160 research outputs found
A new canopy-dwelling species of Dendrelaphis (Serpentes: Colubridae) from Sinharaja, World Heritage Site, Sri Lanka
No full textMendis Wickramasinghe, L. J. (2016): A new canopy-dwelling species of Dendrelaphis (Serpentes: Colubridae) from Sinharaja, World Heritage Site, Sri Lanka. Zootaxa 4162 (3): 504-518, DOI: http://doi.org/10.11646/zootaxa.4162.3.
A new species of Cnemaspis (Sauria: Gekkonidae) from Sri Lanka
No full textMendis Wickramasinghe, L. J. (2006): A new species of Cnemaspis (Sauria: Gekkonidae) from Sri Lanka. Zootaxa 1369: 19-33, DOI: 10.5281/zenodo.17487
A new species of Dryocalamus (Serpentes: Colubridae) endemic to the rainforests of southwestern Sri Lanka
No full textVidanapathirana, L.J. Mendis Wickramasinghe Dulan Ranga, Pushpamal, Vishan, Wickramasinghe, Nethu (2020): A new species of Dryocalamus (Serpentes: Colubridae) endemic to the rainforests of southwestern Sri Lanka. Zootaxa 4748 (2): 248-260, DOI: https://doi.org/10.11646/zootaxa.4748.2.
FIGURE 4 in A new species of Cnemaspis (Sauria: Gekkonidae) from Sri Lanka
No full textFIGURE 4. Right limb of Holotype male of C. ranwellai sp.nov (NMSL2004.1.1)Published as part of Mendis Wickramasinghe, L. J., 2006, A new species of Cnemaspis (Sauria: Gekkonidae) from Sri Lanka, pp. 19-33 in Zootaxa 1369 on page 26, DOI: 10.5281/zenodo.17487
FIGURE 11 in A new canopy-dwelling species of Dendrelaphis (Serpentes: Colubridae) from Sinharaja, World Heritage Site, Sri Lanka
No full textFIGURE 11. Full body of Dendrelaphis caudolineolatus, holotype (BMNH 68.2.17.19).Published as part of Mendis Wickramasinghe, L. J., 2016, A new canopy-dwelling species of Dendrelaphis (Serpentes: Colubridae) from Sinharaja, World Heritage Site, Sri Lanka, pp. 504-518 in Zootaxa 4162 (3) on page 515, DOI: 10.11646/zootaxa.4162.3.5, http://zenodo.org/record/26452
Aspidura desilvai Mendis Wickramasinghe & Bandara & Vidanapathirana & Wickramasinghe 2019, sp. nov.
No full textAspidura desilvai sp. nov. (Figures 1–7) Holotype. NMSL-NH 2019.01.0 2, adult male, 168 mm SVL (Figure 2), from Riverstone, Knuckles, Matale District, Central Province, Sri Lanka (07°31’39” N, 80°44’01” E, elevation 1420 m). Collected by L.J.M. W and D.R.V. on 0 7 July 2018. Paratypes. NMSL-NH 2019.01.0 1, adult female, 208 mm SVL, from Panwila in Knuckles Mountain Range, Kandy District, Central Province in Sri Lanka (07°22'00.36’’ N, 080°41'00.10’’ E, elevation 995 m). Collected by L.J.M. W and I.N.B. on 13 March 2011; DWC 2019.05.0 1, adult female, 157 mm SVL, from Dotulugala, Knuckles Mountain Range, Kandy District, Central Province, Sri Lanka (07°27'00.30” N, 080°45'00.20” E, elevation 1700 m). Collected by L.J.M. W and I.N.B. on 17 March 2011; DWC 2019.05.0 2, juvenile male, 93 mm SVL, from Gombaniya Mountain, Knuckles Mountain Range, Matale District, Central Province, Sri Lanka (07°27'51.76’’ N, 080°45'51.79’’ E, elevation 1375 m). Collected by L.J.M. W and I.N.B. on 13 March 2011. Diagnosis. SVL 94–216 mm; snout to eye distance 2.5 times the eye width (SE/EW); prefrontals touching eye; preocular small, does not touch supraocular; postoculars 2, lower one larger than the upper; temporal 1+2/1+2; supralabials 6/6, 4 th touching eye; infralabials 6/6, first pair in contact, progressively increasing in size from 1 st to 6 th; anterior chin shields 2, large, touching 1–4 infralabials; posterior chin shields 2, anterior half in contact while the posterior half separated by 1 st ventral; ventrals 124–139; subcaudals 16–29; dorsal scale rows 15–15–15; laterally spine like tubercles present on two scale rows nearest to the subcaudals of the ischiadic, anal and tail base regions in adult males, feeble in juvenile males, and absent in females; entire dorsum brown colour, much paler towards anterior; three irregular dotted lines on dorsum. Description of holotype. Adult male; SVL 168 mm; TaL 25.1 mm; TL 193.1 