115 research outputs found
Lepidochlamus Wichard & Müller 2022
No full text<p>Family † Palleptoceridae Wichard & Müller, 2022</p> <p>Type genus and species.</p> <p> <i>Palleptocerus grimaldii</i> Wichard & Müller, 2022.</p> <p>Family diagnosis</p> <p> <b>(based on Wichard and Müller 2022).</b> Tibial spur formula 2/4/4. Antennae longer than forewings, flagellomeres cylindrical, elongate. Maxillary palps are five-segmented, terminal segment not annulated. Male forewing fork I and V present, discoidal cell absent. Hind wing exclusively fork V present.</p>Published as part of <i>Wichard, Wilfried, 2023, Fossil Trichoptera embedded in mid-Cretaceous Burmese amber, pp. 167-179 in Contributions to Entomology 73 (2)</i> on page 167, DOI: 10.3897/contrib.entomol.73.e11025
Cretapsyche Wichard, Neumann, Muller & Wang 2018
No full textGenus Cretapsyche Wichard, Neumann, Müller & Wang, 2018 Type species Cretapsyche circula Wichard, Neumann, Müller & Wang, 2018. Description See family description. So far, four species have been described and two new species are described here: Cretapsyche circula Wichard, Neumann, Müller & Wang, 2018 Cretapsyche elegans Wichard, Neumann, Müller & Wang, 2018 Cretapsyche insueta Wichard, Neumann, Müller & Wang, 2018 Cretapsyche palpinova Wichard & Neumann, 2019 Cretapsyche kachini Wichard & Espeland sp. nov. Cretapsyche myanmari Wichard & Espeland sp. nov.Published as part of Wichard, Wilfried & Espeland, Marianne, 2022, The family Cretapsychidae (Insecta, Trichoptera) from mid- Cretaceous Burmese amber, with descriptions of two new species, pp. 1-11 in European Journal of Taxonomy 833 on page 5, DOI: 10.5852/ejt.2022.833.1879, http://zenodo.org/record/691692
Cretapsyche Wichard, Neumann, Muller & Wang 2018
No full textKey of males of species of Cretapsyche Wichard, Neumann, Müller & Wang, 2018 Family and genus characters: five-segmented maxillary palps with second segment longest; threesegmented labial palps; forewing forks II, III, and V and hindwing forks I, III, and V present; tibial spurs 2/2/4. 1. Forewing with Rs branching before M; in male genitalia, harpago shorter than coxopodite, coneshaped.................................................................................................. Cretapsyche kachini sp. nov. – Forewing with Rs and M branching at about same level; in male genitalia, harpago about as long as coxopodite.......................................................................................................................................... 2 – Forewing with M branching before Rs; in male genitalia, harpago shorter than coxopodite, needleshaped................................................................................................................................................ 3 2. Labial palps with third segment oval, flattened; maxillary palps each with pilifer arising from third segment............................................................. Cretapsyche palpinova Wichard & Neumann, 2019 – Labial palps with third segment simple; maxillary palps simple, without pilifers..................................................................................... Cretapsyche circula Wichard, Neumann, Müller & Wang, 2018 3. Forewing with thyridial cell open, fork V petiolate........................ Cretapsyche myanmari sp. nov. – Forewing with thyridial cell closed, fork V nearly sessile................................................................. 4 4. Median cell open; in male genitalia mesodorsal process of tergum X distinct, ventrally protruded .......................................................... Cretapsyche insueta Wichard, Neumann, Müller & Wang, 2018 – Median cell closed; in male genitalia mesodorsal process of tergum X simple...................................................................................... Cretapsyche elegans Wichard, Neumann, Müller & Wang, 2018Published as part of Wichard, Wilfried & Espeland, Marianne, 2022, The family Cretapsychidae (Insecta, Trichoptera) from mid- Cretaceous Burmese amber, with descriptions of two new species, pp. 1-11 in European Journal of Taxonomy 833 on page 9, DOI: 10.5852/ejt.2022.833.1879, http://zenodo.org/record/691692
Palleptocerus WICHARD & MÜLLER 2022, gen. nov.
