166,126 research outputs found
Diamysis mesohalobia Ariani & Wittmann 2000
Diamysis mesohalobia Ariani & Wittmann, 2000 Figs 14, 15 Diagnosis (sensu lato: covering the three currently known subspecies). Appendix masculina 80–120% the length of terminal segment of antennular peduncle (Fig. 14 B). Eyes normal; eyestalks with fenestra paracornealis weakly developed (Fig. 15 K) or absent (Fig. 15 A), in any case mostly not or poorly visible. Distal segment of maxillary palpus with 4–27 distinct denticles. Presence of fringes on male carapace varies between subspecies; no such fringes in females. Basal segment of thoracic exopods with outer corner spiniform or less frequently ending in an acute edge, occasionally rounded in the first and/or in some of the median and/or posterior exopods. All pereiopods with normal carpopropodus and slender, styliform claw (Figs 14 E, 15B, M). Carpopropodus of thoracic endopods 3–8 with 3 (2; 4), 3–2 (4), 3–2, 2–3, 2–3 and 3–2 segments, respectively. Carpopropodus of endopod 3, if 3- segmented, with basal segment not longer than remaining segments combined (Fig. 15 M). Thoracic endopod 3 and often also endopod 8 with at least one among the four paradactylar setae distally pectinate in (most) females, smooth or pectinate in males. Male pleopod 4 biramous with 2-segmented sympod and with small, 2-segmented endopod (Figs 14 H, 15N); its exopod 2–(3)-segmented, with large modified seta at tip. This exopod with basal segment bearing a smaller smooth seta (Figs 14 H, 15D), in certain populations occasionally with an additional barbed seta (Fig. 15 N). Endopod of uropod with one strong spine below statocyst, statolith composed of vaterite. Telson subquadrangular (Fig. 15 S) to subtriangular (Fig. 15 J); maximum width is 1.4–3.0 times that at apex; its apical cleft with 8–39 laminae; cleft is 5–26% telson length. Taxonomy. Interbreeding experiments by Ariani & Wittmann (2000) indicated mutual crossability between morphologically different Mediterranean populations of D. mesohalobia. The three main morphotypes distinguished were, therefore, described at subspecific level as D. mesohalobia mesohalobia Ariani & Wittmann, 2000, D. mesohalobia gracilipes Ariani & Wittmann, 2000, and D. mesohalobia heterandra Ariani & Wittmann, 2000; each of these subspecies are treated in separate subchapters below. Occurrence (Fig. 16). Marine, brackish and fresh (near)-coastal waters of the E-Mediterranean and Marmora Seas.Published as part of Wittmann, Karl J., Ariani, Antonio P. & Daneliya, Mikhail, 2016, The Mysidae (Crustacea: Peracarida: Mysida) in fresh and oligohaline waters of the Mediterranean. Taxonomy, biogeography, and bioinvasion, pp. 1-70 in Zootaxa 4142 (1) on page 29, DOI: 10.11646/zootaxa.4142.1.1, http://zenodo.org/record/26110
Diamysis lagunaris Ariani & Wittmann 2000
<i>Diamysis lagunaris</i> Ariani & Wittmann, 2000 <p>Fig. 17 A–J</p> <p> <i>Mysis bahirensis</i> G. O. Sars, 1877 (partim: material from La Spezia): Gourret 1897; Sudry 1910.</p> <p> <i>Diamysis bahirensis</i>: Băcescu 1941; Genovese 1956; Drake <i>et al.</i> 1997; Cunha <i>et al.</i> 2000; San Vicente & Munilla 2000; Goulletquer <i>et al.</i> 2002; Munilla & San Vicente 2005.</p> <p> <i>Diamysis bahirensis</i> ssp.: Ariani 1979 (partim: material from Lake Ganzirri).</p> <p> <i>Diamysis</i> sp. B: Wittmann 1999.</p> <p> <i>Diamysis</i> sp.: Wittmann & Ariani 2000.</p> <p> <i>Diamysis lagunaris</i> Ariani & Wittmann, 2000: 2004, 2005; Ariani 2004; Anderson 2008; Petrescu & Wittmann 2009; Petryashov 2009; Wittmann & Ariani 2009, 2010, 2012a; San Vicente 2010; ITIS 2014; Mees 2014; Wittmann <i>et al.</i> 2014.</p> <p> <b>Material examined.</b> Two samples from marine waters of the eastern Mediterranean, 32 samples from brackish and marine waters of the western Mediterranean, plus 3 from the E-Atlantic (Portugal): see Ariani & Wittmann (2000), Wittmann & Ariani (2012a). Among these 37 positive samples only one from the oligohaline reach (<i>S</i> = 3.4): 1 M subad. 4.7 mm from the Mediterranean coast of France, Canal d'Arles à Fos, 43.4663N 004.8338E; previously unpublished sample: 2 M ad. 5.4–5.6 mm, 1 F ad. 6.7 mm, among ~30,000 <i>Mesopodopsis slabberi</i> and 2 <i>Limnomysis benedeni</i>, Mediterranean coast of France, estuary of the Petit Rhône at Tiki, same sample as indicated above for <i>M. slabberi</i>, NHMW reg. no. 25707.</p> <p> <b>Diagnosis</b> (sensu lato: covering the known population range). Eyes normal, eyestalks dorsally with welldeveloped fenestra paracornealis (Fig. 17 B), although not well visible in poorly pigmented eyestalks. Rostrum forms a wide convex angle with broadly rounded tip (Fig. 17 A, B). Carapace without fringes in both sexes (Fig. 17 A). Palpus of maxilla with distal segment subcircular, armed with 5–25 distinct denticles. Pereiopods of moderate length, eighth endopod extending to the maxillae or at most up to mandibles. All pereiopods with normal carpopropodus and slender, styliform claw (Fig. 17 D). Basal segment of thoracic exopods with outer corner spiniform (Fig. 17 C) or occasionally rounded in some of the posterior exopods, most often rounded in last exopod. Pereiopods poorly to markedly slender, with