 4 setae and 1 solenidion, d, l', v', v", ω; seta d 41 long, about 2 times as long as tibia; seta l' 11 long, nearly 0.3 times as long as d; v' and v'' 13 long, about 1.2 times as long as l'; solenidion Φ 13 long, extending beyond base of claw, its base close to distal end of tibia. Tarsus IV 8 long, about 1.3 times as long as wide; with 5 setae and 1 claw, tc', tc", pv', pv", u', claw; seta tc' 19 long, over 2.3 times as long as tarsus; seta tc'' 9 long, about as long as tarsus; seta pv' 7 long, over 0.8 times as long as tarsus; seta pv'' 4 long, about half as long as pv'; seta u' 10 long, nearly 1.2 times as long as tc". Claws strong, finger-like. Description of female (Figs 31–36) Size. 270 long, 110 wide. Gnathosoma. Gnathosomal body 40 long, 40 wide. Seta ch 1 7–8 long; seta ch 2.10–11 long Subcapitular setae 6 long. Palpal femoral seta 14–15 long. Palpal genual seta 7–8 long. Palpal tibiotarsal seta 5 long. Palpal solenidion 2 long. Prodorsum. Prodorsal shield posterior margin concave in middle slightly. Setae v 1 13 long. Setae v 2 13 long. Setae sc 1 20 long; its capitular body length/width 14 long, 9 wide. Setae sc 2 67 long, not reaching posterior margin of C plate. Distance between setae: v 1 – v 2 13; v 2 – sc 2 11–12; v 1 – sc 2 20; sc 2 – sc 2 48; v 2 – v 1 / v 2 – sc 2> 1. Hysterosomal dorsum. Plate C posterior margin slightly concave in middle, striate faintly. Setae c 1 21 long, 0.8 times as long as c 2. Setae c 2 27 long. Distances c 1 – c 1 40, about 0.8 times as wide as sc 2 – sc 2. Plate D posterior margin concave in middle slightly. Setae d 24–25 long. Distance d–d 27. Plate EF posterior margin convex in middle. Setae e 13 long. Setae f 45 long, about 0.7 times as long as sc 2. Distance f–f 20 wide, over 0.7 times as wide as d–d. Setae h 1 20–22 long. Setae h 2 12 long. Distance h 1 – h 1 20, as wide as f–f. Setae ps 1 8–9. Setae ps 2 absent. Setae ps 3 absent. Idiosomal venter. Apodeme 1 not reaching base of trochanter I; forming an angle of 90 ° medially. Apodeme 2 strong, just reaching base of trochanter II. Prosternal apodeme strong. Setae 1a 12 – 13 long, shorter than 1 a– 1 a (15). Setae 1 b 11–12 long, not close to apodeme 2. Setae 2a 22 – 27 long. Setae 2 b 15–17 long. Distance 1 b– 1 b 39, 2a– 2a 16, nearly as wide as 1 a– 1 a, 2 b– 2 b 40. Apodeme 3 as long as apodeme 4. Apodeme 4 strong and long. Apedeme 5 short and weaker in distal end. Setae 3a 27 – 28 long. Distance 3 a– 3a 16, as wide as 2 a– 2 a. Setae 3 b 21–28 long, nearly as long as 3 a. Distance 3 b– 3 b 35. Distance 3 a– 3 b 31–34. Distance 3 a– 3 c 22–24. Distance 3 b– 3 c 18– 21 apart. Setae 3 c 16–18 long. Distance 3 c– 3 c 50. Setae 4a 12 – 17 long. Distance 33, nearly as wide as 3 b– 3 b. Setae 4 b 17–18 long. Distance 4 b– 4 b 15, nearly half as wide as 4 a– 4 a. Tegula subtriangular, posterior margin an concave. Setae ag 15–17 long. Distance ag–ag 13. Legs (Figs 33–36). Leg I 66 long (excluding claw) (Fig. 33). Trochanter I 13 long, about 0.7 times as long as wide, with 1 seta, v' 6 long, not reaching base of segment. Femur I 17 long, nearly 1.2 times as long as wide; with 4 setae, d, l', l", v"; seta d 12 long, not reaching base of l" Ge; seta l' 11 long, about 1.8 times as long as v' Tr, and 0.9 times as long as d; seta l'' 12 long, about as long as d and not reaching base of l" Ge; seta v'' 45 long, reaching mid of tarsus, nearly 0.7 times as long as leg I, the longest in leg I. Genu I 13 long, 16 wide, about 0.8 times as long as wide; with 4 setae, l', l", v', v"; seta l' 29 long, about 2.3 times as long as genu, reaching mid tarsus and the longest in segment; seta l'' 22 long, about as long as l'; seta v' 19 long, nearly 0.7 times as long as l'; seta v'' 14 long, nearly half as long as l'. Tibia I 20 long, 16 wide, nearly 1.4 times as long as wide; with 6 setae and 2 solenidia, d, l', l", v', v", k, 2 φ; seta d 43 long, nearly 2.2 times as long as tibia and nearly 0.7 times as long as leg I; seta l' 23 long, about half as long as d; seta l'' 22 long; seta v' 17 long, about 0.8 times as long as tibia; nearly 0.9 times as long as v' Ge; seta v'' 23 long, as long as l'; nearly 0.9 times as long as v' Ge; microseta k 8 long, about 1.2 times as long as solenidion 1; solenidion Φ 1 7 long, thin throughout length; solenidion Φ 2 4 long. Tarsus I 14 long, nearly 1.2 times as long as wide; with 13 setae and 1 solenidion, pl', pl", pv', pv", ft', ft", tc', tc", p', p", s, u', u", ω; seta pl' 36 long, about 2.6 times as long as tarsus, the longest in tarsus, and nearly 0.6 times as long as leg I; seta pl'' 33 long, about 0.9 times as long as pl'; seta pv' 12 long, nearly 0.4 times as long as pl"; seta pv'' 9 long; seta ft' 10 long, eupathidium, about 0.7 times as long as tarsus; seta ft'' 12 long, eupathidium, about 1.2 times as long as ft'; seta tc' 10 long, eupathidium, as long as ft'; seta tc'' 9 long, eupathidium, 0.9 times as long as tc'; seta p' 10 long, eupathidium, as long as tc'; seta p'' 9 long, eupathidium; seta s 11 long, nearly 0.8 times as long as tarsus; seta u' and u'' abscured; solenidion ω 5 long, thick