1,185 research outputs found

    Nychiodes eberti Wanke & Hausmann & Krogmann & Petrányi & Rajaei 2020, sp. nov.

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    Nychiodes eberti sp. nov. Wanke, Hausmann & Rajaei (figs 73–83, 120–122; map 3) Type material examined. Holotype, ♂, Türkei, Prov. Erzurum, Kopdagi Pass, vic. Ascale, 2200 m, 27.-31.vii.1978, Lichtf., leg. W. Thomas, g.prep. 0267/2019 D. Wanke; in SMNS. Paratypes, 1 ♂, 1 ♀, NE Türkei, Prov.Erzurum, Dogu Karadeniz Daglari, Korga Dagi, Köprüköy, Umg. bei Ispir, 2000 m, w. 28.vii.2001, e.o. 3.xi.2001, leg. J. Gelbrecht, S. Beshkov, R. Busse, A. Kazanci & E. Schwabe, g.preps (♂) 0456/2019 D. Wanke, (♀) 0457/2019 D. Wanke; 1 ♂, NE Türkei, Dogu Karadeniz Daglari, Ovit Dagi, ca. 5 km südl. Ovit Dagi Gecidi, 2400-2500 m, 27.vii.2001, leg. J. Gelbrecht, S. Beshkov, R. Busse, A. Kazanci & E. Schwabe, g.prep. 0241/2019 D. Wanke; all in PCJG. 1 ♂, Turkey, St. 2108, Nigde, Bolkardaglari N Side, SW Maden, 1600 m, 3.viii.1995, leg. D.v.d. Poorten, W. De Prins, g.prep. 0341/2019 D. Wanke; 1 ♀, Turkiye, Konya, 38 km W Konya, 1550 m, 8.-9.vii.[19]88, leg. S. Wagener, g.prep. 0443/2019 D. Wanke; 1 ♂, 1 ♀, Turkey, Erzincan, Caglayan, 4 km S. Kalecik, 1600 m, 13.- 15.vii.2000, leg. K. Larsen, g.prep. (♂) 0339/2019 D. Wanke, (♀) 0444/2019 D. Wanke; 2 ♂, Turkey, Konya, Toros Daglari, Adiller Taskent, 1700 m, 14.vii.1986, leg. Arne Moberg, g.prep. 0343/2019 D. Wanke; 1 ♂, Turkey, Ankara, 1150-1250 m, 10 km NW Kizilcahaman, 6.-7.viii.1989, leg. Fibiger, Esser; 1 ♂, Turkiye, St. 1711, Artvin, 8-10 km SW Yusufeli Coruh valley, 900-1000 m, 4.-9.vii.1991, leg. W. De Prins, D.v.d. Poorten, A. Riemis, g.prep. 0344/2019 D. Wanke; 3 ♂, Turkey, Nigde, Bolkar Daglari, N. s. Maden, 2100 m, 29.vii.1997, K. Larsen, g.prep. 0342/2019 D. Wanke; 1 ♂, Turkey, Sivas, Gökpinar, 10 km S Gürün, 1500 m, 1.viii.1997, leg. K. Larsen, g.prep. 0340/2019 D. Wanke; 1 ♂, Turkey, Sivas, Gökpinar, 12 km S Gürün, 1500 m, 25.vii.1998, leg. K. Larsen; 1 ♀, Turkey, Sivas, Gökpinar, 10 km S Gürün, 1500 m, 11.vii.2000, leg. K. Larsen, g.prep. 0445/2019 D. Wanke; 1 ♀, [Turkey], Maras, Binboga Daglari, Göksun Yalakköy, 1600 m, 17.vii.1986, leg. A. Moberg, g.prep. 0451/2019 D. Wanke; all in PCPS. 2 ♂, Türkei centr., Provinz Sivas, Gökpinar, 1,5 km westlich, N 38°39'21'', O 37°17'19'', 1600 m ü.NN, 02.vii.2008, LF, leg. Ralf & Sylvana Fiebig, g.prep. 2134/2017 H. Rajaei; 1 ♂, 1 ♀, Türkei centr., Provinz Nevsehir, Kappadokien, Göreme, N 38°39', O 34°50', 1080-1150 m ü.NN, 08.-11.vii.2011, LF, leg. R. Fiebig & S. Rothe, g.preps. (♂) 0398/2019 D. Wanke (♀) 0399/2019 D. Wanke; 1 ♂, Türkei centr., Prov.Nevsehir, Kappadok- ien, Aktepe1 km SSO, N 38°40'43'', O 34°52'25'', 1070 m ü.NN, 26.-27.viii.2009 LF, leg. R. & S. Fiebig, g.prep. 0400/2019 D. Wanke; 1 ♂, Türkei centr, Provinz Nigde, Aladag West, 5 km SSO von Sulucaova, N 37°58'13'', O 35°09'58'', 2200-2500m ü.NN, 16.viii.2009, LF, leg. R. Fiebig & S. Rothe, g.prep. 0396/2019 D. Wanke; all in PCRF. 2 ♂, Türkei, Kopdagi, 4.viii.1978, 2100m, leg. Dittrich, Austria; in PCTM. 2 ♀, Türkei, Prov.Erzurum, Kop- dagi Pass, vic. Ascale, 2200 m, 27.-31.vii.1978, Lichtf., leg. W. Thomas, g.preps 0263, 0435/2019 D. Wanke; 1 ♂, [Turkey], Kleinasien, Prov.Erzurum, 40 km NW Erzurum, vic. Egerti, 1850-2000 m, 30.vii.-01.viii.[19]80, g.prep. 0066/2018 D. Wanke; 5 ♂, [Turkey], Kleinasien, Prov.Kars, vic. Karakurt, Aras-Tal, 1500 m, 15.-16.vii.1978, leg. de Freina, g.preps 0076, 0077/2018 D. Wanke, 0260/2019 D. Wanke; 5 ♂, 1 ♀, [Turkey], Kleinasien, Prov.Kars, vic. Kagizman, Kötek, 1550 m, 29.-31.vii.[19]78, leg. de Freina, g.preps (♂) 0078, 0080/2018 D. Wanke, 0261, 0262, 0264/2019 D. Wanke, (♀) 0266/2019 D. Wanke; 2 ♂, [Turkey], Kleinasien, Prov.Erzurum, Umg. Ovacik, Camlika, 2100 m, 01.-02.viii.1980, leg. de Freina, g.prep. 0086/2018 D. Wanke; 1 ♂, [Turkey], Kleinasien, Prov.Kars, vic. Sarikamis, 2000-2300 m, 21-27.vii.[19]80, leg. de Freina, 0079/2018 D. Wanke; 1 ♀, Türkei, Anatolien, 25 km, südl. Sivas, 1500 m, 24.+ 25.vii.1978, leg. W. Thomas, g.prep. 0432/2019 D. Wanke; 2 ♀, [Turkey], Klei- nasien, Prov.Artvin, 5 km SE Sarigöl, 750 m, 31.vii.-09.viii.[19]83, leg. de Freina, g.prep. 2090/2017 H. Rajaei, 0436/2019 D. Wanke; all in SMNS. 6 ♂, [Turkey], Anatolien, Ankara, 1000 m, vi.1934, leg. Herbert Noack, g.preps 0161, 0162/2018 D. Wanke, 0282, 0283, 0284/2019 D. Wanke; 1 ♀, Türk., Ostkurdistan, Van Gölü, ca. 1800 m, 1.-31.Vii.1965, leg. Herbert Noack, g.prep. (♀) 0292/2019 D. Wanke; all in SMNK. 1 ♂, Türkei, vii.[19]95, leg. Gelbrecht, g.prep. 0464/2019, D. Wanke; in ZSM. Description. Wingspan ♂29–40 mm, ♀35–42 mm (forewing length ♂18–22 mm, ♀19–24 mm) (figs 73–83). Antennae bipectinate in both sexes. Frons flat, projecting about one quarter diameter of eye, smoothly scaled. Chaetosemata present as two small patches. Length of labial palpi about 1.0 times diameter of the eye, slightly exceeding frons. Proboscis absent. Ground colour of wings highly variable, varying from sandy grey-yellow and light brown to dark brown. Transverse lines faint or visible. Terminal line from light to dark brown, sometimes faint. Postmedial line of forewing, if present, curved between M1 and M2; antemedial line, if visible, curved outwards. Postmedial line of hindwing absent or slightly visible. Antemedial line often faint or slightly visible as a shadow. Underside of wings variable, from unicolorous to vaguely showing the pattern of the upper side. Discal spots present only on underside. In male genitalia (figs 120–122) uncus tapered and curved, basally broad (fig. 121). Gnathos well developed, strongly sclerotized, medially tongue-shaped. Saccus wide. Costa of valva narrow, strongly sclerotized, apically spinose, digitiform and slightly exceeding apex of valva. Both ampullae located in the distal half of valva, apically spinose; ampulla superior long, basally wide, medially narower; ampulla inferior well developed, of similar length like ampulla superior and narrower than the latter. Juxta anchor-shaped, stalk broad. Aedeagus broad (width-length ratio, 1:9), medially curved; cornutus half length of aedeagus. In female genitalia ovipositor broad, Apophyses anteriores one fifth length of apophyses posteriores (fig. 143). Sternite A9 conical. Lamella postvaginalis antero-posteriorly extended. Ductus bursae sclerotized. Corpus bursae membranous, pear-shaped, signum stellate. Note. Nychiodes eberti sp. nov. is highly variable in wing pattern, similar to the large variability in N. divergaria (see figs 52–83). Therefore, wing pattern and coloration are absolutely unreliable characters for a secure diagnosis of these two species and examination of male and female genitalia is mandatory. Diagnosis. In the distribution range of N. eberti sp. nov. the sympatrically occurring N. rayatica and N. divergaria can be confused when comparing wing pattern only (see figs 41, 42, 52–83). Furthermore, the wing pattern and coloration of N. eberti sp. nov. overlaps with that of N. convergata sp. nov., which is so far only known from Israel (figs 84, 85). Male genitalia of N. eberti sp. nov. with costa of valva characteristically narrow, digitiform at apex, slightly exceeding apex of valva (costa of valva apically very large and broad, clearly exceeding apex of valva in N. rayatica; costa of valva wide, not exceeding apex of valva in N. divergaria; costa of valva basally narrow, apically widely clubbed, clearly exceeding apex of valva in N. convergata sp. nov.) (see figs 105, 113–123). In N. eberti sp. nov. ampulla superior twice as broad as ampulla inferior, both ampullae not curved, located in the distal half of valva (ampulla superior conical, ampulla inferior strongly curved, both ampullae located at the centre of valva in N. rayatica; ampulla superior very long and broad, ampulla inferior highly variable, both ampullae located at the centre of valva in N. divergaria; ampulla superior digitiform, three times as broad as ampulla inferior, both ampullae located at the centre of valva in N. convergata sp. nov.) (see figs 105, 113–123). In N. eberti sp. nov. aedeagus thick and slightly curved, cornutus-aedeagus ratio 1/2 (aedeagus very small and short, cornutus-aedeagus ratio 1/ 2 in N. rayatica; aedeagus thick and strongly curved, cornutus-aedeagus ratio 2/ 3 in N. divergaria; aedeagus narrow and long; cornutus-aedeagus ratio 1/ 3 in N. convergata sp. nov.) (see figs 105b, 113 b–118b, 123b). Female genitalia of N. eberti sp. nov. with apophyses anteriores one fifth length of apophyses posteriores (strongly reduced apophyses anteriores, two strongly sclerotized spherical patches on sternite A 9 in N. divergaria; female of N. rayatica and N. convergata sp. nov. unknown) (see figs 141–143). Phenology. Flying from May to August. Biology. Unknown. Habitat. In altitudes from 50 up to 2500m. Distribution. Distributed in Turkey (map 3). DNA barcoding. Diverging by 3.1% (minimum pairwise distance) from the N. divergaria-subvirida complex. Nearest species (minimum pairwise distances): N. mirzayansi sp. nov. (3.1%) and N. convergata sp. nov. (3.4%) (fig. 145). Etymology. The name of this species is dedicated to Günter Ebert (born in 1935), former curator of the Lepidoptera collection in Karlsruhe State Museum of Natural History and editor in chief of the masterpiece "Die Schmetterlinge Baden-Württenbergs" in 10 volumes. During his many expeditions in Iran and Afghanistan, Günter Ebert collected the most important reference collection for these countries, deposited in SMNK (and parts in ZSM). All Nychiodes specimens, which were collected by Ebert are examined in this paper.Published as part of Wanke, Dominic, Hausmann, Axel, Krogmann, Lars, Petrányi, Gergely & Rajaei, Hossein, 2020, Taxonomic revision of the genus Nychiodes Lederer, 1853 (Geometridae: Ennominae: Boarmiini) with description of three new species-an integrative approach, pp. 1-61 in Zootaxa 4812 (1) on pages 34-36, DOI: 10.11646/zootaxa.4812.1.1, http://zenodo.org/record/394410

