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Tingidae Laporte 1833
No full textFamily Tingidae Laporte, 1833 Subfamily Cantacaderinae Stål, 1873 Tribe Phatnomini Drake & Davis, 1960Published as part of Wappler, Torsten, 2003, New fossil lace bugs (Heteroptera: Tingidae) from the Middle Eocene of the Grube Messel (Germany), with a catalog of fossil lace bugs, pp. 1-26 in Zootaxa 374 on page 4, DOI: 10.5281/zenodo.15676
FIGURE 11 in New fossil lace bugs (Heteroptera: Tingidae) from the Middle Eocene of the Grube Messel (Germany), with a catalog of fossil lace bugs
No full textFIGURE 11. Oblongomorpha lutetia gen. et sp. n., reconstruction of the holotype, (macropterous form), compression fossil; coll. Forschungsinstitut Senckenberg (SMF), Frankfurt am Main (Germany), specimen SMF MeI 10508; scale bar represents 1 mm.Published as part of Wappler, Torsten, 2003, New fossil lace bugs (Heteroptera: Tingidae) from the Middle Eocene of the Grube Messel (Germany), with a catalog of fossil lace bugs, pp. 1-26 in Zootaxa 374 on page 13, DOI: 10.5281/zenodo.15676
A new bark-gnawing beetle (Coleoptera, Trogossitidae) from the middle Eocene of Europe, with a checklist of fossil Trogossitidae
No full textSchmied, Heiko, Wappler, Torsten, Č, Ji Ř Í Kolibá (2009): A new bark-gnawing beetle (Coleoptera, Trogossitidae) from the middle Eocene of Europe, with a checklist of fossil Trogossitidae. Zootaxa 1993: 17-26, DOI: 10.5281/zenodo.27467
FIGURES 17–22 in A description of Promanodes serafini gen. et sp. nov. from Baltic amber, with a review of related New Zealand Promanus Sharp, 1877 (Coleoptera: Trogossitidae)
No full textFIGURES 17–22. Promanus depressus Sharp: (17) wing; (18) female sternite VIII; (19) female tergite VIII; (20) maxillary palp; (21) labial palp; (22) hind leg. (J. Kolibáċ del.).Published as part of Kolibáč, Jiří, Schmied, Heiko, Wappler, Torsten & Kubisz, Daniel, 2010, A description of Promanodes serafini gen. et sp. nov. from Baltic amber, with a review of related New Zealand Promanus Sharp, 1877 (Coleoptera: Trogossitidae), pp. 29-44 in Zootaxa 2620 on page 38, DOI: 10.5281/zenodo.19807
FIGURE 6 in Fossil ants of the genus Gesomyrmex Mayr (Hymenoptera, Formicidae) from the Eocene of Europe and remarks on the evolution of arboreal ant communities
No full textFIGURE 6. Photographs of different casts and species of Gesomyrmex. (A) Gesomyrmex hoernesi Mayr – minor worker GMUG.BST. 04207. (B) Gesomyrmex hoernesi Mayr – worker PMHU.13/221. (C) Gesomyrmex hoernesi Mayr – male MKC.F-010. (D) Gesomyrmex hoernesi Mayr – major worker SIZK.K-419. (E) Gesomyrmex hoernesi Mayr – gyne PMHU.7/229. (F) G. b re v ic e ps sp. nov., gyne, holotype MeI 2305. (G) G. pulcher sp. nov., gyne, holotype MeI 10999. (H) G. curiosus sp. nov., head of gyne, holotype MeI 11953. (I) G. germanicus sp. nov., gyne, holotype PE-1997/29. (J) G. flavescens sp. n., gyne, holotype PE-2000/14. Scale bars = 1 mm.Published as part of Dlussky, Gennady M., Wappler, Torsten & Wedmann, Sonja, 2009, Fossil ants of the genus Gesomyrmex Mayr (Hymenoptera, Formicidae) from the Eocene of Europe and remarks on the evolution of arboreal ant communities, pp. 1-20 in Zootaxa 2031 on page 15, DOI: 10.5281/zenodo.18626
Formicidae Dlussky, Wappler & Wedmann, 2009, comb. nov.
No full textFormicidae (incertae sedis) miegi Théobald, 1937 comb. nov. Gesomyrmex Miegi Théobald, 1937, p. 211, pl. XIV, figs. 22, 23 (3, Ƥ). Gesomyrmex miegi Théobald: Burnham, 1978, p. 114; Bolton, 1995, p. 207. Comments. Described from two imprints from Kleinkembs, Haut-Rhin, France (early Oligocene), both without head and wing venation. The poor preservation of these imprints does not permit the assignment of these ants to any living or fossil genus, so we propose to regard these specimens as Formicidae incertae sedis.Published as part of Dlussky, Gennady M., Wappler, Torsten & Wedmann, Sonja, 2009, Fossil ants of the genus Gesomyrmex Mayr (Hymenoptera, Formicidae) from the Eocene of Europe and remarks on the evolution of arboreal ant communities, pp. 1-20 in Zootaxa 2031 on page 14, DOI: 10.5281/zenodo.18626
Exmesselensis Wappler, 2003, n. gen.