throughout length. Claw strong. Leg II 63 long (Fig. 34). Trochanter II subtriangular, 17 long, about 0.9 times as long as wide; with 1 seta, v' 10 long, extending beyond base of segment. Femur II 18 long, about 1.3 times as long as wide; with 3 setae, d, l', v"; seta d 16 long, just extending beyond destal end of genu, nearly 0.9 times as long as femur, obviously longer than l' and v'; seta l' 12 long, over 0.7 times as long as d, just reaching base of v' Ge; seta v'' 13 long, about as long as l'. Genu II 10 long, over 0.8 times as long as wide; with 3 setae l', l", v'; seta l' and seta l'' subequal, 13 long; seta v' 18 long. Tibia II 14 long, about as long as wide; with 4 setae and 1 solenidion, d, l', v', v", φ; seta d 20 long, over 1.4 times as long as tibia, as long as l'; seta l' 19 long, as long as d; seta v' 22 long, about 1.1 times as long as d; seta v'' 34 long, over 1.7 times as long as d, the longest in segment; solenidion Φ 3 long, thin. Tarsus II 15 long, over 1.5 times as long as wide; with 7 setae and 1 solenidion, tc", tc', pl", pv', pv", u', u", ω; seta tc' 14 long, about 0.6 times as long as tc"; seta tc'' 23 long, over 1.5 times as long as tarsus, nearly 0.4 times as long as leg II; seta pl'' 12 long, about half as long as tc"; seta pv' 17 long, about 1.2 times as long as tc'; seta pv'' 15 long, as long as tc'; seta u' 15 long, as long as tc'; seta u'' 8 long; solenidion ω 4 long. Leg III 76 long (Fig. 35). Trichanter III 19 long, nearly 1.7 times as long as wide; with 1 seta, v' 16 long, about 0.8 times as long as trochanter, extending beyond base of Tr. Femur III 21 long, nearly 1.8 times as long as wide; with 2 setae, d, v'; seta d 15 long, about 0.7 times as long as femur, just reaching base of tibia; seta v' 16 long, about as long as d. Genu III 10 long, over 0.8 times as long as wide; with 3 setae, l', l", v'; seta l' 18 long, not reaching distal end of tibia, just extending base of d Ti, over 1.8 times as long as genu; seta l'' 13 long, about 0.7 times as long as l'; seta v' 22 long, about 1.2 times as long as l'. Tibia III 15 long, nearly 1.2 times as long as wide; with 4 setae and 1 solenidion, d, l', v', v", φ; seta d 31 long, twice as long as tibia and about 0.4 times as long as leg III; seta l' 24 long, nearly 0.8 times as long as d; seta v' 27 long, about 1.8 times as long as tibia, nearly 0.9 times as long as d; seta v'' 22 long, 0.9 times as long as l'; solenidion Φ 2 long. Tarsus III 18 long, over 1.6 times as long as wide; with 7 setae, tc", tc', pl", pv', pv", u', u"; seta tc' 11 long, about 1 / 3 as long as tc"; seta tc'' 34 long, over 0.4 times as long as leg III, over 1.9 times as long as tarsus; seta pl'' 11 long, as long as tc '; seta pv' 15 long, nearly 1.4 times as long as tc'; seta pv'' 12 long, about as long as tc'; seta u' 18 long, about 1.2 times as long as pv'; seta u'' 12 long, as long as pv". Leg IV 93 long (Fig. 36). Trochanter IV 20 long, over 1.4 times as long as wide; with 1 seta, v' 16 long, about 0.8 times as long as trochanter. Femur IV 27 long, twice as long as wide; with 2 setae, d, v'; seta d 18 long, nearly 0.7 times as long as femur, reaching base of tibia; seta v' 20 long, about 1.3 times as long as v' Tr, about 1.1 times as long as d. Genu IV 12 long, about as long as wide; with 2 setae, l", v'; seta l'' 15 long, about 1.3 times as long as genu; v' 22 long, nearly 1.5 times as long as l". Tibia IV 21 long, about 1.9 times as long as wide; with 4 setae, d, l', v', v"; seta d 32 long, reaching tip of tarsus, about 1.5 times as long as tibia, over 0.3 times as long as leg IV; seta l' 23 long, over 0.7 times as long as d; v' 27 long, nearly 0.9 times as long as d; v'' 24 long, as long as l'. Tarsus IV 22 long, nearly 2.1 times as long as wide; with 6 setae, tc", tc', pl", pv', pv", u'; seta tc' 28 long, nearly 0.3 times as long as tc", about 1.3 times as long as tarsus; seta tc'' 105 long, over 1.1 times as long as leg, the longest in leg I, about 4.8 times as long as tarsus; seta pl '' 13 long, about half as long as tc'; seta pv' 25 long, about 0.9 times as long as tc' and as long as pv"; seta pv'' 24 long, as long as pv'; seta u' 23 long, about as long as pv". Material examined Holotype male (slide 21,437) ex culture of moths originating in Turkey in 1977, collected by Robin Wilkin. Paratype female (slide 21,476) and paratypes, 3 males (slides 21,435, 21,439, 21,442), same data as holotype. Holotype and one paratype female deposited in USNM. Other paratypes in NZAC. Etymology Named after Turkey where this species is distributed. Remarks This species can be readily distinguished from two other closely related species of this complex using the key above.Published as part of Yu, Lichen, Zhang, Zhi-Qiang & He, Limin, 2010, Two new species of Pyemotes closely related to P. tritici (Acari: Pyemotidae), pp. 1-40 in Zootaxa 2723 on pages 27-38, DOI: 10.5281/zenodo.19992
Robust factor number specification for large-dimensional elliptical factor model
10.1016/j.jmva.2019.104543JOURNAL OF MULTIVARIATE ANALYSIS17
Pyemotes zhonghuajia Yu, Zhang & He, 2010, sp. nov.