R6 = 4.8–8.1 (Fig. 17 D). Carpopropodus of thoracic endopods 3–8 with 3–2 (4), 2–3, 2, 2, 2, and 2–3 segments, respectively; tarsus slender, with slender, in part feebly serrated claw; carpopropodus 3 longer than 5 times its maximum width (Fig. 17 D). Exopod of fourth male pleopod 2-segmented with a large modified seta and often an additional minute seta at tip; basal segment with smooth seta and one (0–2) additional, small, barbed seta; endopod with distinct subbasal articulation (Fig. 17 E). Scutellum paracaudale terminally well rounded or biconvex with rounded (rarely acute) apex (Fig. 17 F–H), its lower margin occasionally almost straight. Endopod of uropod with one strong spine below statocyst, statolith composed of vaterite. Telson (Fig. 17 J) subquadrangular to subtriangular, length 1.1–1.5 its maximum width or 0.7–1.0 times length of last abdominal somite; maximum width near basis 2.1–2.7 times that at apex; each lateral margin armed with 6–16 spines. Apical cleft 11–19% telson length, cleft lined by 9–23 laminae, its margins straight to convex.</p> <p> <i>Body length.</i> Adult females 4.1–8.1 mm, males 3.6–6.6 mm.</p> <p> <b>Distribution</b> (Fig. 6). Mainly in the western Mediterranean: along the coasts of the Tyrrhenian, Sardinian and Ligurian Seas, Golfe du Lion, Strait of Messina; rare in the eastern Mediterranean: Island of Crete in the Aegean Sea. The populations at the Atlantic coasts of southern Spain and Portugal may have originated from Mediterranean lagoons by transfer in ballast water (Cunha <i>et al.</i> 2000: as <i>D. bahirensis</i>), although an indigenous status of the Atlantic populations is not excluded (Wittmann & Ariani 2012a). Type locality is the mixoeuhaline to weakly metahaline lagoon Lago di Caprolace at the Lazio coast, Tyrrhenian Sea. The species is mostly found in mixoeuhaline to metahaline lagoons, also in marine coastal habitats as well as mesohaline to mixoeuhaline reaches of estuaries. Normal salinity range 14–49; so far only two positive samples from the oligohaline reach (<i>S</i> = 2–3), taken at different stations in the Rhône Delta on the Mediterranean coast of France.</p>Published as part of <i>Wittmann, Karl J., Ariani, Antonio P. & Daneliya, Mikhail, 2016, The Mysidae (Crustacea: Peracarida: Mysida) in fresh and oligohaline waters of the Mediterranean. Taxonomy, biogeography, and bioinvasion, pp. 1-70 in Zootaxa 4142 (1)</i> on pages 36-38, DOI: 10.11646/zootaxa.4142.1.1, <a href="http://zenodo.org/record/261102">http://zenodo.org/record/261102</a>
Abyssomysis Wittmann 2020, gen. nov.
Abyssomysis gen. nov. urn:lsid:zoobank.org:act: 41771FE0-7B94-4422-803C-C4B2292B9BAF Type species Abyssomysis cornuta gen. et sp. nov. by monotypy and present designation. Diagnosis Calyptommini with eyeplate formed by a transverse, median lamina (= ocular symphysis) connecting separate, rudimentary eyestalks; lamina not incised, visual elements absent. Carapace normal, anteriorly rounded.Antennula with well-developed, setose appendix masculina; proximal segment of trunk without ventral carina. Antennal sympod without spiniform extension; antennal scale well developed, entire, its smooth outer margin ending in a prominent tooth, terminal lobe well-developed. Labrum with small, knob-like rostral process. Mandibles with distinct molar part; lacinia mobilis present on left mandible, absent or vestigial on right mandible. First thoracopods normal; basis of endopod with long, setose endite. Females with three pairs of functional oostegites. Penes normal. Pleopods of both sexes reduced to unsegmented, plate- or rod-like endopods, except that the fourth male pleopod is sub-segmented. Both rami of the uropods unsegmented, setose all around, without spines. Telson short, terminally truncate; lateral margins smooth, terminal margin with spines. Etymology The generic name is an amalgamated noun with female ending, referring to the occurrence of this mysid in the abyssal plain.Published as part of Wittmann, Karl J., 2020, Lophogastrida and Mysida (Crustacea) of the " DIVA- 1 " deep-sea expedition to the Angola Basin (SE-Atlantic), pp. 1-43 in European Journal of Taxonomy 628 on page 20, DOI: 10.5852/ejt.2020.628, http://zenodo.org/record/375614
Diamysis mesohalobia subsp. mesohalobia Ariani & Wittmann 2000
Diamysis mesohalobia mesohalobia Ariani & Wittmann, 2000 Fig. 14 Diamysis bahirensis: W. M. Tattersall 1927; Ariani 1966 (partim), 1979 (partim), 1981a (partim); Ariani et al. 1981, 1982, 1983, 1993 (partim); De Matthaeis et al. 1982 (partim); Wittmann et al. 1990, 1993 (partim); Schlacher et al. 1992. Diamysis bahirensis ssp.: Ariani 1981b (partim). Diamysis sp.: Wittmann & Stagl, 1996 (partim); Ariani et al. 1999. Diamysis sp. A: Wittmann 1999. Diamysis mesohalobia Ariani & Wittmann, 2000: 2002, 2004; Ariani et al. 2000; Petrescu & Wittmann 2009; San Vicente 2010; Daneliya & Petryashev 2011; ITIS 2014. Diamysis mesohalobia mesohalobia Ariani & Wittmann, 2000: 2005; Kocataş et al. 2003; Ariani 2004; Özbek et al. 2004; Remerie et al. 2004; Özbek & Ustaoğlu 2006; Anderson 2008; Wittmann & Ariani 2010; Mees 2014. Material examined. 