mm; TaL/TL 0.13; body elongate and cylindrical; head short (SVL/HL 18.3), elliptical, indistinct from thick neck; snout long, narrowing anteriorly, pointed in dorsal aspect, snout to nostril distance about 2.8 (EW/SN) times as long as nostril width; nasal divided; small, triangular nostril, touching divided nasal and first supralabial, not touching rostral; eye larger than horizontal diameter of nostril, distance between snout to eye about 2.6 (SE/EW) times the eye width, round pupil; snout to eye distance 0.3 times head length (SE/HL); tail short (TaL/SVL 0.1), robust at its base, tapering progressively to a single point. Head scalation. Head scalation includes 1 internasal, 2 prefrontals, 2 supraoculars, 1 frontal, and 2 parietals (Figure 3A). Rostral small, convex, wider than long and rounded in lateral, dorsal and ventral aspects. Nasal vertically divided by a groove above nostril (Figure 3B). Internasal large, irregular hexagonal; widely in contact with prefrontals. Two large prefrontals, longer and wider than internasals, largest distance along the longitudinal axis of prefrontals shorter than frontal (Figure 3A) in length, anterior-most corner of prefrontals touching nasal, bordered by 2 nd and 3 rd supralabial, preocular scale, eye, supraocular and frontal. Preocular small, not in contact with supraocular. Loreal and subocular scales absent. Supraocular smaller than frontal. Two postoculars, lower one larger than upper. Two parietals; largest scales on head. Temporals 1+2/1+2. Supralabials 6/6, 4 th touching eye, progressively increasing in size from 1 st to 6 th (Figure 3B). Mental small and triangular, wider than long. Infralabials 6/6, first pair in contact, progressively increasing in size from 1 st to 6 th. Anterior chin shields 2, large, touching 1–4 infralabials. Posterior chin shields 2, anterior half in contact, posterior portion separated by 1 st ventral (Figure 3C). Body scalation. Ventrals 124, 1 st ventral longer than wide; subcaudals 24, all single; anal single and large; dorsal scale rows 15–15–15; laterally prominent spine like tubercles present on two scale rows nearest to the subcaudals, and its protrusion reducing towards upper scale rows in the ischiadic, anal and tail base regions (Figure 4); vertebral rows and first coastal not enlarged; no apical pit. Hemipenis morphology. Based on Holotype specimen: right everted hemipenis extends for length of 3 subcaudals. Everted organ single subcylindrical, globular, sulcus spermaticus simple. Basal to apex region bearing prominent spines which are evenly distributed and are in uniform length (Figure 5). Colour in life. Supralabials and infralabials light yellow, with dark margins separating each scale (Figure 6A). Entire dorsum reddish brown colour, much paler towards anterior and each scale having tiny dark spots (Figure 2). Three irregular dotted lines on dorsum (Figure 6B). These are symmetrically placed and continues from neck to tail end. Prominent light brown stripe continues dorsolaterally from neck to tail end, marked due to much darker regions which constitutes of dotted lines below and above this region. These lines continue from neck to tail end. Venter primarily peach, with black blotching all over; gular region yellow. Colour in alcohol. Colour pattern remains unchanged. Pupil changes to off white. Darker regions fades to a light brown. Variations in colour. In an unpreserved male specimen (Figure 7D) except the head region and ventre the entire body was black. Natural History. Aspidura desilvai sp. nov. have been observed commonly in its habitat (Figure 1). The species is confined to Knuckles conservation area, and is found in and above the lower montane forests of Knuckles. Authors have observed the snake from 995 m up to 1700 m above sea level (Figure 8). The habitat of A. desilvai sp. nov. is closed canopy forests dominated by Syzigium sp. (Figure 9). The