    Die Kirche des Anfangs. Festschrift fur Heinz Schürmann zum 65. Geburtstag. éd. par R. Schnackenburg, J. Ernst et J. Wanke, textes de J. B. Bauer, G. Baumbach, G. Delling, J. Dupont, J. Ernst, J. Gnilka, W. Grundmann, T. Holtz, K. Kertelge, J. Kremer, X. Léon-Dufour, H. Lubsczyk, F. Mussner, R. Pesch, P. Pokorny, I. de la Potterie, E. Ruckstuhl, K.H. Schelkle, R. Schnackenburg, G. Schneider, W. Thusing, W. Trilling, A. Vögtle, N. Walter, J. Wanke, H. Zimmermann. 1977

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    Ponthot Joseph. Die Kirche des Anfangs. Festschrift fur Heinz Schürmann zum 65. Geburtstag. éd. par R. Schnackenburg, J. Ernst et J. Wanke, textes de J. B. Bauer, G. Baumbach, G. Delling, J. Dupont, J. Ernst, J. Gnilka, W. Grundmann, T. Holtz, K. Kertelge, J. Kremer, X. Léon-Dufour, H. Lubsczyk, F. Mussner, R. Pesch, P. Pokorny, I. de la Potterie, E. Ruckstuhl, K.H. Schelkle, R. Schnackenburg, G. Schneider, W. Thusing, W. Trilling, A. Vögtle, N. Walter, J. Wanke, H. Zimmermann. 1977. In: Revue théologique de Louvain, 10ᵉ année, fasc. 1, 1979. p. 98

    Die Kirche des Anfangs. Festschrift fur Heinz Schürmann zum 65. Geburtstag. éd. par R. Schnackenburg, J. Ernst et J. Wanke, textes de J. B. Bauer, G. Baumbach, G. Delling, J. Dupont, J. Ernst, J. Gnilka, W. Grundmann, T. Holtz, K. Kertelge, J. Kremer, X. Léon-Dufour, H. Lubsczyk, F. Mussner, R. Pesch, P. Pokorny, I. de la Potterie, E. Ruckstuhl, K.H. Schelkle, R. Schnackenburg, G. Schneider, W. Thusing, W. Trilling, A. Vögtle, N. Walter, J. Wanke, H. Zimmermann. 1977

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    Ponthot Joseph. Die Kirche des Anfangs. Festschrift fur Heinz Schürmann zum 65. Geburtstag. éd. par R. Schnackenburg, J. Ernst et J. Wanke, textes de J. B. Bauer, G. Baumbach, G. Delling, J. Dupont, J. Ernst, J. Gnilka, W. Grundmann, T. Holtz, K. Kertelge, J. Kremer, X. Léon-Dufour, H. Lubsczyk, F. Mussner, R. Pesch, P. Pokorny, I. de la Potterie, E. Ruckstuhl, K.H. Schelkle, R. Schnackenburg, G. Schneider, W. Thusing, W. Trilling, A. Vögtle, N. Walter, J. Wanke, H. Zimmermann. 1977. In: Revue théologique de Louvain, 10ᵉ année, fasc. 1, 1979. p. 98

    Phaselia sihvoneni Werner, Wanke, Rajaei 2023, sp. nov.