84 samples from mesohaline to mixoeuhaline coastal waters of the Adriatic, Aegean and Levantine Seas (Ariani & Wittmann 2000). Previously unpublished sample: 93 F ad. 5.5–7.1 mm, 223 M ad. 4.3– 6.3 mm, 22 subad., 56 imm., 112 juv., karstic spring with small salinity fluctuations, Fiume Piccolo, near Torre Canne, Adriatic coast of southern Italy, 40.8275N 017.4818E, 2–3 m depth, striped with hand net from brown algae, S = 15, pH 7, 18.5°C, 29 Nov. 2011, leg. A. P. Ariani. One specimen of this sample reminds of a gynandromorph of the 'fore and aft' type (Hollingsworth 1960): with pleopods as typical for adult males, but antennula without appendix masculina and also without plumose setae typical of males or females. Short updated description. The following data cover primarily the type population in the mesohaline spring Fiume Morello (Apulia, Adriatic Sea). Data from remaining populations, as far as different, are given in square brackets. Diamysis mesohalobia with short rostrum forming a wide convex angle with rounded tip (Figs 14 A, C). Fenestra paracornealis poorly developed, rarely visible. Carapace without fringes (Fig. 14 C) in both sexes. Palpus of maxilla with subcircular terminal segment, armed with 6–24 [5–24] denticles along distal margin. Basal segment of thoracic exopods with outer corner spiniform (Fig. 14 D), less frequently ending in an acute or rounded edge, especially in posterior exopods. Pereiopods relatively short, endopod 8, when stretched anteriorly, extending to basis of endopod 1 or at most up to maxillae [mandibles]. Pereiopods stout (Fig. 14 E) to moderately slender, with R6 = 4.4–6.1 [4.4–6.8]. Carpopropodus of thoracic endopods 3–8 with 3 (2), 3 (2), 3–2, 2–3, 2–3, and 3 (2) [3–2] segments, respectively. Thoracic endopod 3 with carpopropodus being longer than 5 times its maximum width; thoracic endopods 3–8 with long and slender claw. Penis with smooth setae only, arranged in a semicircle close to ejaculatory opening (Fig. 14 F). Male pleopod 4 biramous with 2-segmented exopod bearing a modified, strong seta at tip and a smaller, smooth seta subterminally on basal segment (Fig. 14 H). Scutellum paracaudale subtriangular, biconvex; tip pointed or less frequently rounded (Fig. 14 J–M). Telson subquadrangular (Fig. 14 N) [to subtriangular], 0.8–0.9 [0.7–1.0] times length of last abdominal somite; maximum width of telson is 1.6–2.0 [1.6– 3.0] times that at apex; lateral margins concave or rarely straight, armed with 7–12 [7–14] spines. Apical cleft of telson with convex (Fig. 14 N) or rarely straight margins, bottom of cleft rounded, cleft is 12–16% [9–16%] telson length, cleft lined by 15–39 [8–39] laminae. Body length: Adult females 4.6–9.6 mm, males 4.1–7.7 mm. Distribution (Fig. 16). Eastern Mediterranean only: in Adriatic, Aegean, and Levantine Seas. Mostly in mesohaline karstic springs with small salinity fluctuations, also in mesohaline to mixoeuhaline lagoons and estuaries. Predominantly in the salinity range 10–38, locally down to S = 2. Samples of this subspecies were taken by Özbek et al. (2004) in the Köyceğiz lagoon at the Aegean coast of Turkey. Data by Akin et al. (2005) and own measurements at the northern shore (S = 2.3 in 0.5–2 m, 10 June 2006) suggest that the positive station was most likely from the oligohaline range within this large, oligo- to metahaline lagoon with complex salinity patterns.Published as part of Wittmann, Karl J., Ariani, Antonio P. & Daneliya, Mikhail, 2016, The Mysidae (Crustacea: Peracarida: Mysida) in fresh and oligohaline waters of the Mediterranean. Taxonomy, biogeography, and bioinvasion, pp. 1-70 in Zootaxa 4142 (1) on page 32, DOI: 10.11646/zootaxa.4142.1.1, http://zenodo.org/record/26110
Diamysis mesohalobia subsp. heterandra Ariani & Wittmann 2000
Diamysis mesohalobia heterandra Ariani & Wittmann, 2000 Fig. 15 K–S Mysis oculata var. relicta: Zimmer 1927 (partim: Lake Deran). Diamysis bahirensis: Holmquist 1955; Avĉin et al. 1973 (partim); Matjašič & Štirn 1975 (partim); Ariani et al. 1983, 1993 (partim). Diamysis bahirensis ssp.: Ariani 1981b (partim). Diamysis sp.: Wittmann & Stagl 1996 (partim). Diamysis mesohalobia heterandra Ariani & Wittmann, 2000: 2004, 2005; Anderson 2008; San Vicente 2010; Wittmann & Ariani 2010, 2012a, 2012b; Mees 2014. Material examined. 42 samples from oligohaline to metahaline lagoons and karstic springs in diverse parts of the eastern Mediterranean plus 13 samples from freshwater tributaries of the Adriatic Sea, see Ariani & Wittmann (2000) and Wittmann & Ariani (2012b); material studied by Holmquist (1955), previously unpublished reexamination: dissected parts of 1 M ad. 8 mm, 2 egged F ad. with body length 7 or 9 mm, respectively, on a total of 3 slides labelled " Diamysis biharensis. Herzegovina & Timavo, det. Ch. Holmquist, prep. 57–59", SMNH reg. nos. 140108 – 140110. According to Holmquist (1955) this material was collected by Janez Hoenigman in Lake Deran, 2–5 m depth, 27 Apr. 1954, 43.04N 017.75E, Herzegovina, altitude 0 m, sea distance 31 km as calculated along the small effluent and following this along the Neretva River to the east coast of the Adriatic Sea. Short updated description. The following data covers primarily the type population in an oligo- to mixoeuhaline