moist-cooler habitat is densely occupied with large and medium sized trees which are heavily covered with epiphytes. No direct sunlight falls to the forest floor, and the undergrowth was not well established where the individuals were found. Relatively thin litter cover was observed in the habitat. Commonly observed under leaf litter and loose soil while they were also observed under rocks, boulders, and decaying logs. Individuals come out to the surface during the day time. Reddish brown latosolic soil in the locality is more or less similar to the body colour of the snake. Etymology. The species is named in honor of Pilippu Hewa Don Hemasiri de Silva (Dr. P. H. D. H. de Silva), a former Director (1965-1981) of the National Museums of Sri Lanka. In recognition of his tireless services to the country, while in service and through his many publications specially as the author of the book titled “ Snake Fauna of Sri Lanka, with special reference to skull, dentition and venom in snakes ”. The species epithet desilvai is a noun in the genitive case. Suggested common names. desilvage madilla, and de Silva’s Rough-Side Snake in native Sinhala language and English language respectively. Comparison. The new species was compared with all known congeners of the genus Aspidura and the species most closely resembles A. ravanai, and A. trachyprocta, due to the following combination of characters: one preocular, two postoculars, 1+2 temporals, supralabials 6, 4 th supralabial in contact with the eye, infralabials 6, coastals 15, single cloacal scale, and overlapping ventral and subcaudal counts, but can easily be distinguished by the following morphological characters: from A. ravanai: entire dorsum brown colour, much paler towards anterior and each scale having tiny dark spots in Aspidura desilvai sp. nov. (vs. entire dorsum jet black in Aspidura ravanai), ventrolaterally darker region which constitutes of irregular longitudinal dotted lines (vs. ventrolaterally an irregular longitudinal yellow stripe), laterally prominent spine like tubercles present on two scale rows nearest to the subcaudals, and its protrusion reducing towards upper scale rows (vs. entire coastal rows coarsely keeled, with 1–3 peaks on each scale) in males (Figures 4 & 10 A–C), entire coastal rows of the ischiadic, anal and tail base regions smooth (vs. feebly keeled) in females, snout to eye distance about 2.5 times its eye width (vs. 3.2 times in A. ravanai) (Figures 3 & 10 D–E); from Aspidura trachyprocta: entire dorsum brown colour, much paler towards anterior and each scale having tiny dark spots in A. desilvai sp. nov. (vs. reddish-yellow to brown with a longitudinal black stripe on mid dorsum in Aspidura trachyprocta), ventrolaterally darker region which constitutes of irregular longitudinal dotted lines (vs. black stripe), laterally prominent spine like tubercles present on two scale rows nearest to the subcaudals, and its protrusion reducing towards upper scale rows (vs. bulging spine like tubercles prominent laterally which reduces towards dorsum) of the ischiadic, anal and tail base regions in males (Figures 4 & 11 A–C), entire coastal rows of the ischiadic, anal and tail base regions smooth (vs. feebly keeled) in females, snout to eye distance about 2.5 times its eye width (vs. twice in A. trachyprocta) (Figures 3 & 11 D–E); from A. brachyorrhos Boie, 1827, by having 15 coastals (vs. 17), preocular not in contact with supraocular (vs. contact), prefrontal contact with eye (vs. separate), single subcaudals (vs. paired); from A. copei Günther, 1864 by having coastals 15 (vs. 17), single subcaudals (vs. paired), single preocular (vs. absent); from A. deraniyagalae Gans & Fetcho, 1982 by having 15 coastals (vs. 17), ventrals 124–139 (vs. 117–122), single subcaudals (vs. paired); from A. drummondhayi Boulenger, 1904, by having single subcaudals (vs. paired), single preocular (vs. absent); from A. guentheri Ferguson, 1876 by having 15 coastals (vs. 17), ventrals 124–139 (vs. 100–127); from A. ceylonensis (Günther, 1858), by prefrontal touching eye (vs. not touching eye), preocular does not touch supraocular (vs. touches), lower postocular larger than the upper (vs. vise versa), mid body coastals not keeled (vs. coarsely keeled).Published as part of Mendis Wickramasinghe, L. J., Bandara, Imesh Nuwan, Vidanapathirana, Dulan Ranga & Wickramasinghe, Nethu, 2019, A new species of Aspidura Wagler, 1830 (Squamata: Colubridae: Natricinae) from Knuckles, World Heritage Site, Sri Lanka, pp. 265-280 in Zootaxa 4559 (2) on pages 266-272, DOI: 10.11646/zootaxa.4559.2.3, http://zenodo.org/record/262697
Cnemaspis retigalensis Mendis Wickramasinghe & Munindradasa 2007, sp. nov.
No full text<i>Cnemaspis retigalensis</i> sp. nov. <p> <b>Holotype</b>. NMSL 20061201, Adult male, 28.76 mm SVL, from Weweltenna, Retigala, Sri Lanka, (N 08º 06’ 40.3” E 080º 39’ 31.4”, elevation 710m), 30.10.2005, collected by L. J. Mendis Wickramasinghe and D. A. I. Munindradasa.</p> <p> <b>Paratypes</b>. NMSL 20061202, Adult female, 30.87 mm SVL; NMSL 20061203, Adult female, 26.56 mm; NMSL 20061204, Adult male 27.67 mm SVL. Date of Collection 27.08.2006, the same locality and collected by L. J. Mendis Wickramasinghe and Roshan Rodrigo.</p> <p> <b>Diagnosis.</b> A small-sized <i>Cnemaspis</i> (snout to vent length 26–31 mm in an adult males), which can be distinguished from all known congeners by the following combination of characters: postmentals separated by a small scale; nostrils are not in contact with first supralabial; six supra labials to angle of mid-orbit position and end of jaw at 7–8 supra labials; 30–32 interorbitals; throat scales smooth; dorsal tubercles 62–65; dorsal tubercles small, rounded, pentagonal or hexagonal; absence of groups of carinated large scales in dorsal body; presence of conical tubercles, larger than dorsal body scales on the lower part of flank; spine-like tubercles absent on flanks; scales on the thigh intermixed with the tricarinated form; gular scales smooth; midventrals 26–27; ventral smooth and imbricate; subcaudals slightly large; preanal pores absent; 3–4 femoral pores on each side; 11 subdigital lamellae and 3 basal lamellae in the 4 th finger; 11–12 subdigital lamellae and 6 basal lamellae in the 4 th toe.</p> <p> <b>Description of Holotype.</b> Adult male (figs. 5, 13B, 17B, 21B, 25B and 29B) snout to vent length 28.76 mm, head depressed and narrow (HD / HLJ 0.38), head elongated and large (HLJ / SVL 0.29), distinct from the neck. Snout long (SE / HW 0.76), longer than the eye width (EW / SE 0.43). Eye relatively large (EW / HLJ 0.20). Ear opening small (EL / HLJ 0.11), inter ear distance is greater than the width of the eye (EE / EW 2.87).</p> <p>Rostral is large with a groove penetrating 3/4 of the scale. There are three internasals, with the mid scale being large in size to the nostril, and the other two are larger. The supranasal and postnasal consist of one smooth circular scale each and are bigger than the nostril, but equal or smaller than internasal and larger than the mid one. The head is covered with elongated, round, pentagonal or hexagonal shaped tubercle scales from snout to posterior margin of interorbital area and with small granulated scales up to the neck. The size of tubercle scales becomes progressively smaller from the snout to interorbital area. However, a group of large scales (still smaller than that on the snout) is located on upper interorbital area, and a set of very small scales are located in the parietal area. There are 30 interorbital scales of which mid scales are shorter and smaller than that of outer. The supraciliaries are slightly larger than upper interorbital scales. The nostril is oval, and is not connected with the supralabials. The nostril and the first supralabial are separated