Pyemotes zhonghuajia sp. nov. (Figs 1–12) Pyemotes phloeosinus Han et al. 2008: 72 nom. nud. Description of male (Figs 1–6) Size. Idiosoma 185–232 long, 105–128 wide. Gnathosoma. Gnathosomal body 26–29 long, 26–32 wide (Fig. 1). Seta ch 1 3–6 long, antero-medial in location; seta ch 2 5–6 long, postero-lateral to ch 1. Subcapitular setae absent (Fig. 2). Palpal femoral seta (Fe) 13–14 long, much longer than other setae on ventral gnathosoma (Fig. 2). Palpal genual seta (Ge) 5 long. Palpal tibiotarsal seta 3–6 long. Palpal solenidion ω 3 long. Prodorsum. Prodorsal shield longer than wide, anterior margin strongly convex (Fig. 1). Seta v 1 a pit only on each side. Setae v 2 4–8 long. Setae sc 1 6–16 long, longer than v 2. Setae sc 2 73–89 long, half as thick as setae d; long and thin, 1.7–2.1 times as wide as sc 2 – sc 2. Distance between setal bases: v 1 – v 1 18–22; v 2 – v 2 38 –44; v 1 – v 2 12–15; v 2 – sc 1 5–6; sc 1 – sc 1 38–47, nearly as wide as sc 2 – sc 2; sc 2 – sc 2 37–46; sc 1 – sc 2 6–7. Hysterosomal dorsum. Plate CD sub-circular in shape (Fig. 1). Setae c 1 23–33 long. Setae c 2 22–48 long. Distances c 1 – c 1 42–53 wide, 1.1–1.3 times as wide as sc 2 – sc 2. Setae d 100–106 long, long and thick, 1.2–1.3 times as long as sc 2, about half as long as body length. Distance d–d 26–32, 0.5–0.7 times as wide as c 1 – c 1. Plate EF wider than long. Setae e thin and short, 10–12 long. Setae f 66–84 long, thick, 0.7–0.9 times as long as setae d. Distance f–f 11–18 wide, about half as wide as d–d. Setae h 1 7–9 long. Setae h 2 minute, 1 long, bacilliform, much shorter than h 1. Distance h 1 – h 1 7–13 apart. Setae ps 1 6–9 long. Setae ps 2 3–4 long, a thickened peg, shorter than ps 1. Setae ps 3 5–6 long. Idiosomal venter. Apodeme 1 short, anterolaterally just reaching base of trochanter I, the pair forming an angle about> 90 ° medially (Fig. 2). Apodeme 2 strongly tapered in distal 1 / 3, mediad 2 / 3 not as thick as prosternal apodeme. Sejugal apodeme not well developed. Prosternal apodeme psteriorly extending to level of insertion of setae 2 a. Setae 1a 12 – 16 long, close, but sometimes obviously separated from apodeme 1. Setae 1 b 9–11 long, close to anterolateral end of apodeme 2. Distance 1 a– 1 a. 15–19. Setae 2a 14 – 21 long, widely apart from apodeme 2, extending beyond base of 2 b. Setae 2 b 13–15 long. Distance 2 a– 2 b 12–13; 2 a– 2a 22– 29, 1.3–1.7 times as wide as 1 a– 1 a. Apodeme 3 strong, not connecting with each other but connecting with apodeme 4 and apodeme 5. Apodeme 4 strong, not connecting with each other medially, but nearly connecting with apodeme 5 of the same side. Apedeme 5 strong and long. Setae 3a 8 – 18 long, slightly shorter than 3 b. Distance 3 a– 3a 24– 29 wide, 0.8–1.1 times as wide as 2 a– 2 a. Setae 3 b 15–19 long, not reaching bases of 3 a and 4 a. Distance 3 b– 3 b 45–52 wide, 1.7 –2.0 times as wide as 3 a– 3 a. Setae 3 c 16–20 long, close to apodeme 3. Distance 3 c– 3 c 58–70 wide, 2.4–2.8 times as wide as 3 a– 3 a. Setae 4a 8 – 9 long. Distance 4 a– 4a 35– 41, 1.3–1.7 times as wide as 3 a– 3 a. Setae 4 b 6–8 long. Distance 4 b– 4 b 15–19 wide, 0.6–0.7 times as wide as 3 a– 3 a. Legs (Figs 3–6). Leg I 55 –61 long (Fig. 3), shorter than other legs. Trochanter I 13–16 long, 0.6–0.9 times as long as wide (Fig. 3); seta v' 6–10 long, not reaching base of segment, 0.4–0.8 times as long as trochanter. Femur I 17–19 long, 1.3–1.5 times as long as wide, with 4 setae, d, l', l", v"; seta d 17–21 long, extending beyond base of l" Ge and 0.9–1.1 times as long as femur; seta l' 9–11 long, 0.5–0.6 as long as d; 10–15 long; seta v'' 25–45 long. Genu I 9–11 long, 0.8–0.9 times as long as wide, with 4 setae, l', l", v', v"; seta l' 35–42 long, extending beyond distal end of tarsus; seta l" 28–34 long, about 0.8 time as long as l'; seta v' 12–13 long, about 1.2 times as long as genu; seta v'' 12 long, about as long as v'. Tibia I 16 long, 1.1–1.5 times as long as wide, with 6 setae and 1 solenidion, d, l', l", v', v", k, φ (Fig. 3); seta d 40–51 long, 0.8–0.9 times as long as leg I, the longest in leg I, about 4 times as long as tibia; seta l' 