No full textExmesselensis n. gen. Type species: Exmesselensis disspinosus n. sp., here designated. Derivatio nominis: The new genusgroup name is derived from ex (Latin, meaning “out of”), plus the name of the type locality Messel. The name is masculine. Diagnosis: Small specimen, ca. 2.5 mm. Body moderately oblongoval. Head elongated, with one significant interocular (dorsomedial) tubercle. Bucculae elongated forward. Eyes globular, strongly projecting laterally of the head. Antennae thin, 3 rd segment long: just a little shorter than body width (relation 0.91). Pronotum more or less trapezoid, with 3 longitudinal carinae; posterior margin concave. Paranotum narrow. Scutellum completely exposed, triangular. Hemelytra distinctly subdivides by raised veins into costal, subcostal, discoidal, and sutural areas. Stenocostal area absent. Clavus well expressed. Comments: The new genus Exmesselensis combines some morphological characters of the Cantacaderinae and the Tinginae. The features of Cantacaderinae, in particular Phatnomini, includes: (1) the absence of stenocostal area of hemelytra (unfortunately it was impossible to examine in detail the scent gland openings and the morphology of the peritrema), (2) presents of only one dorsomedial (interocular) tubercle (completely absent in Cantacaderini and characteristic for Phatnomini), (3) the absence of a well expressed posterior triangular projection of the pronotum, (4) the exposed scutellum, (5) bucculae elongated forward and not touching anteriorly, and (6) clavi clearly separated by Pcu from corium by commissural. The absence of additional transverse veins in the discoidal area on the other hand is a characteristic feature of the majority of Tinginae contrary to Cantacaderinae in which they are usually well expressed in different degree, especially in Phatnomini (Froeschner 1996). Among those genera of the Phatnomini bearing a dorsomedial tubercle, habitually, the fossil genus Exmesselensis n. gen. is superficially quite similar to the living lace bug of the genus Distocader Froeschner. It shares in particular the head and antennal structure with the former. Exmesselensis n. gen. is distinguished from this endemic genus by relative broader pronotum, shorter carinae of the pronotum (majority of the Phatnomini show a 1 carinated pronotal disc), and the absence transverse veins in the discoidal area (this affiliates the fossil genus with the Phatnomini genera Pullocader Pericart, Ulmus Distant, and Zetekella Drake). Nevertheless, the incomplete state of material, however, does not permit an evaluation of its closer relationships at the present. A reconstruction of the type species of Exmesselensis based on the description below is shown in Fig. 2. The distribution of the extant representatives of the Phatnomini today is restricted to the southern hemisphere. The majority of the 26 described, extant genera are distributed in the Oriental and Ethiopian zoogeographic regions (Froeschner 1996). For instance, Distocader, represented by only a single species, is only known from New Guinea (Drake & Ruhoff 1965 b). Furthermore Exmesselensis shows once more those representatives of the Cantacaderinae were much more significant in the Paleogene fauna and widespread in Europe than today. In particular some genera of the Eocene European Phatnomini (e.g. Intercader Golub & Popov, Tingicader Golub & Popov, Exmesselensis n. gen.) show morphological similarities to the advanced subfamily Tinginae.Published as part of Wappler, Torsten, 2003, New fossil lace bugs (Heteroptera: Tingidae) from the Middle Eocene of the Grube Messel (Germany), with a catalog of fossil lace bugs, pp. 1-26 in Zootaxa 374 on pages 4-6, DOI: 10.5281/zenodo.15676
Oblongomorpha Wappler, 2003, n. gen.