by a postnasal. The loreal region is convex and is covered with 15 large, circular and elongated, smooth tubercle scales. There are seven supralabials at the base of the jaw, with six at the mid orbit point. The first supralabial is equal or small to the second and third. The rest becomes progressively small. The dorsal tubercles are smaller than the upper interorbitals and are rounded, pentagonal or hexagonal in shape, and all are of similar size. There are 62 dorsal tubercles at the mid region of the body. The spine-like tubercles are absent on flanks. The conical tubercles present on the lower part of flank are larger than dorsal body scales and the subconical scales present on the upper part of flank are slightly smaller than the above. The dorsal part of forelimb and hind limb is covered with a flushed and juxtaposed, comparatively large scales with a keel. The scales on the thigh are intermixed with the tricarinated form. The tail is covered with scales larger than the dorsal body and the ventrolateral margin possesses rounded tubercles larger than tail scales. The mental scale is large and sub-triangular. A pair of rounded and pentagonal or hexagonal postmentals (smaller than the mental) is present on either side. The first postmental pair is separated by a small scale, and is connected with the first infralabial. The second postmental pair is smaller, and is connected with the first and second infralabials. There are seven infralabials towards the jaw end, with six of them towards the mid orbit point. The infralabials become progressively smaller in size towards the anterior end. The ear holes are oval shaped, bigger than nostrils, but smaller than eyes. There are 24–26 scales between the eye and ear. The scales in the throat are smooth, rounded, pentagonal or hexagonal in shape, the anterior scales being larger than the posterior scales. The gular scales are smooth. The mid ventral area consists of 26 scales, which are smooth, imbricate and smaller than the postmentals. The scales in ventral portion of fore and hind limbs are smooth, with the scales in the hind limb being relatively larger than those of the forelimb. There are four femoral pores and no preanal pores present. The preanal is smaller than anal scales. There are 70 subcaudals. The mid subcaudals are slightly larger than the other scales in the tail or equal in size. Although the mid subcaudals are circular or overlapping diamond in shape, the lower border appears to be slightly elongated-diamond in shape. This feature becomes prominent towards the end of tail. The keels are absent in subcaudals. The digits are slender, elongated and clawed. The distal sub-digital formulae include 4>3>2>5>1 (fingers) and 4>3>5>2>1 (toes) (Fig.25.B.).</p> <p> <b>Colour in life.</b> The body colour in the dorsal side is light brown. There is a faded black transverse band on either upper interorbital area. A closed contour of black comprised of boundaries of internasal, loreal, upper interorbital and parietal areas and a ‘W’ shaped marking (with a light yellow patch in posterior ‘W’) on anterior neck is on the dorsal head. A black patch is present on the posterior neck. The supraciliaries are light yellow. The eye pupil is circular and black with the surrounding being luminous red. The lateral view of the head and neck consists of three black line segments (one from nasal to mid eye in loreal region, the other along lower parietal boundary – both dorsally seen as part of the closed contour and the third from back of eye to neck on temporal region) in a brownish yellow background with yellow spots in supralabial, lower jaw and lower neck areas. The ventral view of the throat is light grey with irregular yellow markings in ventral jaw. Three faded stripes are present on each lower and upper arm in a brownish yellow background. The black stripe formula of 2,3,3,4 and 3 is present on fingers in