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    Phaselia sihvoneni Werner, Wanke, Rajaei sp. nov. (Figs 43–46, 98–100, 136–137, 144). Type material examined. Holotype, 1 ♀, “ India, J & K [Jammu and Kashmir], Ladakh, Kharbu, 2800 m, 13.viii.1984, leg. W. Thomas ”, g. prep (♀) 1213/2021 D. Wanke (in SMNS). Paratypes (7 ♁♁, 1 ♀): 1 ♁, “ Tajikistan, W. Pamir, Rushan distr. (border with Afghanistan), Jazgulemskiy range, Shipad vill, 2000 m, 29.vii.2014, Leg. Valentina Zurilina ”, g. prep. (♁) 0130/2021 M. Werner (in PCPS). 2 ♁♁, “N-Pakistan, Naltar, 2950 m, N36°08', E74°12', Nr. 19, 18.vi.1992, leg. M. Hreblay & G. Csorba ”, g. prep. (♁) 0158/2021 M. Werner; 3 ♁♁, “N-Pakistan, 8 km N of Sost, 2800 m, N36°74', E74°49', Nr. 17, 15.vi.1992, leg. M. Hreblay & G. Csorba ”, g. prep. (♁) 0159/2021 M. Werner (in PCMS). 1 ♀, “ Pakistan, Baltistan, Satpara lake N of Skardu, A. LT., 2620 m, 15.vii.1990, F. Aluombard & J. Plante leg.”, g. prep. (♀) 0165/2021 M. Werner (in NHMUK). 1 ♁, “N-Pakistan, 2 km N of Karimabad, 2900 m, N36°25', E74°40', Nr. 15, 13.vi.1992, leg. M. Hreblay & G. Csorba ”, g. prep. (♁) 0153/2021 M. Werner (in PCMS). Description. Wingspan, males: 32–36 mm (average 32 mm, n = 5), females: 32.5–33 in females (average 32.6 mm, n = 3). Forewing. Ground colour warm white, slightly speckled with very fine brown to pale grey dots; basal area and postmedial area darker than medial area; antemedial line darker than basal area, clearly zigzagged with rounded to acute tips, all three teeth are of similar length; medial line present as a diffuse darker brown shadow close to postmedial line; postmedial line thin, dark brown, strongly kinked inwards on costal area; subterminal line dark brown, either continuous, serrate with well-pronounced triangular spikes, or interrupted in the middle, serrate with weakly pronounced triangular spikes, distally accompanied by a white shadow; terminal line dark brown, wavy. Hindwing. Ground colour white, speckled with brown towards terminal area; postmedial line brown, medially projected outwards; sometimes interrupted or even invisible; subterminal line barely visible as a light shadow. Discal spots usually visible as brown, short streaks or tiny dots on all wings, sometimes faded. Fringes in all wings white and brown, dark at the end of the veins. Ventral side generally paler than dorsal side, pattern of upper side shines through the wings, visible as a shadow (figs 43–46). Male genitalia. Uncus triangular, tip incurved; gnathos well-sclerotized, medially not fused; valva trapezoidal; costal part of valva slightly sclerotized, distally finger-shaped, extended over the apex of valvula; sacculus small, oval, with a few equally distributed spines, not elongated at the ventral tip of sacculus; juxta formed as two ovals fused in the upper half; saccus wide, triangular, terminally curved. Aedeagus thick and short, straight or slightly curved, vesica with two separate, well-sclerotized cornuti: a claw-like curved cornutus and a more or less twisted, spinose plate-like cornutus (figs 98–100). Female genitalia. Ovipositor large and wide; apophyses posteriores up to three times longer than apophyses anteriores; lamella antevaginalis wide, evenly sclerotized throughout its width, with slightly ruffle-like sclerotization; ductus bursae membranous, very large and round, wider than lamella antevaginalis, slightly tapered towards corpus bursae; corpus bursae membranous, ornamented with a small but well-sclerotized, round, stellate signum (figs 136–137). Diagnosis. Phaselia sihvoneni sp. nov. can be confused with P. serrularia, P. erika and P. kasyi. For differentiation of P. sihvoneni sp. nov. from P. serrularia and P. erika, see the diagnosis of the respective species above. Characters of the compared species are given in parentheses. P. sihvoneni sp. nov. vs. (P. kasyi) External characters (figs 43–49). No diagnostic characters. Male genitalia (figs 98–102). Sacculus oval; ventral tip not elongated; spinose plate-like cornutus well-twisted, about as long as claw-like curved cornutus (sacculus triangular, ventral tip slightly elongated; spinose plate-like cornutus slightly twisted, clearly shorter than claw-like curved cornutus). Female genitalia (136–139). Antrum very large, round and only slightly tapered towards corpus bursae (antrum spherical, ductus bursae formed from two semispherical lateral halfs fused together, strongly tapered towards corpus bursae). Phenology. Possibly uni- or bivoltine. Investigated specimen collected from June to August. Biology. Unknown. Habitat. Investigated specimens have been collected at altitudes from 620 m up to 3600 m. Distribution. Central Tajikistan, North Pakistan, and North India (fig. 144). DNA barcoding. No data available. Etymology. The name sihvoneni is dedicated to our friend and well-known Geometridae specialist, director of the Department of Zoology at Finnish Museum of Natural History, long-term cooperation partner Pasi Sihvonen.Published as part of Werner, Maria Johanna, Hausmann, Axel, Kostjuk, Igor, Wanke, Dominic & Rajaei, Hossein, 2023, Integrative taxonomic revision of the genus Phaselia Guenée, [1858] (Geometridae: Ennominae) in the Middle East and Central Asia, pp. 1-66 in Zootaxa 5326 (1) on pages 21-22, DOI: 10.11646/zootaxa.5326.1.1, http://zenodo.org/record/824381