15–21 long, 1.1–1.5 times as long as tibia; nearly 0.4 times as long as d; seta l'' 14–21 long, about long as l', nearly 0.4 times as long as d; seta v' 14–16 long, 1.0– 1.2 times as long as tibia; seta v'' 18–19 long, 1.1–1.2 times as long as v'; microseta k 8–10 long, over half as long as tibia; solenidion Φ 1 4 long, about 0.4 times as long as k; solenidion Φ 2 absent. Tarsus I 13 long, 1.4–1.6 times as long as wide, with 12 setae and 1 solenidion, pl', pl", pv', pv", ft', ft', tc', tc", p', p", s, u', ω (Fig. 1); seta pl' 26–35 long, 2.0– 2.7 times as long as tarsus; seta pl'' 22–30 long, about 0.8 times as long as pl'; seta pv' 8–9 long, about 0.6 times as long as tarsus; seta pv'' 5-6 long, 0.6–0.7 times as long as pv'; seta ft' 9–11 long, eupathidial, 0.7–0.8 times as long as tarsus; seta ft'' 10–13 long, eupathidial, 1.0– 1.2 time as long as ft'; setal pair tc' 9 long, eupathidial, about as long as ft'; and tc'' 10-12 long, eupathidial; seta p' 9–11 long, eupathidial, about equal length of ft'; seta p'' 8–9 long,eupathidia, about equal length of ft'; seta s 4–7 long, as long as pv''; seta u' obscured; u'' 3–5 long; solenidion ω 6 long, expanded in mid length and then tapered distally. Claw a small hook. Leg II 63 –74 long (Fig. 4). Trochanter II 14–16 long, 0.5–0.6 times as long as wide; with 1 seta, v', 10–14 long, reaching or extending beyond base of trochanter. Femur II 20–24 long, 1.6–1.7 times as long as wide, with 3 setae, d, l', v"; seta d 26–34 long, extending beyond base of d Ti; seta l' 10–18 long, 0.4–0.5 as long as d; seta v'' 14–16 long, about half as long as d. Genu II 10 long, 0.8–0.9 times as long as wide; with 3 setae, l', l", v'; seta l' 12–17 long, 1.2–1.7 times as long as genu; seta l'' 18–22 long, 2.0– 2.2 times as long as genu, longer than l'; seta v' 16–19 long, about as long as l'. Tibia II 13–15 long, 1.2–1.3 times as long as wide; with 4 setae, d, l', v', v"; seta d 48–60 long, 0.7–0.9 times as long as leg II, the longest in leg II; seta l' 17–22 long, 0.7 – 0.4 times as long as d; seta v' 17–31 long, 1.4–1.7 times as long as l'; seta v'' 17–37 long, 1.6 –2.0 times as long as l'; solenidion Φ absent. Tarsus II 17–21 long, 2.0– 2.3 times as long as wide; with 7 setae and 1 solenidion, tc", tc', pl", pv', pv", u', u", ω; seta tc' 18–26 long, about 0.7 times as long as tc"; seta tc'' 23–33 long, about 2.2 times as long as tarsus; seta pl'' 11–13 long, 0.5–0.7 as long as tc'; setal pair pv' 11–14 long, 1.0– 1.2 times as long as pl"; pv'' 11-12 long, about time as long as pl"; seta u' 12–18 long, nearly half as long as tc"; and u'' 13–17 long, about as long as u' and pv"; solenidion ω 5–6 long, shorter than Ta I, thin throughout most of length and tapered in distal end. Claw not strongly built (relatively thin). Leg III 104–117 long (Fig. 5), the longest of all legs. Trichanter III 29–35 long, 1.40–1.6 times as long as wide; seta v' 15–22 long, longer than 1 / 2 length of segment (Fig. 5). Femur III 35 –41 long, 2.1–2.9 times as long as wide; with 2 setae, d, v' (one specimen has 3 setae on one leg but 2 on the other); seta d 63–84 long, nearly 0.7 times as long as leg III, extending beyond tip of leg III (including claw), as long as d Ti; seta v' 15–25 long, 0.4–0.7 times as long as femur. Genu III 13–16 long, 1.0– 1.4 times as long as wide; with 3 setae, l', l", v'; setal pair l' 33–50 long, 2.1–3.7 times as long as genu, extending beyond mid-length of tarsus; seta l'' 21–29 long, 1.4 –2.0 times as long as genu, extending beyond mid-length of tarsus; seta v' 17–21 long, 1.0– 1.5 times as long as genu. Tibia III 19–22 long, 1.8 –2.0 times as long as wide; with 4 setae, d, l', v', v"; seta d 71–91 long, 0.7–0.8 times as long as leg III, long and thick, thickest in leg III, as long as d of Fe; seta l' 26–31 long, about 1 / 3 as long as d; seta v' 25–31 long, 0.8 –1.0 times as long as l'; seta v'' 28–34 long, 0.9–1.3 times as long as l'. Tarsus III 20–26 long, 2.4–2.6 times as long as wide; with 7 setae, tc", tc', pl", pv', pv", u', u"; seta tc' 38 long, 0.8–0.9 times as long as tarsus; seta tc'' 39–42 long, 1.6–1.8 times as long as tarsus; seta