No full textOblongomorpha n. gen. Type species: Oblongomorpha lutetia n. sp., here designated. Derivatio nominis: The new genusgroup name is a reference to the oblong shape of the body. The name is feminine. Diagnosis: Body oblong, surface areolate. Head very short, protruding little in front of the antennal insertion. No cephalic spines or tubercles. Eyes globular, strongly projecting laterally of the head. Antennae long and thin, first antennal segment 3 to 4 times as long as second, 3 rd prolonged, 4 th clubbed and covered with dense setae. Labium short, not reaching middle of metasternum. Pronotum with a ringlike vesicular at the anterior margin. Paranota absent. Triangular areolate posterior pronotal process is well developed, extremely elongated backwards along hemelytra covering whole clavi and approximately half of discoidal area. Pronotum with 3 longitudinal carinae. Macropterous form. Outline of hemelytra distinctly sinuate about midlength, apical rounded. Costal and subcostal area inconsicuous. Discoidal occupying nearly 0.54 of hemelytron length. Sutural area (membrane) in broadest place with 11–12 of angled cells. Legs moderately long. Tarsi 2 segmented, with second segment definitely broader and longer than segment I. Upper surface of segment II slightly convex, lower surface flat and covered with bristlelike hairs. Comments: The development of a celllike hind outgrowth of the pronotum that completely covers the clavi, a short and broad head, and the absence of additional protruding transverse veins on the hemelytra (plesiomorphic!) clearly classify Oblongomorpha n. gen. in the subfamily Tinginae. The general structure of the antennae and especially the presence of clearly modified second tarsal segment, with bristlelike hairs in a ventral concavity, unequivocally place Oblongomorpha in the tribe Litadeini. Even if the second tarsal segment is not quite as wide as in most of members of the tribe (e.g. Litadea China) it is even elongated and decidedly broader than the first tarsal segment. The decision of describing a new genus for the fossil species instead of including it in the genus Oecharis (only genus among the Litadeini with the same tarsal modification as in the fossil specimen) is mainly based on the absence of clearly prominent cyst formed by the paranota and the general structure of the hemelytra with small cells. The current classification of this tribe contains 13 extant genera (Froeschner 2001). So far no representative of the Litadeini has been reported from the fossil record. Lace bugs belonging to the tribe Litadeini range to be PanTropical without exception (Froeschner 2001). The oldest representative of the Tinginae (Archepopovia yurii Golub) has yet been recorded from Baltic amber. Judging on the characteristic features of this specimen the subfamily Tinginae with their main features was in the stage of formation during the Eocene (Golub 2001). Oblongomorpha n. gen. and Chorotingioites n. gen. clearly show that the differentiation of the Tinginae has taken place in this period or even earlier as already supposed by Golub (2001), and Golub & Popov (1999). A reconstruction of the type species of Oblongomorpha based on the description below is shown in Fig. 11.Published as part of Wappler, Torsten, 2003, New fossil lace bugs (Heteroptera: Tingidae) from the Middle Eocene of the Grube Messel (Germany), with a catalog of fossil lace bugs, pp. 1-26 in Zootaxa 374 on pages 11-12, DOI: 10.5281/zenodo.15676
Promanus subcostatus Broun 1909
No full textPromanus subcostatus Broun, 1909 (Figs 10–12, 24, 27, 31, 34, 36) Material examined. Female: “ Type ” [in red circle], “ 2761.” [handwritten], “Southland”, “ New Zealand./ Broun coll./ Brit. Mus./ 1922 - 482.”, “ Promanus subcostatus.” [handwritten]. The specimen deposited in The Natural History Museum, London (British Museum Natural History). Diagnosis. Largest of the species: body length 8.8 mm; dorsal surface reddish; elytra and pronotum nearly bare, head with pale pubescence; head dorsal surface very coarsely and densely sculptured; antenna 11 - segmented, with distinct 3 -segmented club; eyes distinctly elevated; gular area transversely wrinkled, with fine punctation along gular sutures only; elytra with four distinct carinae; anterior margin of mesosternum sharply incised; coxitae of ovipositor with distinct pubescence and coxitae smaller than those in P. auripilis; female tergite VIII (pygidium) with evenly rounded emargination of base; sternite VIII shown in Fig. 11.Published as part of Kolibáč, Jiří, Schmied, Heiko, Wappler, Torsten & Kubisz, Daniel, 2010, A description of Promanodes serafini gen. et sp. nov. from Baltic amber, with a review of related New Zealand Promanus Sharp, 1877 (Coleoptera: Trogossitidae), pp. 29-44 in Zootaxa 2620 on page 36, DOI: 10.5281/zenodo.19807
Gesomyrmex flavescens Dlussky, Wappler & Wedmann, 2009, sp. nov.
No full text<i>Gesomyrmex flavescens</i> sp. nov. <p>(Fig. 3 B and Fig. 6 J)</p> <p> <b>Derivation of the name.</b> <i>Flavescens</i> is Latin for yellowish.</p> <p> <b>Holotype.</b> NHMM PE- 2000/14 (Ƥ).</p> <p> <b>Type locality and horizon.</b> Eckfeld, Germany. Middle Eocene, ca. 44 Ma (Mertz <i>et al.</i> 2000).</p> <p> <b>Description.</b> Gyne. BL about 13 mm. Head subrectangular, 1.2 times longer than wide, with nearly parallel sides, sharp posterolateral corners and straight posterior margin. Anterior margin of the clypeus straight. Head 3.5 times longer than maximum eye diameter. Ocelli moderate large, distance between them some times greater than their diameter. Mesosoma a little wider than head. Scutum about as long as wide. Petiole with thick scale, longer than wide. Wing venation not preserved. Posterior part of the head, mesosoma, petiole and base of first gastral segment yellowish brown, anterior part of head and gaster varying from black to dark brown. Wings colorless with brown veins and pterostigma.</p> <p> <b>Measurements (in mm).</b> Holotype PE-2000/14: AL 4.0, HL 2.4, HW 2.0, ED 0.65.</p>Published as part of <i>Dlussky, Gennady M., Wappler, Torsten & Wedmann, Sonja, 2009, Fossil ants of the genus Gesomyrmex Mayr (Hymenoptera, Formicidae) from the Eocene of Europe and remarks on the evolution of arboreal ant communities, pp. 1-20 in Zootaxa 2031</i> on page 11, DOI: <a href="http://zenodo.org/record/186261">10.5281/zenodo.186261</a>
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