    Scopula sacraria subsp. ariana

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    <i>Scopula sacraria ariana</i> <p> 4 ♁/ ♀, S– Iran, prov. Fars, Tange Surkh, 50 km NW Ardekan, 12.–15.vi.1975, 2250 m NN, leg. Ebert & Falkner; 42 ♁/ ♀, S– Iran, prov. Khuzestan, Yasudj, Sisakht, 50 km NW, 15.–18.vi.1975, leg. Ebert & Falkner, g. preps (♁) 0598, 0599/2020 D. Wanke; 1 ♁, N– Iran, Elburs – Mts. S– Rand, Tehran – Evin, 25.–28.vi.1972, 1800 m, leg. Ebert & Falkner, g. prep. 0683/2020 D. Wanke; 1 ♀, same data, but 24.ix.1972, 1600 m, leg. Ebert & Falkner, g. prep. 0684/2020 D. Wanke; 1 ♁/ ♀, same data, but 29.ix.1972, 1600 m; 3 ♁/ ♀, same data, but 30.vi.–5.vii.1972, 1600 m; 1 ♀, same data, but 30.–31.vi.1972, 1600 m; 1 ♁/ ♀, N– Iran, 70 km s. Teheran, 29.v.1969, 1300 m, leg. Ebert; 3 ♁/ ♀, Iran, Derbend, 25 km N v. Teheran, 1.–10.vi.1962, 2000 m, E. & A. Vartian, g. prep. (♁) 0608/2020 D. Wanke; 1 ♀, Iran, prov. Hamadan, 8 km S of Arak, 02. – 03.vi.2005, leg. P. Gyulai & A. Garai, g. prep. 0807/2020 D. Wanke; 57 ♁/ ♀, Iran, prov. Boyerahmad –va– Kohgiluyeh, SE – Zagros, 35 km SE of Yasuj, 06.–07.vi.2005, 2600 m, leg. P. Gyulai & A. Garai, g. preps (♁) 0506, 0509, 0555, 0556, 0557, (♀) 0507, 0508, 0754, 0755/2020 D. Wanke; 24 ♁/ ♀, W– Iran, Lorestan, Dorud, 5 km SE Saravand, “Kohyeh”, 29–30.vii.1975, 2300 m, leg. Ebert & Falkner, g. preps (♁) 0561, 0693/2020 D. Wanke; 8 ♁/ ♀, W– Iran, Lorestan, 28 km E Borudjerd, 2300 m, 27.7. 1975, leg. Ebert & Falkner, g. preps (♀) 0691, 0692/2020 D. Wanke; 3 ♁/ ♀, W– Iran, Lorestan, Dorud, 4 km SE Saravand, “Nermyeh”, 4.–6.viii.1975, 2400 m, leg. Ebert & Falkner; 5 ♁/ ♀, W– Iran, Lorestan, 14 km E Dorud, 6.viii.1975, 1990 m, leg. Ebert & Falkner; 4 ♁/ ♀, W– Iran, Kordestan, Strasse Baneh–Marivan, 86 km SE Baneh, 5.vii.1975, 1950 m, leg. Ebert & Falkner, g. preps (♁) 0643, (♀) 0644/2020 D. Wanke; 6 ♁/ ♀, W– Iran, Kordestan, Strasse Baneh–Marivan, 25 km E Baneh, 4.vii.1975, 1950 m, leg. Ebert & Falkner, g. prep. (♀) 0649/2020 D. Wanke; 7 ♁/ ♀, W– Iran, Kordestan, Strasse Saghez– Baneh, 21 km NE Baneh, 1950 m, 30–2.vii.1975, leg. Ebert & Falkner; g. preps (♁) 0651, (♀) 0652/2020 D. Wanke; 2 ♁/ ♀, W– Iran, 51 km westl. Kermanshah, 1500 m, Quercetum, 17.vi.1975, leg. H. G. Amsel; 3 ♁/ ♀, W– Iran, Kermanshahan, Ghalladje Pass, 40 km S Schahabad, 1880 m, 13.vii.1975, leg. Ebert & Falkner, g. prep. (♀) 0659/2020 D. Wanke; 5 ♁/ ♀, NW– Iran, 100 km Strasse Mahabad – Sardascht, 1300 m, Quercetum, 13.vi.[19]75, leg. H. G. Amsel, g. prep. (♁) 0659/2020 D. Wanke; 2 ♁/ ♀, W– Iran, Kordestan, 95 km N Kermanschah, Strasse nach Sanandaj, 11.vii.1975, 1350 m, leg. Ebert & Falkner, g. prep. (♁) 0645/2020 D. Wanke; 3 ♁/ ♀, W– Iran, Kordestan, Ariz, 27 km W Sanandaj, 10.vii.1975, 2200 m, leg. Ebert & Falkner, g. prep. (♁) 0646/2020 D. Wanke; 2 ♁/ ♀, W– Iran, 15 km nördl. Kermanshah, 1350 m, 16.vi.1975, leg. H. G. Amsel, g. prep. (♁) 0657/2020 D. Wanke; 2 ♁/ ♀, W– Iran, 60 km nördl. Kermanshah, 1400 m, 18.vi.1975, leg. H. G. Amsel, g. prep. (♀) 0656/2020 D. Wanke; 5 ♁/ ♀, W– Iran, W– Azarbaijan, 2 km W Sardasht, 1650 m, 3.vii.1975, leg. Ebert & Falkner, g. prep. (♁) 0656/2020 D. Wanke; 17 ♁/ ♀, S– Iran, Strasse Shiraz– Kazerun, Imam Sade, 1200 m, 3.vi.1969, leg. G. Ebert, g. preps (♀) 0689, 0690/2020 D. Wanke; 101 ♁/ ♀, S– Iran, Miyan Kotal, 4.–7.vi.1969, 1900 m, östl. Kazerun, 51°40’ öL., 29°30’ nB., leg. G. Ebert, g. prep. (♀) 0600/2020 D. Wanke; 32 ♁/ ♀, S– Iran, Fars, Kaserun, Mian –Kotal, 1900 m, 11.vi.1972, leg. Ebert & Falkner, g. preps (♁) 0633, 0634 (♀) 0632, 0635, 0636/2020 D. Wanke; 6 ♁/ ♀, Iran, prov. Fars, S– Zagros, 40 km SW of Sivand, 09.– 10.06. 2005, leg. P. Gyulai & A. Garai, g. prep. (♁) 0537/2020 D. Wanke; 3 ♁/ ♀, Iran, Fars, Shiraz ESE, Darab N (Pass), 1850 – 2100 m, 19.v.2005, leg. T. & A. Hofmann, g. preps (♁) 0541, (♀) 0750/2020 D. Wanke; 1 ♁, Iran, prov. Fars, Shiraz–Kazerun road, 5 km before Dashte Arjan, N 29°40’34’’, E 052°02’18’’, 2158 m, 23.v.2009, leg. Hossein Rajaei, g. prep. (♁) 0550/2020 D. Wanke; 1 ♁/ ♀, Iran, Fars, Umgebung von Chiraz, 10.iv.1937, ca. 1600 m, coll. Brandt; 2 ♀, Iran, Fars, Strasse Chiraz– Kazeroun, Fort Sine –Sefid, ca. 2200 m, 4.ix.1937, coll. Brandt, g. preps 0581, 0590/2020 D. Wanke; 1 ♁, [Iran], Fars, Kazeroun, Mian –Kotal, 1900 m, 11.vi.1972, Mi.Li, leg. Ebert & Pazouki, g. prep. 0681/2020 D. Wanke; 1 ♁, S– Iran, Khusestan, Yassudj, Sisakht, 2250 m, 13./ 14.vi.1972, leg. Ebert & Falkner, g. prep. 0680/2020 D. Wanke; 6 ♁/ ♀, Iran, prov. Chahar Mahal, Zagros mts. NW Samsami, 2800 mNN, N 32°09’, E050°11’, 13.vii.2003 (lux), leg. G. Ebert & R. Trusch, g. prep. (♀) 0756/2020 D. Wanke; 1 ♁/ ♀, NE– Iran, prov. Ostan –e Khorasan, Kopet Dagh, NW Mashad, N Tschenaran, N Radkan, Dolmeh Olia, N 36°55’56.6’’, E 59°02’18.6’’, 8.v.2008, 1.560 mNN, lux, leg. R. Trusch M. Falkenberg & B. Müller; 10 ♁/ ♀, same data, but 9.v.2008, g. preps (♁) 0512, (♀) 0749/2020 D. Wanke; 24 ♁/ ♀, same data, but 10.v.2008, g. preps (♁) 0511, 0513, 0514, 0515/2020 D. Wanke; 9 ♁/ ♀, same data, but 11.v.2008, g. prep. (♁) 0517/2020 D. Wanke; 4 ♁/ ♀, Iran NE, Kopet Dagh, prov. Khorasan, ca. 50 km N Bojnurd, S Izmansufla, N 37°44’20’’, E 57°25’53’’, 1.240 mNN, 17.v.2005, leg. Trusch, Petschenka, Müller, g. preps (♁) 0529, (♀) 0530/2020 D. Wanke; 1 ♁/ ♀, Iran, prov. Tehran, Elburz mts. 3 km NNW Shemshak, N36°02’, E051°28’, 2860 mNN, 24.vii.2003 (lux), leg. G. Ebert & R. Trusch; 1 ♀, Iran, prov. Hamadan, 8 km S of Arak, 02–03.vi.2005, leg. P. Gyulai & A. Garai, g. prep. 0540/2020 D. Wanke; 2 ♁, Iran, prov. Hamadan, 8 km S of Arak, 02–03.vi.2005, leg. P. Gyulai & A. Garai, g. prep. 0544, 752/2020 D. Wanke; 1 ♀, Iran N, prov. Semnan, 30 km NW Damghan, Cheschme Ali, N 36°15’07’’, E 54°04’20’’, 1560 mNN, 23.v.05, leg. Trusch, Petschenka, Müller, g. prep. 0532/2020 D. Wanke; 1 ♁, Iran, prov. Lorestan, Oshtorankuh, Dorud –Gahar, lake road, before Cheshmeh Khorram, 2360 mNN, N33° 22’41’’, E49°11’13’’, 22.–24.vi. 2009, leg. H. Rajaei, J.U. Meineke & A. Hofmann, g. prep. 0551/2020 D. Wanke; 1 ♁, Iran, prov. Kohkiluyeh –va– Boyer –ahmad, 30 km S Yassuj, road Abshare–Tange–Tamoradi, 8km before Abshar [=waterfall], N30°31’53’’; E51°25’11’’, 2254 mNN, 24.v.2009, leg. Hossein Rajaei, g. prep. 0552/2020 D. Wanke; 2 ♁/ ♀, S– Iran, 160 km n. Shiraz, 1900 m, s. Didegan, 8.vi.1969, leg. G. Ebert, g. prep. (♁) 0658/2020 D. Wanke; 1 ♁, [Iran], Fadiheh –T–H, 20.7.1971, leg. Paz., Ayat., g. prep. 0562/2020 D. Wanke; 1 ♀, N– Iran, Elburs – Mts., prov. Tehran, Arangeh 25 km N Karadj, 1550 m, 1.–6.vi.1972, leg. Ebert & Falkner, g. prep. 0687/2020 D. Wanke; 2 ♁/ ♀, NW– Iran, 17 km nw. Maku, 1400 m, 4.vi.1975, leg. H. G. Amsel, g. prep. (♀) 0661/2020 D. Wanke; 1 ♀, NW– Iran, 15 km sö. Maku, 1050 m, 3.vi.1975, leg. H. G. Amsel, g. prep. 1143/2021 D. Wanke; 2 ♁/ ♀, Iran – Centr., Prov. Yazd, N Yazd, Chak Chak, N 32°20’ 07.8’’, E 54°22’58.0’’, 1.550 mNN, 10. – 11.iv.2007, leg. R. Trusch, SMNK E– Lep. 234, g. prep. (♀) 0545/2020 D. Wanke; 1 ♀, N– Iran, Elburs Mts., Prov. Tehran, 15 km E Gatschsar, 17.viii.1972, 2800 m, leg. Ebert, g. prep. 1164/2021 D. Wanke; 1 ♁/ ♀, [Iran], Karkas, 9.vi.1970, [leg.] Mirz. & Abai; 1 ♁, [Iran], Darrehgaz, 3.vii.1971, [leg.] Paz. & Abai, g. prep. 1147/2021 D. Wanke; <b>all in SMNK</b> <b>.</b></p> <p> 3 ♁/ ♀, Iran, prov. Kohkiluyeh –va– Boyer –ahmad, 30 km S Yassuj, road Abshare–Tange–Tamoradi, 8km before Abshar [=waterfall], N30°31’53’’; E51°25’11’’, 2254 mNN, 24.v.2009, leg. Hossein Rajaei, g. preps (♁) 0714, 0715/2020 D. Wanke; 4 ♁/ ♀, Iran, prov. Lorestan, Oshtorankuh, Dorud –Gahar, lake road, before Cheshmeh Khorram, 2360 mNN, N 33°22’41’’, E 49°11’13’’ 22.–24.vi.2009, leg. H. Rajaei, J.U. Meineke & A. Hofmann, g. prep. (♁)0724/2020 D. Wanke; 1 ♁/ ♀, Iran, Shahrud, Shahkouh, Tash, Ayoub Hosseini region, 2588m, 36°37’18’’N, 54°33’ 42.6’’E, 11.07.2016, leg. Sh. Feizpour; 1 ♁, Iran, prov. Chaharmahal –va– Bakhtiyari, Dehnau, 2248m, 12.v.2010, Leg. G. Petrányi, P. Hentschel, g. prep. 0747/2020 D. Wanke; 1 ♁, Iran, prov. Esfahan, near Qamsar, 1781m, 06.v.2010, Leg. G. Petrányi, P. Hentschel, g. prep. 0745/2020 D. Wanke; 4 ♁/ ♀, Iran, prov. Hamedan, Nehavand, 1855m, 13.v.2010, Leg. G. Petrányi, P. Hentschel, g. prep. (♁) 0743/2020 D. Wanke; 8 ♁/ ♀, Iran, prov. Zanjan, Ab Bar, 1053m, 17.v.2010, Leg. G. Petrányi, P. Hentschel, g. preps (♁) 0739, (♀) 0738, 0740/2020 D. Wanke; <b>all in SMNS</b> <b>.</b></p> <p> 2 ♁/ ♀, N– Iran, S. Elburs, Varamin, 700 m, semidesert, 11.–21.vii.1996, leg. Müller; <b>in ZSM</b> <b>.</b></p>Published as part of <i>Wanke, Dominic, Hausmann, Axel, Lee, Kyung Min, Murillo-Ramos, Leidys, Sihvonen, Pasi & Rajaei, Hossein, 2023, Systematics and integrative taxonomic revision of the tribe Scopulini Duponchel, 1845 in Iran (Lepidoptera: Geometridae: Sterrhinae), pp. 1-96 in Zootaxa 5359 (1)</i> on pages 94-95, DOI: 10.11646/zootaxa.5359.1.1, <a href="http://zenodo.org/record/10147920">http://zenodo.org/record/10147920</a&gt