pl'' 14–20 long, 0.6–0.9 times as long as tarsus; seta pv' 15–17 long, 0.8 –1.0 times as long as pl"; seta pv'' 13–16 long, about 0.9 times as long as pv"; seta u' 20–26 long, 0.5–0.7 times as long as tc'; seta u'' 10–18 long, 0.7–1.4 times as long as pv". Claw not strong. Leg IV 86 –97 long (Fig. 6). Trochanter IV 22–28 long, 1.0– 1.3 times as long as wide; with 1 seta, v', 11–16 long (Fig. 6). Femur IV 37 –42 long, 2.5 –3.0 times as long as wide; ventral margin concave; with 2 setae, d, v'; seta d 65–82 long, the thickest and the longest in leg IV, 0.8–0.9 times as long as leg IV, 1.8–2.3 times as long as femur; seta v' 9–11 long, 0.2–0.4 times as long as femur. Genu IV 13–15 long, 1.1–1.5 times as long as wide; with 2 setae, l", v'; seta l'' 11–22 long, 0.8–1.3 times as long as genu; v' 11–18 long. Tibia IV 19–21 long, 1.8–2.1 times as long as wide; with 4 setae and 1 solenidion, d, l', v', v", ω; seta d 24–37 long, 1.2–1.4 times as long as tibia; seta l' 12–15 long, 0.4–0.5 times as long as d; v' 12–13 long, as long as l'; v'' 13–16 long, 1.0– 1.1 times as long as v'; solenidion Φ 10–12 long, extending beyond base of claw, its base close to or at distal end of tibia. Tarsus IV 5–7 long, about as long as wide; with 5 setae and 1 claw, tc', tc", pv', pv", u', claw; seta tc' 20–30 long, 3.2 –6.0 times as long as tasus; seta tc'' 8–9 long, 1.1–1.7 times as long as tasus; seta pv' 5–8 long, as long as tarsus; seta pv'' 4–5 long, 0.5–0.8 slightly shorter than pv'; seta u' 5–12 long, 0.8 –1.0 times as long as tc". Claw strong, finger-like. Description of female (Figs 7–12) Size. 216–294 long, 92–113 wide. Gnathosoma. Gnathosomal body 39–43 long, 35–37 wide. Seta ch 1 8 long; seta ch 2 9–10 long Subcapitular setae 8–10 long. Palpal femoral seta (Fe) 14–19 long. Palpal genual seta 7–8 long. Palpal tibiotarsal seta 5–8 long. Palpal solenidion 2–3 long. Palpal supracoxal seta (e) slightly shorter than distance between setal bases of e and Fe. Prodorsum. Prodorsal shield posterior margin convex, striate faintly. Setae v 1 16 –20 long. Setae v 2 12 –15 long. Setae sc 1 17–18 long; its capitular body 11–12 long, 7 wide. Setae sc 2 74–88 long, reaching and ofte extending beyond posterior margin of C plate. Distance between setae: v 1 – v 2 12–14; v 2 – sc 2 11–13; v 1 – sc 2 19–21; sc 2 – sc 2 43–46; v 2 – v 1 / v 2 – sc 2 often 1, rarely> 1. Hysterosomal dorsum. Plate C posterior margin emarginated medially (Fig. 7). Setae c 1 20–24 long, 0.7–0.9 times as long as c 2. Setae c 2 24–29 long. Distances c 1 – c 1 41–46, about as wide as sc 2 – sc 2. Plate D posterior margin concave in middle slightly (not straight). Setae d 21–30 long. Distance d–d 29–40. Plate EF posterior margin straight or slightlyconvex in middle. Setae e 9–16 long. Setae f 43–47 long, 0.5–0.6 times as long as sc 2. Distance f–f 19–24, 0.6–0.8 times as wide as d–d. Setae h 1 25–29 long. Setae h 2 9–11 long. Distance h 1 – h 1 15–19 wide, 0.7– 0.8 times as wide as f–f. Setae ps 1 minute, less than half distance ps 1 – ps 1. Setae ps 2 absent. Setae ps 3 absent. Idiosomal venter. Apodeme 1 just reaching, sometimes slightly extending beyond base of trochanter I; pair forming an angle of> 90 ° medially. Apodeme 2 curved abruptly close to posterolateral angle of trochanter I, and weakened medially, not connecting with prosternal apodeme. Prosternal apodeme strong, extending posteriorly beyong level of insertions of setae 2 a. Setae 1a 16 – 18 long, at most as long as 1 a– 1 a (16–20). Setae 1 b 14–15 long, close to apodeme 2. Setae 2a 29 – 31 long. Setae 2 b 16–20 long. Distance 1 b– 1 b 40 –43, 2a– 2a 16– 19, 0.8 –1.0 times as wide as 1 a– 1 a, 2 b– 2 b 40–41. Apodeme 3 shorter than apodeme 4. Apedeme 5 short and weaker in distal end. Setae 3a 23 – 27 long. Distance 3 a– 3a 18– 20, 1.0– 1.2 times as wide as 2 a– 2 a. Setae 3 b 29–33 long, obviously longer than 3 a. Distance 3 b– 3 b 35–41. Distance 3 a– 3 b 26–29. Distance 3 a– 3 c 18–21. Distance 3 b– 3 c 16–21. Setae 3 c 20–22 long. Distance 3 c– 3 c 49–52. Setae 4a 16 – 30 long. Distance 32–39, about 0.8–0.9 times as wide as 3 b– 3 b. Setae 4 b 16–22 long. Distance 