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The odontocerid genera Psilotreta and Palaeopsilotreta (Insecta, Trichoptera) embedded in mid-Cretaceous Burmese amber
No full textWICHARD, WILFRIED, MÜLLER, PATRICK, XU, CHUN-PENG (2021): The odontocerid genera Psilotreta and Palaeopsilotreta (Insecta, Trichoptera) embedded in mid-Cretaceous Burmese amber. Palaeoentomology 4 (4): 331-338, DOI: 10.11646/palaeoentomology.4.4.8, URL: http://dx.doi.org/10.11646/palaeoentomology.4.4.
FIGURE 2 in The odontocerid genera Psilotreta and Palaeopsilotreta (Insecta, Trichoptera) embedded in mid-Cretaceous Burmese amber
No full textFIGURE 2. Palaeopsilotreta kachini sp. nov., male holotype, NIGP175454. A, Imago in dorsal view; scale bar = 2 mm. B, Drawing of forewing venation; scale bar = 2 mm. C, Male genitalia embedded in dusty amber; scale bar = 0.3 mm. D, Drawing of inferior appendages; scale bar = 0.3 mm.Published as part of WICHARD, WILFRIED, MÜLLER, PATRICK & XU, CHUN-PENG, 2021, The odontocerid genera Psilotreta and Palaeopsilotreta (Insecta, Trichoptera) embedded in mid-Cretaceous Burmese amber, pp. 331-338 in Palaeoentomology 4 (4) on page 335, DOI: 10.11646/palaeoentomology.4.4.8, http://zenodo.org/record/550799
Psilotreta fossilis WICHARD & MÜLLER & XU 2021, sp. nov.
No full textPsilotreta fossilis sp. nov. (Fig. 1) Type material. Male, NIGP175453, deposited in the Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences. Well preserved in mid-Cretaceous Burmese amber, Kachin State, Myanmar. Body well preserved; forewing in dorsal view, hindwing covered dorsally by forewing. Antennae and legs present. Male genitalia visible in ventral view. Etymology. This new species is the first fossil Psilotreta described from the mid-Cretaceous Burmese amber. Diagnosis. Filamentous antennae about three times longer than forewing lengths. In forewings media M reduced, here without base, apical part originating from R 4+5. In male genitalia harpagoes originate mesally on coxopodites. Therefore the extinct species belong very probably to the Psilotreta trimeresuri diagnostic speciesgroup (sensu Oláh & Johanson, 2010). Description. Embedded in amber, conspicuous imago by long antennae, which about three times as long as elongated forewings with a length of 7 mm. Extant Psilotreta species often with antennae longer than their forewings. In forewings (7 mm in lengths) fork I about 1/3 times longer than fork II, discoidal cell closed. Fork II sessile, arising at junction of R 4+5 and crossveins. Media M reduced, simple (M 1, 2, 3, 4), without base, represented by single apical branch, originating from R 4+5; forks III and IV absent. Cu 1 running straight, apically bifurcated to fork V; Cu 2 simple, running unforked to wing margin. Male genitalia visible ventrally, but through very cloudy amber and difficult to recognize. Only inferior appendages can be described. In ventral view inferior appendages appear narrow, elongated and two-segmented consisting of coxopodite and harpago. Shorter harpago arising from coxopodite laterally in middle and not fully reaching apical level of coxopodite. Inferior appendages covered with relatively long hairs; spines not visible at apex of harpago. Although other structures (e.g., phallic apparatus, parameres) of male genitalia cannot be clearly described, unambiguous organization of inferior appendages suggests that Psilotreta fossilis sp. nov. likely belongs to the diagnostic species group Psilotreta trimeresuri. Remarks. The wing venation of this fossil agrees largely with those of extant Psilotreta species that also have simple media (M 1, 2, 3, 4), e. g., Psilotreta baureo Malicky, 1989, Psilotreta illuan Malicky, 1989, Psilotreta enikoae Oláh & Johanson, 2010, Psilotreta kurenschikovorum Arefina-Armitage & Armitage, 2011, all extant species from Southeast Asia.Published as part of WICHARD, WILFRIED, MÜLLER, PATRICK & XU, CHUN-PENG, 2021, The odontocerid genera Psilotreta and Palaeopsilotreta (Insecta, Trichoptera) embedded in mid-Cretaceous Burmese amber, pp. 331-338 in Palaeoentomology 4 (4) on page 332, DOI: 10.11646/palaeoentomology.4.4.8, http://zenodo.org/record/550799
Palaeopsilotreta succini WICHARD & MÜLLER & XU 2021, sp. nov.