    PLATE 14 in Systematics and integrative taxonomic revision of the tribe Scopulini Duponchel, 1845 in Iran (Lepidoptera: Geometridae: Sterrhinae)

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    PLATE 14: Figures 1–3. Male genitalia of Scopula species. 1: S. ornata enzela (N-Iran, Bandar Pahlavi, g. prep. 0797/2020 D. Wanke); 2: S. orientalis (Iran, Elburz, g. prep. 1192/2022 D. Wanke); 3: S. decorata (W Iran, Kordestan, Strasse Zandjan-Bijar, g. prep. 0792/2020 D. Wanke). a = genitalia capsule; b = aedeagus; c = 8th sternite. Scale-bar 1 mm.Published as part of <i>Wanke, Dominic, Hausmann, Axel, Lee, Kyung Min, Murillo-Ramos, Leidys, Sihvonen, Pasi & Rajaei, Hossein, 2023, Systematics and integrative taxonomic revision of the tribe Scopulini Duponchel, 1845 in Iran (Lepidoptera: Geometridae: Sterrhinae), pp. 1-96 in Zootaxa 5359 (1)</i> on page 61, DOI: 10.11646/zootaxa.5359.1.1, <a href="http://zenodo.org/record/10147920">http://zenodo.org/record/10147920</a&gt