4 b– 4 b 10–16, 0.3– 0.5 times as wide as 4 a– 4 a. Tegula subtriangular, posterior margin slightly convex (Fig. 8). Setae ag 16–20 long about twice distance ag–ag (8–10). Legs (Figs 9–12). Leg I 70 –73 long (excluding claw) (Fig. 9). Trochanter I 15–17 long, about 0.9 times as long as wide, with 1 seta, v'; 7–8 long, extending beyond base of segment. Femur I 20–21 long, nearly 1.3 times as long as wide; with 4 setae d, l', l", v"; seta d 14–18 long, extending beyond base of l" Ge; seta l' slgihtly longer than v' TrI; seta l'' 18-19 long, often longer than d and reaching base of l" Ge; seta v'' 46–59 long, extending beyond distal end of tarsus, over 0.8 times as long as leg I, the longest in leg I. Genu I 11–14 long, nearly as long as wide; with 4 setae, l', l", v', v"; seta l' 36–37 long, 2.6–3.4 times as long as genu, nearly reaching distal end of tarsus and the longest in segment; seta l'' 31-34 long, 0.8-0.9 times as long as l'; seta v' 16–17 long, 0.4–0.5 times as long as l'; seta v'' 17 long, as long as v '. Tibia I 20 – 12 long, nearly 1.5–1.6 times as long as wide; with 6 setae and 2 solenidia, d, l', l", v', v", k, 2 φ; seta d 55–59 long, 2.8 –3.0 times as long as tibia and 0.8 times as long as leg I; seta l' and seta l'' 22–28 long, 0.4–0.5 times as long as d; seta v' 20–25 long, 1.0– 1.2 times as long as tibia; 1.1–15 times as long as v' of genu; seta v'' 15- 20 long, about 0.7 times as long as l '; microseta k 8 long, about 0.9 –1.0 times as long as solenidion 1; solenidion Φ 1 8–9 long, thin to expanded distally; solenidion Φ 2 2 long. Tarsus I 14–16 long, 1.6–1.7 times as long as wide; with 13 setae and 1 solenidion, pl', pl", pv', pv", ft ', ft", tc', tc", p', p", s, u', u", ω; seta pl' 39–47 long, about 2.8–3.3 times as long as tarsus, the longest in tarsus, and 0.6 times as long as leg I; seta pl'' 33–41 long, 0.8–0.9 times as long as pl '; seta pv' 13–16 long, about 0.3–0.5 times as long as pl"; seta pv'' 7–10 long, 0.4 0.8 times as long as pv'; seta ft' 11–13 long, eupathidium, 0.7–0.8 times as long as tarsus; seta ft'' 10–15 long, eupathidium, about 1.2 times as long as ft'; seta tc' eupathidium, about as long as ft'; seta tc'' 9-14 long, eupathidium, 0.8–1.2 times as long as tc'; seta p' eupathidium, about as long as tc '; seta p'' eupathidium, 0.7 –1.0 times as long as tc"; seta s 9–11 long, 0.6 times as long as tarsus; seta u' 2–4 long; u'' 3–4 long; solenidion ω 5–6 long, slightly expanded and distally tapered. Claw strong. Leg II 65 –73 long (Fig. 10). Trochanter II subtriangular, 18.5 long, nearly 1.3 times as long as wide; with 1 seta, v' 13 long, extending beyond base of segment. Femur II 19–20 long, nearly 1.5–1.7 times as long as wide; with 3 setae, d, l', v"; seta d 17–20 long, just extending beyond distal end of genu, nearly 0.9 times or as long as femur, obviously longer than l' and v'; seta l' 10–14 long, over 0.6 times as long as d, nearly reaching base of l' Ge; seta v'' 12–15 long, 1.1 times as long as l '. Genu II 11–14 long, 1.0– 1.2 times as long as wide; with 3 setae l', l", v'; seta l' 19–22 long, nearly 1.5–1.7 times as long as genu; seta l'' 16-25 long; seta v' 16–22 long, about as long as l". Tibia II 13–16 long, over 1.4–1.6 times as long as wide; with 4 setae and 1 solenidion, d, l', v', v", φ; seta d 31– 43 long, 1.9–3.3 times as long as tibia, as long as v '; seta l' 16–25 long, about 0.3–0.7 times as long as d; seta v' 24– 43 long, as long as d; seta v'' 38–41 long; solenidion Φ 2–3 long, thin. Tarsus II 17–22 long, 2.2–2.7 times as long as wide; with 7 setae and 1 solenidion, tc", tc', pl", pv', pv", u', u", ω; seta tc' 14–17 long, 0.3–0.4 times as long as tc"; seta tc'' 40–48 long, nearly 2.3–2.5 times as long as tarsus, 0.5– 0.7 times as long as leg II; seta pl'' 11–15 long, about 1 / 4 as long as tc"; seta pv' 14–17 long, 0.8–1.2 as long as tc'; seta pv'' 16–21 long, nearly 1.0– 11 times as long as tc '; seta u' 16–21 long, nearly 1.0– 11 times as long as tc '; seta u'' 11–13 long; solenidion ω 3–4 long, expanded slightly. Leg III 88 –92 long (Fig. 11). Trichanter III 25–28 long, 1.8–2.4 times as long as wide; with 