lagoon with brackish spring, Limni Antinioti (Island of Corfu, Ionian Sea). Data from remaining populations, as far as different, are given in square brackets. Diamysis mesohalobia with short rostrum mostly forming a wide convex angle with broadly rounded tip (Fig. 15 K, L). Fenestra paracornealis weakly developed, mostly visible (Fig. 15 K) in well preserved material [mostly visible in well preserved material from Lake Deran (near E-Adriatic coast) or rarely from Schiavetti Springs (Gulf of Trieste, N-Adriatic)]. Carapace of adult males with fringes arranged in two submedian stripes plus one subterminal stripe (Fig. 15 K, L). The submedian stripes may be differentiated as two separate stripes each (Fig. 15 K). Palpus of maxilla with subcircular terminal segment, armed with 8–27 denticles along distal margin. Basal segment of all thoracic exopods normally with spiniform outer corner, rounded only in some of the posterior exopods of small individuals ( 6 mm) often with a minute additional seta; basal segment subterminally with a smooth seta (Fig. 15 N–P). Large males (> 7 mm) with 0–1 [0–4] additional small barbed [and/or smooth] seta (Fig. 15 N, P) on terminal margin of basal segment of exopod. Scutellum paracaudale subtriangular, mostly biconvex [or with upper margin convex and lower margin concave]; tip pointed (Fig. 15 Q, R) or less frequently rounded, rarely bifid. These margins mainly smooth in small specimens [or undulate in large ones (> 8 mm; Fig. 15 Q)]. Telson mostly subquadrangular (Fig. 15 S), but subtriangular in small specimens (<6 mm), 0.7–0.9 [0.7–1.0] times length of last abdominal somite; lateral margins concave [to straight], armed with 8–13 [6–11] spines; maximum width of telson is 1.8–2.4 [1.4–2.4] times that at apex; its apical cleft with straight to strongly convex margins. Bottom of cleft angular to rounded. Cleft is 10–19% [10–26%] telson length, cleft lined by 12–31 [9–38] laminae (Fig. 15 S). Body length. Adult females 3.7–9.7 mm, males 3.0– 8.7 mm. Distribution (Figs 12, 16). In fresh and brackish waters of springs, estuaries, lagoons, and lakes all around the Adriatic Sea (Fig. 12), salinity range S = 0–42. Outside the Adriatic (Fig. 16) known only from oligo- to polyhaline waters on the coasts of the Ionian and Marmora Seas, so far not from fresh-water (Ariani & Wittmann 2000, Wittmann & Ariani 2012a, b).Published as part of Wittmann, Karl J., Ariani, Antonio P. & Daneliya, Mikhail, 2016, The Mysidae (Crustacea: Peracarida: Mysida) in fresh and oligohaline waters of the Mediterranean. Taxonomy, biogeography, and bioinvasion, pp. 1-70 in Zootaxa 4142 (1) on pages 33-36, DOI: 10.11646/zootaxa.4142.1.1, http://zenodo.org/record/26110
Ipirophthalmus Wittmann 2020, gen. nov.
Ipirophthalmus gen. nov. urn:lsid:zoobank.org:act: CC 361198 -98 AD- 4399-8483-3912 AA 62D2A0 Petalophthalmus – Pillai 1965: 1684 (partim). — O.S. Tattersall 1968: 273 (partim). — Panampunnayil 1982: 643 (partim). — Wang 1998: 202–203. — San Vicente & Corbari 2015: fig. 8 (partim). Type species Petalophthalmus liui Wang, 1998, by present designation. Etymology The generic name is a Latinized noun with masculine ending, formed as a condensate of the Ancient Greek adjective ύπηρεΤΙΚός (functional) with the noun ὀφθαλμὸς (eye), underlining the difference from the reduced visual elements in the genus Petalophthalmus. Diagnosis Petalophthalminae with antennal scale setose all around; eyes with well-developed, functional cornea; anterior margin of carapace with rostrum flanked by a pair of processes; thoracic endopods 5–8 slender, each with more than three segments; flagellum of thoracic exopods 3–8 with 10–14 segments; male pleopods biramous, female pleopods uniramous or biramous; telson large, subquadrangular, terminal margin truncate without or with distinct mid-terminal emargination, terminal margin with two submedian groups of small teeth, with 7–8 large, toothed spines and 1–2 pairs of long setae, these setae arising from the emargination, if present, otherwise from homologous submedian position. Species included – Ipirophthalmus liui (Wang, 1998) gen. et comb. nov., recombined from Petalophthalmus liui Wang, 1998, known only from the northern South China Sea, depth 194–219 m; – Ipirophthalmus caribbeanus (O.S. Tattersall, 1968) gen. et comb. nov., recombined from Petalophthalmus caribbeanus O.S. Tattersall, 1968, from the Caribbean, mesopelagic in 329–366 m depth; – Ipirophthalmus oculatus (Illig, 1906) gen. et comb. nov., recombined from Petalophthalmus oculatus, Illig, 1906, from the NW-Arabian Sea, W-Indian Ocean, off Japan, mesopelagic to bathybenthic, depth Ĺ 1200 m; – Ipirophthalmus macrops (Tchindonova & Vereshchaka, 1991) gen. et comb. nov., recombined from Petalophthalmus macrops Tchindonova & Vereshchaka, 1991, from the W-Indian Ocean: off Madagascar, benthopelagic in 360–555 m depth.Published as part of Wittmann, Karl J., 2020, Lophogastrida and Mysida (Crustacea) of the " DIVA- 1 " deep-sea expedition to the Angola Basin (SE-Atlantic), pp. 1-43 in European Journal of Taxonomy 628 on pages 14-15, DOI: 10.5852/ejt.2020.628, http://zenodo.org/record/375614
Mysidium (Occimysidium) Wittmann & Wirtz, 2019, subgen. nov.