    Nychiodes rayatica Wiltshire 1957

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    <i>Nychiodes rayatica</i> Wiltshire, 1957 <p>1 ♂, [Turkey], Kleinasien, Prov. Hakkari, Hakkari-Daglari, 10 km, östl. Gecitli, 2100-2300 m, 13.vii.-14.vii.[19]80, leg. de Freina, g.prep. 0254/2019 D. Wanke; in SMNS.</p> <p>1 ♂, W-Iran, Kordestan, Strasse Baneh-Marivan, 25 km E Baneh, 1950 m, 4.vii.1975, leg. Ebert & Falkner, g.prep. 0197/2018 D. Wanke; in SMNK.</p>Published as part of <i>Wanke, Dominic, Hausmann, Axel, Krogmann, Lars, Petrányi, Gergely & Rajaei, Hossein, 2020, Taxonomic revision of the genus Nychiodes Lederer, 1853 (Geometridae: Ennominae: Boarmiini) with description of three new species-an integrative approach, pp. 1-61 in Zootaxa 4812 (1)</i> on page 51, DOI: 10.11646/zootaxa.4812.1.1, <a href="http://zenodo.org/record/3944102">http://zenodo.org/record/3944102</a&gt

    Scopula immorata subsp. riloensis

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    <i>Scopula immorata riloensis</i> <p> 1 ♀, Turkey, Erzurum, 20 km nördl. Askole, Kop Gec, 2300 m, 16.vii.1992, g. prep. 1226/2021 D. Wanke; 1 ♁, Persia sept., Elburs mts.c.s., Tacht i Suleiman, Hecarcal – Tal, 28–3200 m, 3.–7.vii.[19]37, E. Pfeiffer & W. Forster leg., München, g. prep. 1225/2021 D. Wanke; <b>all in SMNK</b> <b>.</b></p> <p> 1 ♁, Turkey, Erzurum, Kopdagi gec., 2400 m, 10.–14.vii.1983, leg. W. Thomas, g. prep. 1230/2021 D. Wanke; 1 ♁, Kleinasien [Turkey], prov. Tokat – Sivas, Camlibel–Pass, 1700 m, 1.–10.vii.1978, leg. deFreina, g. prep. 1231/2021 D. Wanke; 1 ♁, Mongolia, Chentej aimak Tsenhernandel, Modoto Chentej Mts. 1600–1800 m, 47°48’N, 109°04’E, 9.–14.vii.1984, leg. Cerny, g. prep. 1232/2021 D. Wanke; <b>all in SMNS</b> <b>.</b></p> <p> 1 ♁, [Iran] Persia sept., Elburs mts.c.s., Tacht i Suleiman, Hecarcal–Tal, 28–3200 m, 3.–7.vii.[19]37, E. Pfeiffer & W. Forster leg., München, g. prep. 1259/2022 D. Wanke; <b>in ZFMK</b> <b>.</b></p> <p> 1 ♀, Persia sept. [Iran], Elburs mts.c.s., Tacht i Suleiman, Särdab–Tal (Tanakarud), 29–3200 m, 19.–23.vii.[19]37, leg. E. Pfeiffer & W. Forster, München, g. prep. 1284/2022 D. Wanke; 1 ♁, Persia sept. [Iran], Elburs mts.c.s., Särdab–Tal, 3000 m, 19.–23.vii.[19]37, leg. E. Pfeiffer & W. Forster, München, g. prep. 1285/2022 D. Wanke; 1 ♁, Persia sept. [Iran], Elburs mts.c.s., Särdab–Tal, Vandarban, 19–2200 m, 10.–14.vii.[19]37, leg. E. Pfeiffer & W. Forster, München, g. prep. 1286/2022 D. Wanke; <b>all in ZSM</b> <b>.</b></p>Published as part of <i>Wanke, Dominic, Hausmann, Axel, Lee, Kyung Min, Murillo-Ramos, Leidys, Sihvonen, Pasi & Rajaei, Hossein, 2023, Systematics and integrative taxonomic revision of the tribe Scopulini Duponchel, 1845 in Iran (Lepidoptera: Geometridae: Sterrhinae), pp. 1-96 in Zootaxa 5359 (1)</i> on page 81, DOI: 10.11646/zootaxa.5359.1.1, <a href="http://zenodo.org/record/10147920">http://zenodo.org/record/10147920</a&gt

    Synopsia phasidaria subsp. phasidaria phasidaria (Rogenhofer 1873, comb. nov.