1 seta, v' 15–22 long, about 0.6–0.8 times as long as trochanter, extending beyond base of Tr III. Femur III 23–25 long, 2.3–2.4 times as long as wide; with 2 setae, d, v'; seta d subditally located, 20–25 long, 0.7–0.8 times as long as femur, extending beyond base of tibia; seta v' 20–24 long, about as long as d. Genu III 12–13 long, 1.2–1.4 times as long as wide; with 3 setae, l', l", v'; seta l' 21–28 long, extending beyond distal end of tibia, over 1.7–2.5 times as long as genu; seta l'' 21–28 long, extending beyond distal end of tibia, over 1.7–2.5 times as long as genu; seta v' 15–21 long. Tibia III 13–20 long, 1.5–1.9 times as long as wide; with 4 setae and 1 solenidion, d, l', v', v", φ; seta d 42–48 long, 2.4–2.8 as long as tibia, about half as long as leg III; seta l' 26–30 long, 0.5–0.7 times as long as d; seta v' 33 long, nearly 1.9–2.1 times as long as tibia, 1.1–1.4 times as long as l'; seta v'' 36 long, about as long as v'; solenidion Φ 2–3 long. Tarsus III 21 long, about 2.7 times as long as wide; with 7 setae, tc", tc', pl", pv', pv", u', u"; seta tc' 17 long, about 1 / 3 as long as tc"; seta tc'' 45–52 long, 0.5–0.6 times as long as leg III, 2.0– 2.4 times as long as tarsus; seta pl'' 11–12 long, about 0.7 –08 times as long as tc'; seta pv' 17–20 long, 1.0– 1.2 times as long as tc'; seta pv'' 15–17 long, about 0.9 times as long as tc'; seta u' 20–25 long, 1.1–1.6 times as long as pv'; seta u'' 14 long, about as long as pv". Leg IV 97 –106 (Fig. 12). Trochanter IV 22–25 long, nearly 1.3–1.8 times as long as wide; with 1 seta, v' 18–25 long, 0.8–0.9 times as long as trochanter. Femur IV 29–32 long, 1.8–3.6 times as long as wide; with 2 setae, d, v'; seta d 25 long, about 0.8 times as long as femur, just reaching base of tibia; seta v' 21–27 long, 1.1–1.3 times as long as v' Tr. Genu IV 13 long, 1.3–1.5 times as long as wide; with 2 setae, l", v'; seta l'' 20–28 long, 1.7 –2.0 times as long as genu; v' 17–27 long, 0.7–1.3 times as long as l". Tibia IV 18–20 long, about 1.8–1.9 times as long as wide; with 4 setae, d, l', v', v"; seta d 43–58 long, extending beyond tip of leg, 2.4 –3.0 times as long as tibia, about half as long as leg IV; seta l' 29–32 long, 0.5–0.7 times as long as d; v' 25–32 long, 0.5–0.6 times as long as d; v'' 27–31 long, nearly as long as l'. Tarsus IV 21–25 long, 2.6–3.4 times as long as wide; with 6 setae, tc", tc', pl", pv', pv", u'; seta tc' 24–27 long, about 0.2 times as long as tc", 1.0– 1.2 times as long as tarsus '; seta tc'' 112–138 long, about 1.1–1.4 times as long as leg, the longest in leg I, 4.6–6.6 times as long as tarsus; seta pl'' 14–16 long, 0.6–0.7 times as long as tc'; seta pv' 22–27 long, 0.9–1.3 times as long as tc'; seta pv'' 24–32 long, 0.9–1.3 times as long as pv'; seta u' 29–36 long, 1.0– 1.2 times as long as pv". Specimens examined Holotype male, collected on 7 June 1999 from adults of Sinoxylon japonicum Lesne on Diospyros kaki by Changxin XU in Lianhe Viliage, Ji County, Tianjing, China (sample number as 57.11.1.xiong-beijing- 1 1); paratype female and 6 males, same data as holotype. Paratypes 8 males (slide id 1993 #1, 2, 3), collected on 22 June 1993, Zunhua County, Hebei, China, parasitic on Phloeosinus hopehi Schedl on Biata orientalis. Paratypes 1 male and 1 female, collected from Phloeosinus hopehi in Beijing, China by Liong 2 in 2003 (slides 45644, 45650) Holotype and some paratypes deposited in IZCAS, one paratype each in NZAC and BMNH. Etymology The specific name is from Chinese words: zhonghua (China) and jia (beetle)—referring to beetles being the main host of this parasitic species found in China; used as a noun in apposition. Remarks This new species was previously identified by Dr John Moser as P. tritici based on material from China submitted by Mr Deping Xiong (e.g. Zhang et al. 2008; see also footnote 1). Our detailed studies of morphology revealed numerous features in both adult females and males that show differences between this new species, the P. tritici from the USA and P. turkeyensis sp. nov. (see key to species above). In addition, femur IV (female) of P. zhonghuajia sp. nov (L/W