Occimysidium Wittmann subgen. nov. urn:lsid:zoobank.org:act: EE9F05D3-29B5-4B95-BC0D-2A6756AAECE5 Type species Mysidium pumae Ortiz, Hendrickx & Winfield, 2017, by monotypy. Etymology Noun in nominative singular with neutral ending, formed by amalgamation of the Latin noun ‘ occidens ’ with the generic name Mysidium, referring to the occurrence on the west coast of America. Diagnosis Subdivision of the genus Mysidium Dana, 1852, characterized by two-segmented carpopropodus of third thoracic endopod; merus of this endopod without serrated setae. Third male pleopod with distinct endite at about ¾ distance from basis of medial margin. Sympod of fourth male pleopod without setae on rostral face. Its exopod four-segmented; basal segment with 0–1 seta, second segment without seta, subterminal segment with bifid modified seta, terminal segment with undivided modified seta. Telson entire (not incised). Distribution As for type species. Mysidium (Occimysidium) pumae Ortiz, Hendrickx & Winfield, 2017 comb. nov. Mysidium pumae Ortiz et al., 2017a: 114 –117, table 1, figs 1–4. Mysidium pumae – Ortiz et al. 2017b: 77. — Ortiz & Lalana 2018: 80. Material examined None. Type locality Pacific coast of Mexico, Mazatlán Harbor, 23.1814° N, 106.4242° W (Ortiz et al. 2017a). Revised definition Adapted to the scheme in Table 2 by using published data (Ortiz et al. 2017a, 2017 b) on adults of both sexes: all features diagnosed above for the genus Mysidium Dana, 1852, and its new subgenus Occimysidium. Cornea crescent-like in dorsal view, with ‘diameter’ 1.5 times as long as terminal segment of antennular trunk. Eyestalks smooth. Rostrum triangular, apically bluntly pointed, not extending beyond eyestalks. Appendix masculina with separate inner and distal tufts of setae; its length 2.0–2.2 times terminal segment of antennular trunk. Length of antennal scale six times maximum width; scale reaching well beyond antennular trunk. Median segment of mandibular palp with setae on both margins. Carpopropodus of first to eighth thoracic endopods 2, 2, 2, 3, 3, 3, 2, and two-segmented, respectively. Basal segment occupies 0.8 times total length of carpopropodus of endopod 3. Pleopods 1–2, 5 rodlike, not considering the reduced endopodal portion (pseudobranchial lobe); male pleopod 3 with an additional, small endite. Length of male pleopods increases in order of pleopods 1, (subequal 2, 3, 5), 4. Sympod of male pleopod 4 without endite. Exopod with basal segment occupying 54% total length. Endopod reduced to lobe with = 22% sympod length; apically with one long seta and more proximally four additional shorter setae. Uropodal endopod 0.7 times as long as exopod. Telson spatulate, caudally gradually narrowing; length 2.3 times maximum width near basis; terminal margin convex, evenly rounded. Proximal 51–52% of lateral margins smooth, distal portion of each margin with continuous series of 20–22 acute laminae. Terminal margin lined by 23–25 more densely set, acute laminae. These latter laminae more slender than the neighboring latero-terminal laminae. Distribution So far known only from type locality.Published as part of Wittmann, Karl J. & Wirtz, Peter, 2019, Revision of the amphiamerican genus Mysidium Dana, 1852 (Crustacea: Mysida: Mysidae), with descriptions of two new species and the establishment of two new subgenera, pp. 1-48 in European Journal of Taxonomy 495 on pages 25-26, DOI: 10.5852/ejt.2019.495, http://zenodo.org/record/258486
Data to "Effects of Active Exploration on Novelty-Related Declarative Memory Enhancement'
Memory data as reported in: Schomaker, J., & Wittmann, B. C. (2021). Effects of Active Exploration on Novelty-Related Declarative Memory Enhancement. Neurobiology of Learning and Memory, 107403
Parvimysis laminata Wittmann 2020, sp. nov.
Parvimysis laminata sp. nov. http://zoobank.org/ urn:lsid:zoobank.org:act: 9347D7BE-4D61-472B-8319-4777299A16EC (Figs 3–5) Parvimysis sp. B, Wittmann & Ariani, 2019: figs 1E, F, Table in Suppl. Type specimens. Holotype ♂ ad. with 2.7 mm body length (NHMW reg. no. 26570), paratypes 57 ♀ ad. 2.6–3.0 mm, 34 ♂ ad. 2.0– 2.6 mm, 14 subad. (AMNH IZC 331514, NHMW 26569, ZMBN 135401), Curaçao, Playa Lagun, 12°19’06’’N 69°09’07’’W, 5–6 m, in loose aggregation above sand, hand net, 28 Aug. 1997, leg. Peter Wirtz. Non-types from Curaçao (SW-coast, S = 36, leg. Karl J. Wittmann). 47 ♀ ad. 2.3–3.2 mm, 25 ♂ ad. 2.0– 3.1 mm, 5 subad. (NHMW 26574), Porto Mariebaai (Playa Portomarie), 12°13’06’’N 69°05’12’’W, sand flat and reef slope, 1–30 m, 24–26°C, swept with hand net from sand, also from small mysid swarms in rock recesses, 24 Feb. 2014, 15:00 local time; 33 ♀ ad. 2.9–3.3 mm, 17 ♂ ad. 2.3–2.7 mm, 4 subad. (NHMW 26572), Boca Sint Michiel, Sun Reef, reef slope, 12°08’21’’N 68°59’53’’W, 10–32 m, swept with hand net from brown algae and sand, 26°C, 16 Feb. 2014, about 08:20 local time; 17 ♀ ad. 2.7–2.9 mm, 17 ♂ ad. 1.9–2.7 mm, 7 subad. (NHMW 26573), Vaersebbaai = Kokomo Beach, reef slope, 12°09’38’’N 69°00’21’’W, swept with hand net from brown and red algae, also from mysid swarms over sand, 8–28 m, 24°C, 18 Feb. 2014, 14:30–15:20 local time. Non-type from Bonaire. 1 ♀ subad. 2.1 mm (NHMW 26571), Bachelor’s Beach, 12°07’34’’N 68°17’16’’W, 3–26 m, in pooled sample from the sea floor; small hand net, 7 June 2004, daytime, leg. Peter Wirtz. Type locality. Sublittoral marine waters of Curaçao, Playa Lagun, 12°19’06’’N 69°09’07’’W. Derivatio nominis. The species name is a Latin adjective with female ending, referring to the series of comparatively long and numerous laminae on the terminal margin of the telson. Diagnosis. Parvimysis with short, broadly rounded rostrum. Freely projecting portion of the rostrum 0.3–0.5 times the length of the terminal segment of the antennular trunk. Antero-lateral edges of the carapace produced into a short acute process. Cornea large, maximum diameter 1.8–2.4 times the length of the terminal segment of the antennular trunk in dorsal view. Antennular trunk extending beyond trunk of antennal flagellum. Antennal scale with basal segment extending beyond antennular trunk and to varying degree also beyond trunk of antennal flagellum. Cardiac portion of foregut with simple, smooth spines; additional stout, modified spines on lateralia, not so on dorso-lateral infolding. Mandibular palp with terminal segment making up 10–17% total palp length; median segment subterminally with two barbed setae; more proximal portions of this segment smooth or at most with one seta on inner margin, none