No full textPalleptocerus gen. nov. Type species. Palleptocerus grimaldii sp. nov. Etymology. The genus name is composed of the name of the related genus Leptocerus and with the abbreviation “Pal” = “paleo” for the mid-Cretaceous age of the extinct genus. Diagnosis. As for the family.Published as part of WICHARD, WILFRIED & MÜLLER, PATRICK, 2022, Palleptoceridae fam. nov., an extinct leptoceroid family in mid-Cretaceous Burmese amber (Insecta, Trichoptera), pp. 468-474 in Palaeoentomology 5 (5) on page 469, DOI: 10.11646/palaeoentomology.5.5.8, http://zenodo.org/record/733375
Palerasnitsyninae Wichard, 2023, stat. nov.
No full text<p>Subfamily † Palerasnitsyninae stat. nov.</p> <p>Type genus.</p> <p> <i>Palerasnitsynus</i> Wichard, Ross & Ross, 2011.</p> <p>Subfamily diagnosis.</p> <p>The species of the extinct subfamily Palerasnitsyninae are characterised by the combination of the fore and hind wings’ characters: in forewings by the presence of forks II, IV, V and by the absence of forks I and III and in hind wings by the presence of forks II and V and by the absence of the forks I, III, IV (Fig. 3B). Furthermore, the adults possess latero-apical dark sporns at the 3rd maxillary palp segments (Fig. 3A).</p> <p>Systematic position.</p> <p> The extinct genus <i>Palerasnitsynus</i>, with currently 10 species, was found in the Oriental mid-Cretaceous Burmese amber and initially placed in the family Psychomyiidae (Wichard et al. 2011). The wing venation is remarkably reduced, with only apical forks II, IV, V in forewings and apical forks II and V in hind wings. The reduction of forewing venations with the absence of forks I and III and of hind wing venations with the absence of forks I, III and IV is observed in some Xiphocentronids, but not in the family Psychomyiidae. This combination of reduced forewings and hindwings is significantly true for the family Xiphocentronidae. Based on these characters, it is proposed to transfer the genus <i>Palerasnitsynus</i> to family Xiphocentronidae and to establish an extinct subfamily Palerasnitsyninae stat. nov. Its species are the oldest Xiphocentronids with a geological age of about 100 million years.</p> <p> In Burmese amber, the species of the subfamily Palerasnitsyninae are certainly among the smallest caddisflies, reaching forewing lengths of only 1.8-2.6 mm (Wichard et al. 2011, 2018b). With an abundance of almost 40% of caddisflies, <i>Palerasnitsynus</i> species dominate the picture of caddisflies in the Burmese amber. They obviously have a tendency to swarm, as up to 100 embedded individuals have been counted in some ambers (Fig. 3C).</p>Published as part of <i>Wichard, Wilfried, 2023, Fossil Trichoptera embedded in mid-Cretaceous Burmese amber, pp. 167-179 in Contributions to Entomology 73 (2)</i> on page 167, DOI: 10.3897/contrib.entomol.73.e11025
Bipectinata orientalis Wichard et al., 2020 b 2023, comb. nov.
No full text<p>Bipectinata orientalis Wichard et al., 2020b comb. nov.</p> <p>Fig. 5</p> <p>Systematic position.</p> <p> <i>Bipectinata orientalis</i> was not originally placed in the family Odontoceridae, but was initially assigned to Calamoceratidae (Wichard et al. 2020), because the presence of wing fork IV on the forewing is not common to extant Odontoceridae. Nevertheless, characteristic features of the family Odontoceridae are synapomorphically present in the genus <i>Bipectinata</i>, such as five-segmented maxillary palps with a terminal segment not flexible or annulated, lack of ocelli, tibial spur formula 2/4/4; in forewings fork I present, discoidal cell closed and median cell absent. In addition, genus <i>Bipectinata</i> is closely related to the odontocerid genus <i>Palaeopsilotreta</i>, whose common synapomorphies involve a variable forewing media and the bipectinate antennae (Fig. 5B).</p> <p> In trichopteran adults a complete set of five apical forks on the forewings is clearly a plesiomorphic character (Comstock 1918; Holzenthal et al. 2007). A reduction of the original wing venation and the reduction of apical forks are derived in many adults, especially within the superfamily Leptoceroidea. However, the extinct <i>Bipectinata orientalis</i> from the middle Cretaceous is characterized by the original arrangement of five apical forks in the forewing venation (Fig. 5C). This feature distinguishes it from the closely related species of the genus <i>Palaeopsilotreta</i>.</p>Published as part of <i>Wichard, Wilfried, 2023, Fossil Trichoptera embedded in mid-Cretaceous Burmese amber, pp. 167-179 in Contributions to Entomology 73 (2)</i> on page 167, DOI: 10.3897/contrib.entomol.73.e11025
Cretapsyche myanmari Wichard & Espeland 2022, sp. nov.