ratio 2.7-3.4) is slender than that in P. tritici (2.1–2.5). Genetically, the new species differs from P. tritici in sequences of tox 34 gene by 11.25 % (Han et al. 2008). We also found that biologically, the new species does not parasitize the larvae of Plodia interpunctella (Hübner), whereas P. tritici is a natural enemy of that species. Hosts and distribution In addition to Sinoxylon japonicum and Phloeosinus hopehi, this species was also found on Dermestidae, Buprestidae, Stenhomalus taiwanus, Phloeosinus sp., Cryptolestes turcicus, Scolytus japonicus, S. seulensis. In laboratory rearing for mass-releases in the field, this species also feeds on the following: Coleoptera (Lasioderma serricorne Fabricius, Martianus dermestoides, Sitophilus zeamais Motschulsky, Sitophilus oryzae, Cryptorrhynchus lapathi L., Semanotus bifasciatus Motschulsky, Dendroctonus valens LeConte, Brontispa longissima Gestro, Tenebrio molitor, Scarabaeidae, De
Processina topoarmata He & Yang & Yu 2018, sp. nov.
Processina topoarmata sp. nov. ( Figs. 5–8, 13–14, 34–44) Description. Crown yellowish white, with large black spot medially and black anterior margin (Figs. 5, 7). Eyes and ocelli pale. Pronotum tawny, infused anteriorly and posteriorly with dark. Scutellum yellowish white, basal triangles dark brown (Figs. 5, 7). Forewing dark brown, translucent, basal half of costal margin black, veins dull yellow (Figs. 5, 6, 7, 8). Face pale with dark brown wide median longitudinal band across frontoclypeus, complete in male and interrupted medially in female (Figs.13, 14). Venter of thorax pale, legs pale with fore tibiae and tarsi, mid tarsi, and apices of hind tarsi dark brown. Male genitalia. Male pygofer (Fig. 34) well produced posteriorly, in lateral view with posterior margin angularly convex, ventral margin with small setae; pygofer process slender and short, extending posteriorly only to middle part of lobe, unbranched. Subgenital plates (Fig. 37) extending to pygofer apex, surface with uniseriate macrosetae medially, with many microsetae mostly on outer lateral portion. Aedeagus (Figs. 35, 36) with shaft bent dorsad subbasally, with pair of apical long process extended laterad and sinuate, and thorn-shaped median ventral process at midlength; basal apodeme well developed, broad in lateral view. Connective (Fig. 38) Y-shaped. Style (Fig. 39) slender and short, apex narrowly rounded with dense fine setae. Female Genitalia. Female seventh sternite (Fig. 40) with posterior margin parabolically rounded; first valvulae (Figs. 41, 42) relatively slender, slightly tapered through most of length then strongly and evenly tapered to acute apex, dorsal and ventral preapical sculpturing densely scaly; second valvulae (Figs. 43, 44) similar to that of P. chandrai but with dorsal teeth very poorly developed. Measurement. Male: 5.1 mm, female: 5.6 mm. Material examined. Holotype: ♂, THAILAND, Chiang Mai Doi Phahompok NP Kewl om 1/m ontane forest, 20°3.549'N 99°8.552'E, 2174 m, Malaise trap, 31.vii–7.viii.2007, Wongchai. P. leg. T2953. Paratype: 1♀, THAILAND, Mae Hong Son Namlok Mae Surin NP Beside Small stream, 19°20.803 'N 97 °59.014'E, 293 m, Malaise trap, 28.x–4.xi.2007, Areerat Kumkhun leg. T2817. Etymology. The specific name combines "topo-" (place) with "arma" (weapon) and refers to the thorn-shaped preapical process of the aedeagus. Remarks. The new species is similar to P. dashahensis in appearance, but differs from the later in having a dark brown longitudinal wide band on the frontoclypeus, the veins of the forewing off-yellow, the aedeagus with only an unpaired thorn-shaped process at midlength and the pygofer process unbranched. Distribution. Thailand.Published as part of He, Hongli, Yang, Maofa & Yu, Xiaofei, 2018, Three new species of the leafhopper genus Processina (Hemiptera: Cicadellidae: Mileewinae) from Thailand, pp. 279-287 in Zootaxa 4531 (2) on pages 281-283, DOI: 10.11646/zootaxa.4531.2.9, http://zenodo.org/record/261464
Projected estimation for large-dimensional matrix factor models
10.1016/j.jeconom.2021.04.001Journal of Econometric
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