a brownish yellow background. The ventral view of lower and upper arm is light brown. The mid dorsal area of the body is light brown, with four faded ‘W’ marks between fore and hind limbs. The black spots in upper flank and yellow and back spots in lower flank are present in mid lateral view. The mid ventral view is light grey with yellow irregular markings on ventrolateral margin of mid body. Three faded stripe are present on each femur and tibia in a brownish yellow background. The black stripe formula of 2,3,4,5 and 4 is present on toes in a brownish yellow background. The ventral femur and tibia are light grey in colour with yellow scale boundaries. The original part of the tail is light grey, with 13 transverse marks of faded black, of which the mark at the base and the next are ‘W’ shaped, and next two are hourglass shaped, and the rest is straight. The ventral aspect of tail is grey.</p> <p> <b>Colour in alcohol.</b> All yellow in life is turned to white while the rest is conserved.</p> <p> <b>Etymology.</b> The species epithet <i>retigalensis</i> is derived from Latin for “Retigala” referring to the forest where the species nov. is discovered. The vernacular names assigned for the species nov. are <i>Retigala diva huna, Retigala pahal palli</i> and <i>Retigala day gecko</i> in native languages Sinhala, Tamil and in English respectively.</p> <p> <b>Remarks.</b> <i>C. retigalensis</i> sp. nov. is congener with <i>C. kandiana</i> and <i>C. kumarasinghei</i> sp. nov. from morphological characters. However, <i>C. retigalensis</i> can easily be distinguished from <i>C. kandiana</i> by the absence of preanal pores and having smooth gula scales and, from <i>C. kumarasinghei</i> by absence of preanal pores and presence of scales on the thigh intermixed with the tricarinated form, and also from morphometric analysis. Specimens with yellow vertebra line are found rarely. <i>C. retigalensis</i> is often found in hill tops, on the lower 2m of trees and rocks.</p>Published as part of <i>Mendis Wickramasinghe, L. J. & Munindradasa, D. A. I., 2007, Review of the genus Cnemaspis Strauch, 1887 (Sauria: Gekkonidae) in Sri Lanka with the description of five new species, pp. 1-63 in Zootaxa 1490 (1)</i> on pages 10-12, DOI: 10.11646/zootaxa.1490.1.1, <a href="http://zenodo.org/record/5087387">http://zenodo.org/record/5087387</a>
FIGURE 3 in A new species of Dryocalamus (Serpentes: Colubridae) endemic to the rainforests of southwestern Sri Lanka
No full textFIGURE 3. Dryocalamus chithrasekarai sp. nov., holotype, NMSL-NH 2019.26.01, 328 mm SVL, head scalation: (A) lateral, (B) dorsal, (C) ventral aspects, respectively.Published as part of Vidanapathirana, L.J. Mendis Wickramasinghe Dulan Ranga, Pushpamal, Vishan & Wickramasinghe, Nethu, 2020, A new species of Dryocalamus (Serpentes: Colubridae) endemic to the rainforests of southwestern Sri Lanka, pp. 248-260 in Zootaxa 4748 (2) on page 252, DOI: 10.11646/zootaxa.4748.2.2, http://zenodo.org/record/369875
FIGURE 6 in A new species of Cnemaspis (Sauria: Gekkonidae) from Sri Lanka
No full textFIGURE 6. Ventral view femoral pores of Cnemaspis ranwellai sp. nov. (NMSL2004.1.1)Published as part of Mendis Wickramasinghe, L. J., 2006, A new species of Cnemaspis (Sauria: Gekkonidae) from Sri Lanka, pp. 19-33 in Zootaxa 1369 on page 28, DOI: 10.5281/zenodo.17487
FIGURE 1 in A new species of Rhinophis Hemprich, 1820 (Reptilia: Serpentes: Uropeltidae) from Rakwana massif, Sri Lanka
No full textFIGURE 1. Map showing the type locality of Rhinophis erangaviraji sp. nov.Published as part of Mendis Wickramasinghe, L. J., Vidanapathirana, Dulan Ranga, Wickramasinghe, Nethu & Ranwella, P. Nayanaka, 2009, A new species of Rhinophis Hemprich, 1820 (Reptilia: Serpentes: Uropeltidae) from Rakwana massif, Sri Lanka, pp. 1-22 in Zootaxa 2044 on page 2, DOI: 10.5281/zenodo.18640
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