No full textCretapsyche myanmari sp. nov. urn:lsid:zoobank.org:act: 468AEDBE-C3E8-4C1A-9AA7-578660AC0A16 Fig. 4 Diagnosis Cretapsyche myanmari sp. nov. can be distinguished from all other species in the genus by having M branching before Rs in the forewing and an open thyridial cell. Cretapsyche myanmari sp. nov. can also be distinguished in the male genitalia by the mesodorsal processes of tergum X, scale-shaped, each with two black spines. Etymology Cretapsyche myanmari sp. nov. is named after Myanmar, where the amber inclusion was found in Kachin State. Material examined Holotype MYANMAR – Kachin State • ♂; exact locality unknown; Mid-Cretaceous Burmese amber inclusion; deposited in the amber collection of the ZFMK; former Patrick Müller Collection, BUB 3597; ZFMK-TRI000830. Body well preserved; forewing in dorsal view, hindwing covered dorsally by forewing. Antennae and legs present. Male genitalia visible in ventral view. Description Male Ocelli absent. Antennae little longer than forewings. Scapus about as long as head, smaller pedicellus, followed by more than 30 cylindrical flagellomeres gradually shortening in length apically. Maxillary palps 5-segmented, 2 nd segment longest, following three segments successively smaller and shorter. Body and wings light brown. Forewing with length about 2.8 mm. In forewings, apical forks II, III and V present, discoidal and thyridial cells absent, Cu2 running continuously to Cu1b. In hindwings, apical forks I, III and V present (holotype hindwings partially spread laterally and visible). Tibial spur formula 2/2/4. In male genitalia, two-segmented inferior appendages characterized by finger-shaped, curved coxopodite and by short and cone-shaped harpago. Two appendages, each with two dark spines, located centrally in genitalia interpreted as probable mesodorsal pair of lobes of tergum X (Wichard 2021).Published as part of Wichard, Wilfried & Espeland, Marianne, 2022, The family Cretapsychidae (Insecta, Trichoptera) from mid- Cretaceous Burmese amber, with descriptions of two new species, pp. 1-11 in European Journal of Taxonomy 833 on page 7, DOI: 10.5852/ejt.2022.833.1879, http://zenodo.org/record/691692
Palleptoceridae WICHARD & MÜLLER 2022, fam. nov.
No full textPalleptoceridae fam. nov. Type genus. Palleptocerus gen. nov. Diagnosis. Ocelli absent. Antennae longer than forewings, scapus conically thickened. Maxillary palps five-segmented, lengths of palp segments in a row: short (1), long (2), long (3) short (4) and longest (5); terminal segment not annulated (Figs 2A, 3B inset). Wings: Forewings slightly narrow and apically rounded, hind wings shorter and broader. Sexual dimorphism in terms of wing length and wing venation: Forewing lengths in males a little shorter than in females. The light brown wings of the males are darker in the females. In males forewing venation with forks I and V present (Fig. 2B); in females forks I, III, and V present (Fig. 2C). In both sexes, discoidal cells open and thyridial cells closed, long. The hind wing venation reduced in both sexes; forks I, II, III, and IV absent, exclusively fork V present; discoidal cells open; crossveins not present (Fig. 2D). Tibial spurs: 2/4/4, metatibia, each with a pair of apical and preapical spurs (Fig. 1B) Palleptoceridae fam. nov. is characterized by the apomorphic combination of the following characters: Tibial spur formula 2/4/4. Forewing fork I present without discoidal cell and in hind wing exclusively fork V present.Published as part of WICHARD, WILFRIED & MÜLLER, PATRICK, 2022, Palleptoceridae fam. nov., an extinct leptoceroid family in mid-Cretaceous Burmese amber (Insecta, Trichoptera), pp. 468-474 in Palaeoentomology 5 (5) on page 469, DOI: 10.11646/palaeoentomology.5.5.8, http://zenodo.org/record/733375
Palaeopsilotreta Wichard & Wang 2017