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    Synopsia phasidaria phasidaria (Rogenhofer, 1873) comb. nov. (figs 8–22, 29–33, 36, 38–40, 42) Nychiodes phasidaria Rogenhofer, 1873. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien 23, 572. Syntypes (Caucasus, Achalzik [Akhaltsikhe]) (deposition of type specimen(s) unknown). Scodonia tekkearia Christoph, 1883. In Romanoff, Mémoires sur les Lépidoptères 2, 123. Holotype ♂ ([Turkmenistan], near Nuchur [Nokhur]). According to Wehrli (1954) deposited in museum of St Petersburg. Wehrli (1941) combined this taxon with Synopsidia. This subspecies is regarded as a junior synonym of Synopsidia phasidaria by Viidalepp (1996). Here confirmed as junior synonym of Synopsia phasidaria phasidaria comb. nov., based on morphological examination and sympatric occurrence of these forms. Synopsia znojkoi Djakonov, 1935. Lambillionea 35 (7), 142. Syntypes ♂ (Transcaucasus: Paraga, NW of Ordubad). Deposition of type specimen(s) unknown. Examination after Djakonov´s description and illustration of the male genitalia. This subspecies is regarded as a junior synonym of Synopsidia phasidaria by Viidalepp (1996). Here confirmed as junior synonym of Synopsia phasidaria phasidaria comb. nov., based on morphological examination and sympatric occurrence of these forms. Hashtaresia jodes Wehrli, 1936. Lambillionea 36 (12), 276. Holotype ♂ (NHMUK, examined). Wehrli (1941) combined this taxon with Synopsidia. Hereby regarded as a new synonym of Synopsia phasidaria phasidaria based on morphological examination and sympatric occurrence of these forms. Synopsidia phasidaria ardschira Brandt, 1938. Entomologische Rundschau 55 (51), 602. Holotype ♂ (NHRS examined). Here-by regarded as a new synonym of Synopsia phasidaria phasidaria based on morphological examination and sympatric occurrence of these forms. Synopsidia phasidaria chiraza Brandt, 1938. Entomologische Rundschau 55 (51), 602. Syntypes (Iran, Sine-Sefid). (NHRS). Hereby regarded as a new synonym of Synopsia phasidaria phasidaria based on morphological examination and sympatric occurrence of these forms. Synopsidia phasidaria mirabica Wehrli, 1941. In: Seitz, A. (Ed.), Die Grossschmetterlinge der Erde. Supplement zu Band 4, 464. Syntypes ♂, ♀ (ZFMK, examined). Hereby regarded as a new synonym of Synopsia phasidaria phasidaria based on morphological examination and sympatric occurrence of these forms. Synopsidia phasidaria alvandi Wiltshire, 1966. Zeitschrift der Wiener Entomologischen Gesellschaft, 51, 146. Holotype ♂ (NHMUK, examined). Hereby regarded as a new synonym of Synopsia phasidaria phasidaria based on morphological examination and sympatric occurrence of these forms. Type material examined. Synopsidia phasidaria alvandi: Holotype, ♂, Iran, Mt. Alvand, 9000ft. [2743 m], 27.vii.1938, E.P. Wiltshire, g.prep. E.P. Wiltshire 1470; in NHMUK (fig. 9). Synopsidia phasidaria ardschira: Paratype, 1 ♂, Iran, Fars, Strasse Ardekan-Talochosroe, Comèe [Komehr] (Barm i Firus), ca 3750 m, 4. und 5.vi.1937, coll. Brandt, g.prep. 11026; in NHRS (fig. 8). Synopsidia phasidaria jodes: Holotype, ♂, Iran, Hashtar, Demawend, 2500 m, viii.1935, F. Fusek, g.prep. 7100; in NHMUK (fig. 10). Synopsidia phasidaria mirabica: Syntype, 1 ♂, Nordost-Iran, Kuh i Mirabi, Waldzone, 1600–1900 m, Juli, Exp. Wernicke, g.prep. 0501/2020 D. Wanke; Syntype, 1 ♀, Nord-Iran, Schahkuh, Westabhang, Geröllzone, 1800–2000 m Juni, Exp. Wernicke, g.prep. 0502/2020 D. Wanke; all in ZFMK (figs 11–12). Additional material examined. 2 ♀, Iran, Mt. Alvand, 9000 ft. [2743 m], 27.vii.1938, E.P. Wiltshire; in NHMUK. 1 ♂, Iran, Fars, Strasse Chiraz-Kazeroun, Fort Sine-Sefid, ca. 2200 m, Sept.1937, coll. Brandt, g.prep. 11027; 1 ♂, Iran, Fars, Strasse Ardecan-Talochosroe [Ardakan-Talle Khosrow], Comèe [Komehr] (Barm i Firus), ca. 3750 m, 4. und 5.vi.1937, Brandt, g.prep. 11028; in NHRS. 1 ♂, Iran, Kordestan, Baneh 15 km NE, Gharawol-Kaneh [Khaneh], 2000 m, 26./ 27.vi.2009, leg. A. Hofmann, J.-U. Meineke, H. Rayai [Rajaei], g.prep. 0213/2019 D. Wanke; in PCJM. 1 ♂, Türkei, Cukurca, Prov. Hakkari, 20.vi.1986, leg. Dittrich, Austria, 16273 coll. R.Wimmer, g.prep.0382/2019 D. Wanke; 1 ♂, 2 ♀, Republic Armenia, Yeranos Mts. 1600 m, Dvin village suburbs, Arat district, 11.–13.vi.2009, leg. Yuriy Shevnin, g.prep. 0381/2019 D. Wanke; 1 ♂, 1 ♀, Republic Armenia, Yeranos Mts. 1600 m, Dvin village suburbs, Arat district, 14.–15.vi.2009, leg. Yuriy Shevnin, g.preps (♂) 0492/2019 D. Wanke, (♀) 0493/2019 D. Wanke; 2 ♂, Republic Armenia, Yeranos Mts. 1600 m, Dvin village suburbs, Ararat district, 11.vi.2009, leg. Yuriy Shevnin, coll Danny Nilsson, g.prep. 0494/2019 D. Wanke; 1 ♂, Republic Armenia, Aiotzdorsky range, 2000 m, Yeghegnadzor suburbs, 150 km to SE from Yerevan, Mozrov Village, Mountain steppes, 12.vii.2009, leg. Yuriy Shevnin; 2 ♂, Iran, Prov. Esfahan, Zagros Mts, Fereidun Shar, 3000 m, 15.–17.vi.2010, leg. B. Benedek & T. Hácz, g.preps 0383, 0495/2019 D. Wanke; 1 ♂, NW-Iran, Provinz Chahar Mahal, ca. 35 km SE Aligudarz Seitental SW Murak, Umg. Dorf Darreh Hoz, 1800–2000 m, 13.–15.vi.2001, leg. de Freina, g.prep. 0384/2019 D. Wanke; 1 ♂, [Iran], Azarbeijan-e Gharbi, Takab, Takht-e soleiman, N36°35’40.0’’, E047°19’43.5’’, 2528 m, 1.vii.2013, leg. M. Afsarian, g.prep. 0385/2019 D. Wanke; 1 ♂, Iran, Zanjan, Zanjan-Gilvan, Gargovol Dag, 2500 m, 26./ 27.vi.2001, leg. A. Hofmann, J.-U. Meineke, W.G. Tremewan, g.prep. 0386/2019 D. Wanke; all in PCPS. 1 ♂, W-Iran, Lorestan, Dorud, Pass S Darrya-che Gahar, Partsche Kabud, 2800 m, 1.–3.viii.1975, leg. Ebert & Falkner, g.prep. 0195/2018 D. Wanke; 2 ♂, 1 ♀, W-Iran, Kordestan, Strasse Saghez-Baneh, 21 km NE Baneh, 1950 m, 30.[vi.]– 2.vii.1975, leg. Ebert & Falkner, g.preps (♂) 0196/2018 D. Wanke, (♀) 0487/2019 D. Wanke; 2 ♂, 1 ♀, Nordiran, Elbursgebirge östl. Shemshak, 50 km nördl. Teheran, 2100–2500 m, 8.–24.vi.1973, leg. G. Junge, g.preps (♂) 0209/2018, 0489/2019 D. Wanke; 1 ♂, NE-Iran, prov. Ostan-e Khorasan, Kopet Dagh, ca 50 km N Bojnurd, Sizmansufla, N 37°44’20’’, E 57°25’53’’, 10.v.2008, 1240 mNN, lux, leg. R. Trusch, M. Falkenberg & B. Müller, SMNK E-Lep. 247, g.prep. 0488/2019 D. Wanke; 1 ♂, N-Iran, Elburs-Mts., Prov. Tehran, 15km E Gatschsar, 2600 m, 5.viii.1972, leg. Ebert; 1 ♂, N-Iran, Elburs-Geb., 12 km v. Keregi, 1650 m, 27.v.1969, leg. G. Ebert, g.prep. 0490/2019 D. Wanke; 1 ♂, Persia s. [Iran], Elburs, Kendevan, ca. 3000 m, 3.– 9.vii. [year missing], coll. Wagner, Wien; all in SMNK. 3 ♂, 1 ♀, [Turkey], Kleinasien, Prov. Hakkari, 20km E Uludere, Suvarihalil-Pass, 2300–2600 m, 14.–15.vii.[19]85, leg. de Freina, g.preps (♂) 0465, 0466/2019 D. Wanke, (♀) 0467/2019 D. Wanke; 2 ♀, [Turkey], Kleinasien, Prov. Hakkari, Tanin-Tanin-Pass, 2000–2300 m, 12.–13.vii.[19]85, leg. de Freina, g.preps 0468, 0480/2019 D. Wanke; 1 ♂, 1 ♀, [Turkey], Kleinasien, Hakkari, Ö-Uludere, 1900 m, 28.