No full textPalaeopsilotreta succini sp. nov. (Fig. 3) Type material. Male, ZFMK-TRI000822, kept in the amber collection of the Museum Zoological Research Museum Alexander Koenig, Bonn, Germany (former Patrick Müller collection, BUB 3865); Mid-Cretaceous Burmese amber inclusion; Kachin State, Myanmar. Body well preserved, but a fracture across body impairs viewing. Forewing in dorsal view, hindwing covered dorsally by forewing. Antennae and legs present. Male genitalia visible in ventral view, but one hind leg across genitalia interferes with view. Etymology. The new species is named after the amber in which the fossil species is embedded. Diagnosis. Fossil extinct species embedded in mid- Cretaceous Burmese amber. Male bearing bipectinate antennae. In forewing forks I, II, V present, discoidal cell closed. Media M reduced, without base, originating from R 4+5, running to wing margin, simply forked at apex into M 1+2 and M 3+4. Cu 1 running straight, apically bifurcated to fork V (Cu 1a + Cu 1b). Cu 2 simple, running unforked to Cu 1b and further (Cu 2 + Cu 1b) straight to wing margin. In male genitalia harpago small, originating mesally on point of bifurcation of strong and slightly curved coxopodite, dorso-apically covered with irregular, dark spines. Description. Head:Antennae bipectinate, longer than forewings, consisting of a strong scapus, a small pedicellus and a long flagellum, first half with 13 flagellomeres bipectinate, last half of flagellum slightly toothed and filiform. Each of bipectinate flagellomeres bearing rami on both sides; rami partially significantly longer than flagellomere, basally slender and gradually expanded towards rounded apex. Maxillary palps five-segmented; 1 st segment smaller than 2 nd segment, 3 rd longest segment, 4 th and 5 th segments of about equal length and smaller than 3 rd segment. Wings: forewing length 6 mm; apical forks I, II and V present, M without base, (pseudo-) originating in R 4+5 and simple forked in M 1+2 and M 3+4. Discoidal cell closed, forks I and II sessile. Male genitalia: inferior appendages two-segmented. Coxopodites in ventral view uniform, strong and slightly curved towards genital centre. Harpago originating lateromesally on coxopodite, dorso-apically covered with irregular, dark spines.Published as part of WICHARD, WILFRIED, MÜLLER, PATRICK & XU, CHUN-PENG, 2021, The odontocerid genera Psilotreta and Palaeopsilotreta (Insecta, Trichoptera) embedded in mid-Cretaceous Burmese amber, pp. 331-338 in Palaeoentomology 4 (4) on pages 334-337, DOI: 10.11646/palaeoentomology.4.4.8, http://zenodo.org/record/550799
Palaeopsilotreta kachini WICHARD & MÜLLER & XU 2021, sp. nov.
No full textPalaeopsilotreta kachini sp. nov. (Fig. 2) Type material. Male, NIGP175454, deposited in the Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences. Well preserved in mid Cretaceous Burmese amber, Kachin State, Myanmar. Body well preserved, but embedded in turbid and unclear amber. Head and antennae incomplete. Forewing in dorsal view, hindwing overlapping the forewing from ventrally. Legs present. Male genitalia turbid, visible in ventral view. Etymology. The new species is named after the Kachin State (Myanmar), where the Burmese amber is found. Diagnosis. Male bearing bipectinate antennae. In forewing forks I, II, III and V present; media M apically three-branched in M 1 + M 2 and M 3+4. Cu 1 running straight, apically bifurcated to fork V; Cu 2 simple, running unforked to wing margin. Male genitalia with two-segmented inferior appendages; harpago arising laterally in middle from coxopodite and overtopping coxopodite in length. Description. Head: antennae bipectinate, longer than forewings, consisting of a long scapus, about 2/3 as long as head, a small pedicellus and more than 22 bipectinate flagellomeres, before antennae cut off. Each flagellomere bearing rami on both sides; rami longer than flagellomeres, basally slender and gradually expanded towards rounded apex. Maxillary palps five-segmented; 1 st segment smaller than 2 nd segment, 3 rd and 4 th segments of about equal lengths and longer than 2 nd and 5 th segments, both also of same length. Wings: forewing with a length of 5.5 mm; forewing venation with apical forks I, II, III and V present, M apically forked in fork III (M 1 + M 2) and M 3+4. Discoidal cell not visible, forks I and II sessile. Crossveins difficult to recognize. Male genitalia (visible ventrally):inferior appendages two-segmented. Coxopodite elongate, digitiform, in ventro-lateral view basally slender and gradually expanded towards rounded apex. Harpago originating latero-mesally on coxopodite; also basally slender and gradually expanded towards rounded apex and overtop coxopodite. Segments of inferior appendage irregularly hairy and obviously of same lengths. So far as visible: Segment X apparently covering phallic apparatus dorsally with two oblong scales. Lateral preanal appendages (or parameres?) long and straight.Published as part of WICHARD, WILFRIED, MÜLLER, PATRICK & XU, CHUN-PENG, 2021, The odontocerid genera Psilotreta and Palaeopsilotreta (Insecta, Trichoptera) embedded in mid-Cretaceous Burmese amber, pp. 331-338 in Palaeoentomology 4 (4) on page 334, DOI: 10.11646/palaeoentomology.4.4.8, http://zenodo.org/record/550799
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