No full textGenus Palaeopsilotreta Wichard & Wang, 2017 Type species Palaeopsilotreta xiai Wichard & Wang, 2017, by monotypy. Diagnosis Palaeopsilotreta can be placed in the family Odontoceridae based on the following combination of characters: ocelli absent; spur formula 2/4/4; males with five segmented maxillary palps, first segment without apicomesal nodule, terminal segment not flexible or annulated; mesoscutum with a pair of ovoid setal warts, mesoscutellum covered by a single wart, large, dome-like and ovoid; presence of forewing forks I, II and V in both wings; discoidal cells closed, median and thyridial cells absent. It can be separated from other odontocerid genera by the Cu 2 in the male forewings reaching the wing margin and not being reduced or absent as in all other genera, and by the unique combination of spur formula, number of wing forks in both wings of the males and the absence of a thyridial cell. Additions to description Antennae partially bipectinate, about as long as forewings. Rami of bipectinate flagellomeres slim and about as long as flagellomeres. Female antennae simple, not bipectinate. Maxillary palps 5-segmented, terminal segment not flexible or annulate. Head ocelli absent; frontal, antennal and posterior setose warts present (Fig. 4E). Pronotum with 2 pronotal setose warts, mesoscutum bearing pair of small setose warts, mesoscutellum with one dome-shaped wart. In fore- and hindwings: forks I, II and V present. In male: media simple, unbranched, running to apical wing margin, or media branched in M1+2 and M3+4. In female: forewing media branched in M1+2 and M3+4. Discoidal cells closed or open, medial and thyridial cells lacking. Tibial spurs: 2/4/4.Published as part of Wichard, Wilfried, Espeland, Marianne, Müller, Patrick & Wang, Bo, 2020, New species of caddisflies with bipectinate antennae from Cretaceous Burmese amber (Insecta, Trichoptera: Odontoceratidae, Calamoceratidae), pp. 1-17 in European Journal of Taxonomy 653 on pages 4-5, DOI: 10.5852/ejt.2020.653, http://zenodo.org/record/386247
Cretapsyche kachini Wichard & Espeland 2022, sp. nov.
No full textCretapsyche kachini sp. nov. urn:lsid:zoobank.org:act: 7B10F380-13D1-4885-9A19-0414DE8F4975 Fig. 3 Diagnosis Creatapsyche kachini sp. nov. can be distinguished from all other species in the genus by having Rs branching before M in the forewing. In all other species, Rs and M branch at about the same level, or M branches before Rs. Additionally, in the male genitalia the harpago is two-thirds the length of the coxopodite. Etymology Cretapsyche kachini sp. nov. is named after Kachin State in Myanmar, where the amber inclusion was found. Material examined Holotype MYANMAR – Kachin State • ♂; exact locality unknown; Mid-Cretaceous Burmese amber inclusion; deposited in the amber collection of the ZFMK; former Patrick Müller Collection, BUB 540; ZFMK-TRI000829. Body well preserved; forewing in dorsal view, hindwing covered dorsally by forewing. Antennae and legs present. Male genitalia visible in ventral view. Description Male Ocelli absent. Antennae probably about as long as forewings (holotype antennae incomplete). Scapus about as long as head. Maxillary palps 5-segmented, 2 nd segment longest, following segments decreasing in length, 3 rd segment basally barrel-shaped and extended laterad to form pointed spur corresponding in length to 4 th and 5 th segments. Body and wings light brown. Forewing with length 2.8 mm longer than small hindwing. In forewings, apical forks II, III and V present, discoidal cell absent, thyridial cell present, Cu2 running continuously to Cu1b. In hindwings, apical forks I, III and V present (difficult to see, holotype forewing hides hindwings in dorsal view). Tibial spur formula 2/2/4. In male genitalia, two-segmented inferior appendages characterized by basally broad coxopodite, tapering and curving mesad. Harpago elongate and tapering continuously towards apex, two-thirds of coxopodite length.Published as part of Wichard, Wilfried & Espeland, Marianne, 2022, The family Cretapsychidae (Insecta, Trichoptera) from mid- Cretaceous Burmese amber, with descriptions of two new species, pp. 1-11 in European Journal of Taxonomy 833 on pages 5-7, DOI: 10.5852/ejt.2022.833.1879, http://zenodo.org/record/691692
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