on outer margin. Maxillary palp with proximal segment 0.3 times total palp length. Exopod of maxilla extends shortly beyond middle of the terminal segment of the palpus. Thoracic exopods 2–5 with 9-segmented flagellum, exopods 6, 7 with (9–8)-segmented flagellum. Endopods 1, 2 without claw; endopods 3–8 with long, slender claw. Length of claw 5 is 11–12 times its width at basis. Endopods 3–8 with oblique articulation between carpus and propodus. Carpus of endopod 5 with comb-like series of setae on inner margin in males, normal setation in females. Oostegites normal. Exopod of fourth male pleopod 3-segmented, ending in large modified seta plus a minute lobe bearing a minute seta. Apical segment of exopod is 0.4–0.5 times the length of the median segment. Scutellum paracaudale terminally well rounded or triangular with broadly rounded tip. Telson short, length 2.6–4.3 times distance between the latero-terminal spines. Shape roughly trapezoid, lateral margins slightly sinusoid, tapering, each armed with 2–5 short spines; each lateral margin ending in a short latero-terminal lobe with apical spine, the latter clearly longer than the lateral spines; margin between these lobes transversely straight or slightly concave, bearing 9–16 laminae with 0.4–0.8 times length of latero-apical spines; no medio-terminal teeth or spines. Description of types. All features of the diagnosis and those reported further above as common to the six Caribbean species of the present study. Size of adult females 2.6–3.0 mm (n = 54), males 2.0– 2.6 mm (n = 34). Cephalic region (Figs 3, 5 A–D). Cornea dorsoventrally not or only slightly compressed, calotte-shaped in dorsal, (sub)spherical to oviform in lateral view, apparent length 1.1–1.3 times height in lateral view (Fig. 3B). Basal segment of antennular trunk longer than combined median and apical segments in females, about equal or longer in males.Antennal scale with apical segment 22–30% length of the basal segment. Third segment of the trunk of the antennal flagellum with 70–82% length of the second segment. Mandibles as in Fig. 3G. Terminal segment of palp with 1–2 smooth setae, 4–5 modified setae bilaterally bearing series of stiff, spine-like barbs; and one longer seta with barbs along most of its distal 70%. Basal segment of maxillary palp with three barbed setae on inner margin, apical segment densely setose at tip and on terminal 50–70% of inner margin, but lined by small hairs in more proximal portions. Each side of the foregut (Fig. 5 A–D) with one large, apically pronged, serrated spine on mid-posterior part of lateralia; this spine with total of 11–18 teeth, among which 1–4 large teeth at tip (Fig. 5C). Group of 4 (3–5) centroapically serrated spines (Fig. 5D) in even more posterior position on lateralia; the largest, most proximal spine with total of 9–14 small teeth distributed over distal 50–70% spine length. Thorax (Figs 4, 5E). Length increases from exopod 1 to 5 and decreases from 5 to 8; length of endopods increases from 1 to 8. Basal plate of thoracic exopods 1–8 terminally well rounded (Figs 4A, 5E). Flagellum of exopod 1 with 8 segments; flagellum of exopod 8 with 8 segments in females, and 8–9 in males. Thoracic endopod 8 slender; when stretched, extending forwards shortly beyond the eyes or backwards to or shortly beyond the telson. Dactylus of endopod 2 large, equipped with 4–9 modified setae (Fig. 4B) plus a few smooth setae, no spine-like setae. Length of claws increasing in series of thoracic endopods 3–5, not clearly increasing in more caudal endopods. Slenderness not clearly differing among endopods 3–8: claw 3 is 10–13, claw 8 is 10–12 times its width at basis. Penes (Fig. 5E) terminally with 5–6 minute setae facing the ejaculatory opening. Pleon (Fig. 5 F–R). Male pleopod 5 slender, 1.7–2.0 times length of pleopod 3. Male pleopod 4, when stretched, reaching to end of pleonite 5, its apical seta ends at 80–100% length of pleonite 6. Fourth endopod 0.7–0.9 times length of basal segment of exopod; endopod with 8–10 barbed setae. Uropods with endopod 0.9 times length of exopod or 1.6–1.7 times length of telson (without spines). Length of exopod 6–7 times maximum width. Statoliths mineralized with vaterite, a metastable polymorph of calcium carbonate; shape elliptical to spherical in dorsal view; diameter 60–120 µm (n = 40). Statolith formula 2 + 3 + (4–6) + (5–6) = 14–17. Scanning electron micrographs of a statolith available as Parvimysis sp. B in Wittmann & Ariani (2019: figs 1E, F). Telson length 1.2–1.5 times its width at basis, or 0.5–0.6 times length of exopod of uropods, or 0.6–0.8 times pleonite 6. Laminae with 0.6–1.2 times median length of lateral spines. Gut contents. Most stomachs empty. Five full foreguts contained mineral particles (‘sand’), fragments of detritus, macerated, unidentifiable material, and diatoms. Content of midgut and hindgut contained mineral particles and particulate to fully macerated, unidentifiable material. The contents suggest that the mysids graze ‘detritus’ on the sediment surface. They also indicate that the secondary filter does not fully prevent infiltration of mineral particles into the midgut. Eggs and larvae. Among 150 adult females examined in this respect, 65 showed empty brood pouch, 38 were with eggs, 30 with nauplioid larvae, and 17 with postnauplioid larvae. Size of breeding females 2.4–3.3 mm, each carrying 3–4 (5) eggs or larvae. Egg diameters 0.23–0.32 mm, length of nauplioids 0.5–0.6 mm at substage N2, 0.5–0.7 mm at N3, 0.6–0.8 mm at N4, postnauplioids 0.7–0.9 mm at P2, 0.8–1.0 mm at P3 (n = 50 eggs; 39, 37, 34, 22, or 35 larvae, respectively). The larvae attain 23–35% parental length (n = 10 parents) shortly before they moult to the free-living juvenile stage. Distribution and habitat. Coasts of Curaçao and Bonaire, both islands located in the south-western part of the Lesser Antilles. In euhaline waters, on the sea floor in 1–32 m depth. Observed in swarms and loose aggregations hovering a few cm above the substrate (sand, algae, overgrown rock), over and inside rock recesses. Without direct observation, swept with hand net from the substrate surface during the day.Published as part of Wittmann, Karl J., 2020, Revision of the genus Parvimysis (Mysida, Mysidae) with descriptions of five new species from coastal waters of the Caribbean, pp. 1-30 in Zootaxa 4742 (1) on pages 10-14, DOI: 10.11646/zootaxa.4742.1.1, http://zenodo.org/record/367435
Mysidium (Orientomysidium) Wittmann & Wirtz, 2019, subgen. nov.
Orientomysidium Wittmann subgen. nov. urn:lsid:zoobank.org:act: E269F7A6-0187-4AD0-9D64-24C700D5126D Type species Diamysis columbiae Zimmer, 1915, by present designation. This taxon acknowledged in revised combination as Mysidium (Orientomysidium) columbiae (Zimmer 1915). Etymology Noun in nominative singular with neutral ending, formed by amalgamation of the Latin adjective ‘ orientale ’ with the generic name Mysidium, referring to the occurrence on the east coast of America. Diagnosis Subdivision of the genus Mysidium Dana, 1852, characterized by three-segmented carpopropodus of third thoracic endopod; merus of this endopod with serrated setae. Third male pleopod with strong endite at about 50–60% distance from basis of medial margin. Sympod of fourth male pleopod with series of setae on rostral face. Its exopod three-segmented; proximal segment without seta; the two distal segments each with one unbranched modified seta. Telson with terminal cleft. Distribution Littoral and sublittoral waters at Atlantic coasts of tropical America (24° N to 8° S), including Gulf of Mexico and Caribbean. Mysidium (Orientomysidium) columbiae (Zimmer, 1915) subgen. et comb. nov. Figs 9, 13E–J Diamysis columbiae Zimmer, 1915a: 172 –174, figs 23–29 (senior synonym in outdated generic combination). Diamysis columbiae – Zimmer 1915b: 215 (in synonymy). — Illig 1930: 599 (in synonymy). — Prieto et al. 2009 (in synonymy). Mysidia columbiae – Zimmer 1915b: 215, fig. 18 (combination with generic junior homonym); 1918: 26. — Illig 1930: 599 (in key). — W.M. Tattersall 1951: 222 (in homonymy). Mysidium columbiae – W.M. Tattersall 1951: 222, 223 (revised combination). — Davis 1966: figs 1–5. — Brattegard 1969: 86, fig. 27B; 1974a: 69; 1974b: 102, table 1; 1975: 112. — Mauchline 1972: table 3; 1973: tables 2, 4; 1980: 338 (in list). — Băcescu & Ortiz 1984: 22 (in key). — Harrison & Bowman 1987: 678 (in comparison). — Modlin 1987: 115; 1990: figs 1, 3; 1993: 25. — Ortiz & Lalana 1988: 14 (in list); 2018: fig. 10A. — Zoppi de Roa et al. 1989: fig. 2B. — Markham et al. 1990: 411. — Băcescu 1991: 3, fig. 1J. — Escobar-Briones & Soto 1991: table 1. — Ariani et al. 1993: table 1. — Ortiz 2001: 100 (in list). — Price et al. 2002: 44. — Price & Heard 2004: 154. — Sorbe et al. 2007: table I. — Prieto et al. 2009: 33. — Miyashita & Calliari 2014: 9. — Wittmann et al. 2014: 350. — Ortiz et al. 2017a: table 1; 2017b: fig. 3A. — Esteves-Astudillo et al. 2017: 62. Mysidium colombiae – Harrison & Bowman 1987: 674, 678 (invalid spelling variant). — Price & Heard 2004: fig. 3I; 2009: 938 (in list). non Mysidium columbiae – Brattegard 1969: fig. 27E. Type material examined The three vials are from the same sample labelled “Cartagena Kolumbien G. Gagzo leg. 18. IX. 1905 ded. 7. XI. 1906 ”, this material is brittle and delicate, limiting examination. The two specimens from vial 14074 with body parts and appendages sticking together. The respective missing appendages suggest that the lectotype has been partly dissected and figured by Zimmer (1915a: figs 23–29). Lectotype (by present designation) COLOMBIA • ♂ ad. bl 4.4 mm; registered at the ZMH as “ Diamysis columbiae Zimmer, lectotype by present designation”; 10.37° N, 75.52° W; ZMH K-14073a Paralectotypes COLOMBIA • 1 ♀ ad. bl 4.8 mm carrying two postnauplioid larvae; ZMH K-14073b • 1 ♂ ad. bl 4.6 mm, 1 ♀ ad. bl 5.0 mm; ZMH K-14074. Other material examined CURAÇAO • 26 ♀♀ ad bl. 4.4–7.0 mm, 19 ♂♂ ad. bl 3.3–7.4 mm, 11 subad.; #C3; MINGA MYS 431 • 13 ♀♀ ad. bl 5.9–7.0 mm, 7 ♂♂ ad. bl 5.8–7.8 mm, 8 subad.; #C5; NHMW 26479. BONAIRE • 24 ♀♀ ad. bl 4.6–5.3 mm, 22 ♂♂ ad. bl 3.9–6.9 mm, 61 subad.; #B11; ZMH K-55263 • 13 ♀♀ ad. bl 5.0– 7.9 mm, 26 ♂♂ ad. bl 5.7–6.5 mm, 7 subad.; #B12; NHMW 26480. Type locality Cartagena at the Caribbean coast of Colombia (Zimmer 1915a). Coordinates estimated by present authors are about 10.37° N, 75.52° W. Revised definition All features diagnosed above for the genus Mysidium Dana, 1852, and its new subgenus Orientomysidium. Cornea globose in lateral view; calotte-shaped in dorsal view, with diameter 1.5–2.0 times as long as terminal segment of antennular trunk. Eyestalks smooth. Rostrum triangular, with concave lateral margins, apically bluntly to acutely pointed, not or weakly extending beyond basis of eyestalks. Anterolateral edges of carapace rounded. Only males with anterior margin of antennular trunk dorsally with rounded, shield-like, mediodistal extension (Fig. 9A–B) bearing 2–4 setae. The largest seta 0.1–0.2 times extension length. Appendix masculina bilobate, 2.0–3.3 times as long as terminal segment of antennular trunk; all along with brush of long setae. Length of antennal scale 7–8 times maximum width; scale reaching far beyond antennular trunk. Median segment of mandibular palp with setae on both margins. Distally angular hump on outer face of terminal segment of the maxillula. Carpopropodus of thoracic endopods 1–8 with 2, 2, 3, 3–2, 2–3, 2–3, 2–3 or 2–3 segments, respectively. Series of 2–6 serrated setae (Fig. 9D) along merus of third thoracic endopod. Basal segment occupies half total length of carpopropodus of endopod 3. Pleopod 1 rod-like in both sexes, not considering the weak (sub) median widening at its vestigial endopodal portion (Fig. 9F). Sympod of male pleopod 4 with strong endite at 2/5 sympod length from basis and with longitudinal series of 3–6 plumose setae on rostral face. Exopod with basal segment occupying 64–72% total length. Endopod reduced to lobe with 27–48% sympod length; with 7–10 barbed setae of various sizes. Endopod of uropods 0.6–0.7 times as long as exopod. Telson subrectangular, length 1.4–1.8 times maximum width near basis. Its lateral margins convex along proximal third; proximal 61–73% of lateral margins smooth, distal portion of each margin with continuous series of 8–12 acute spines. Cleft penetrating 13–15% telson length; cleft separating two broadly rounded, apical lobes; cleft and terminal portion of lobes densely lined in continuous series by a total of 22–30 stout, blunt laminae. Description of type (Fig. 9A) All available features fit within the diagnosis. Antennular trunk extends 25–47% its length beyond (artificially aligned) eyes. Sympod of antenna produced into spiniform extension on outer distal corner. Apical segment of maxillary palp about two times as long as basal segment. Carapace posteriorly emarginate, with well-rounded latero-terminal lobes; cervical sulcus strong. Dactylus of thoracic endopods 1–2 with moderately long, strong claw; dactylus 3–7 with long, needle-like claw; dactylus 8 with short seta-like claw. Sympod of fourth male pleopod with 3–4 plumose setae (this range wider in non-types examined) on rostral face plus 1–2 small, barbed setae on inner margin. Pleonites 1–5 are 0.5–0.8, 0.5–0.8, 0.6–0.8, 0.6–0.8 or 0.8–1.1 times the length of pleonite 6, respectively. Scutellum paracaudale triangular with blunt to acute apex; both margins weakly concave to weakly convex. Uropodal endopod 0.9–1.1 times, exopod 1.2–1.5 times as long as pleonite 6. Exopod extends 0.3–0.4 times its length beyond endopod, or 0.7–0.8 times beyond telson; endopod 0.4–0.6 times its length beyond telson. Telson with 0.5–0.7 times as long as exopod of uropods, 0.5–0.9 times endopod or 0.6–0.9 times pleonite 6. Notes on additional material (Figs 9B–J, 13E–J) Adult females of present material with 4.4–7.9 mm (n = 76) body length, males 3.3–7.8 mm (n = 74). Numbers of serrated setae (Fig. 9C–D) along the merus of thoracic endopod 3 increase with increasing body size; also numbers of endopods with any setae of that type increase with body size, i.e., pertaining only to thoracopod 3 in smaller individuals and successively to thoracopods 4–7 as individuals become larger. Endopod 8 (when stretched) reaching backwards at most to the end of pleonite 4 and forwards to mandibles; its carpopropodus measures 0.4–0.5 times telson length. Male pleopod 4 comparatively short; exopod reaching at most to basal third of pleonite 6, its subapical seta up to terminal third. Endopod of male pleopod 4 (Fig. 9H) with the most distal seta 1.4–4.4 times endopod length. Sympod with field of scales on its endite. Statoliths composed of fluorite. Foregut (Fig. 13E–G) closely similar to that of M. antillarum sp. nov. (Fig. 11A–E). As main differences from this species, M. columbiae shows different structure and variable numbers of large spines: on each lateral half there are 2–4 apically pronged spines on posterior part of lateralia, the teeth of these spines with dense sets of secondary denticles (Fig. 13F); and there are 1–2 larger, apically pronged, serrated spines on dorso-lateral infolding, the latter spines flatter, more clavate, and with more slender teeth on basal ¾ (Fig. 13G). Notes on individual development Nauplioid larvae (Fig. 13H) with smooth cuticle all around, except for a pair of minute furcal processes and a number of minute setae on the blunt end of the abdomen (Fig. 13J). Davis (1966: fig. 4) already reported apical, furcal ‘spines’ in nauplioids of M. columbiae from Jamaica. In non-types of the present material, the differentiation of male sexual characteristics is visible in early subadults as small rudiments of penis and appendix masculina and of a bifid fourth pleopod. At this stage the exopod of pleopod 4 is less than 3/2 as long as endopod and the endite of the sympod is missing or indistinct. In the course of further development a distinct endite (representing an important diagnostic character) becomes visible as soon as the exopod exceeds two times the length of the endopod. Distribution and habitat From 24° N to 8° S in coastal waters of Bahamas, Caribbean, southern Gulf of Mexico, and Brazil (Price & Heard 2004; Miyashita & Calliari 2014). The present samples from Curaçao and Bonaire fit within this geographic range. The present material was encountered at a depth of 3–26 m, during daytime in swarms hovering around and between corals. During the night dispersed over the sea floor.Published as part of Wittmann, Karl J. & Wirtz, Peter, 2019, Revision of the amphiamerican genus Mysidium Dana, 1852 (Crustacea: Mysida: Mysidae), with descriptions of two new species and the establishment of two new subgenera, pp. 1-48 in European Journal of Taxonomy 495 on pages 26-30, DOI: 10.5852/ejt.2019.495, http://zenodo.org/record/258486
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