–29.vi.1982, leg. W. Thomas, g.prep. (♀) 0469/2019 D. Wanke; 1 ♂, [Turkey], Kleinasien, Hakkari, Tanin-Tanin-Pass, 2000 m, 3.–5.vii.1983, leg. W. Thomas; 2 ♀, [Tur-key], Kleinasien, Prov. Artvin, NO-Anatolisches Randgebirge SE Seite, Barhal Tal, 4 km NE Altiparmak, 1100 m, 31.vii.–03.viii.[19]83, leg. de Freina; 1 ♂, Türkei, Prov. Van, Kuzgun Kiran-Gecidi, 1900 m, 21.vi.–8.vii.[19]81, ex. coll. A. Hofmann; 1 ♂, Türkei, Prov. Agri, 2km S Dogubayasit, 2100 m, 25.vii.[19]92, leg. P. Kautt & V. Weiss, g.prep. 0470/2019 D. Wanke; 1 ♂, 2 ♀, Iran, Ostan Tehran, Reshteh Ye Alborz, Dizin Gardaneh, 2700–3000 m, 5.–8.vii. 1978, leg. W. L. Blom, g.preps (♂) 0471/2019 D. Wanke, (♀) 0472/2019 D. Wanke; 1 ♂, 1 ♀, Iran, Elburz-Geb., Dizin, 2600 m, vii.1976, leg. Czipka; 1 ♂, Iran, Elburs, Shemshak, 2700 m, 10.–11.viii.1978, leg. W. Thomas, g.prep. 0473/2019 D. Wanke; 1 ♂, Iran, Elburs, Kendevan, 8.–15.viii.1978, 2300–2800 m, leg. W. Thomas; 1 ♀, Iran, Elburs, vic. Kendevan, 7.–9.viii.1977, 2500–3000 m, leg. W. Thomas, g.prep. 0474/2019 D. Wanke; 9 ♂, 2 ♀, Iran, Zanjan prov., E Zanjan, road to Gilvan, Alt. 1889 m, N36°45’21.8’’, E48°49’20.7’’, 6.vii.2013, leg. H. Rajaei, J.-U. Meineke, B. Hafezi, g.preps (♂) 0105, 0106/2018, 0475, 0476/2019 D. Wanke, (♀) 0477/2019 D. Wanke; 5 ♂, 2 ♀, Iran, Koh-kiluye va Boyerahmad, Yasuj, Sisakht, Dena, 2799 m, 30°57’23.6’’N, 51°23’28.9’’E, 30.vii.2016, leg. Sh. Feizpour, g.preps (♂) 0118/2018, 0484, 0485/2019 D. Wanke, (♀) 0486/2019 D. Wanke; 1 ♂, 4 ♀, Iran, Lorestan, Dorud, As-taneh door, 1801 m, 33°24’48.61’’N, 49°08’42.5’’E, 25.vii.2016, leg. Sh. Feizpour, g.preps (♂) 0123/2018 D. Wanke, (♀) 0478, 0496/2019 D. Wanke; 1 ♀, Iran, Azerbaijan-e Gharbi prov., Khoy to Ghotur road, Esteran vill., Alt. 1637 m, N 38°27’03.1’’, E 44°44’33.6’’, 1.vii.2013, leg. J.-U. Meineke, H. Rajaei, B. Hafezi, g.prep. 0299/2019 D. Wanke; 2 ♀, Iran, Shahrud, Shahkouh, Tash, Ayoub, Hosseini region, 2588 m, 36°37’18’’N, 54°33’42.6’’E, 11.vii.2016, leg. Sh. Feizpour, g.prep. 0481/2019 D. Wanke; 2 ♂, Iran, Prov. Golestan, Shahrud-Golestan road, Shahkuh, 2585 m, 36°38’36’’N, 54°31’31’’E, 16.vii.2015, leg. Sh. Feizpour, g.preps 0482, 0483/2019 D. Wanke; all in SMNS. 1 ♂, Iran, Khorassan, Kouh i Binaloud (Meched), 3000 m, 30.vii.1938, coll. Brandt, g.prep. 0503/2020 D. Wanke; in ZFMK. 1 ♂, 1 ♀, Iran, Fars, Strasse Chiraz-Kazeroun, Fort Sine-Sefid, ca. 2200 m, 26.v.1937, coll. Brandt, both labelled as ‘Paratypoid’; 2 ♂, Iran, Fars, Strasse Ardekan-Talochosroe [Ardakan-Talle Khosrow], Comè [Komehr] ca. 2600 m, 2.viii.1937, coll. Brandt; all in ZSM. Taxonomic remark on the subspecies. Based on the sympatric occurrence of the forms, absence of clear and constant morphological characters and evidence from the molecular data, most subspecies (except Synopsia phasidaria afghana comb. nov.) are treated as synonym to Synopsia phasidaria phasidaria comb. nov.. All subspecies and forms of S. phasidaria have been described based on small series of specimens, mainly referring to a different wing pattern and differences in the male genitalia (e.g. shape of the uncus, the number of cornuti and differences in the shape of the harpe). Nonetheless, in S. phasidaria all these characters are highly variable even within a single population, which has been stated also by different previous authors (Brandt 1938, Wiltshire 1966) and thus are inadequate as diagnostic characters (see fig. 36). Furthermore, we observed two kinds of bipectinate antennae in females (short or long pectination) (figs 1 H–I, 2), even in the same populations. However, these specimens with different shape of antennae do not show any correlated diagnostic differences in the wing pattern, genitalia characters, and DNA barcodes (fig. 41). All evidence therefore support the synonymy of these subspecies and lead us to conclude that the structurally different female antennae are intraspecific variation. Diagnosis. Wings and body (figs 8–22). Wingspan ♂ 26–38 mm, ♀ 32–42 mm, female specimens rarely smaller (forewing length ♂ 15–20 mm, ♀ 17–23 mm). Wing pattern highly variable, ground colour of wings from bright beige or sandy yellow with some brown spots, to brown (sandy yellow with some brown and orange spots in S. phasidaria afghana; similar in S. centralis; beige to brown, spotted dark in S. sociaria). Transverse lines faint or slightly visible (brown in S. phasidaria afghana and S. centralis; black in S. sociaria) (figs 5–7, 23–26). Male genitalia (figs 29–33, 36). Uncus sclerotized, slightly bifurcate or tip only concave in the centre (sclerotized tip concave in S. centralis; sclerotized and strongly bifurcate in S. sociaria). Gnathos well developed, arms fused, forming a tongue-shaped upturned plate, variable in shape (similar in S. centralis; almost similar in S. phasidaria afghana and S. sociaria). Costa of valva with central projection usually bearing one to three spines (similar in S. phasidaria afghana; without central projection in S. centralis; central projection bearing one to two spines in S. sociaria). Harpe sickle-shaped (similar in S. phasidaria afghana; thumb-shaped in S. centralis; without any harpe in S. sociaria). Aedeagus thick, bearing a small amount of cornuti, from 4 to 8 (5–6 cornuti in S. phasidaria afghana; 8 cornuti in S. centralis; vesica with two groups of cornuti, with a total amount of up to 25 spines in S. sociaria) (figs 27–28, 34–36). Female genitalia (figs 38–40). Apophyses anteriores about half the length of apophyses posteriores (similar in S. sociaria). Ductus bursae sclerotized, short and wide, with longitudinal foldings (wide and sclerotized, hardly any longitudinal foldings in S. sociaria). Corpus bursae, posteriorly sclerotized, anteriorly membranous (membranous in S. sociaria). Signum irregularly star-shaped (similar in S. sociaria, see fig. 37b). Phenology. Investigated specimens collected from May to September, suggesting that this species has more than one generation per year. Biology. Larva feeding on Apiaceae and Fabaceae (e.g. Trifolium sp.) (Wehrli 1954). Habitat. In altitudes from 1100 up to 3750 m. Distribution. Anatolian-Iranian, from eastern Turkey and southern Armenia, to western, northern and southeastern Iran. In Afghanistan rarely recorded, occurrence so far restricted to the type locality of the subspecies S. phasidaria afghana (see fig. 42). Genetic data. BIN: BOLD:AAC8929 (n=25). Nearest species: S. sociaria at a distance of only 3.5 %, which underpins the close phylogenetic relationship and thus the synonymy of Synopsidia with Synopsia. Genetic distance of Synopsia (sensu lato) from species of Palearctic related genera is much larger, i.e. Hypoxystis (represented by pluviaria) (7.1 %), Chariaspilates (represented by formosaria) (8.8 %), Angerona (represented by prunaria) (8.9 %), Siona (represented by lineata) (8.9 %) (fig. 41).Published as part of Wanke, Dominic, Hausmann, Axel, Sihvonen, Pasi, Krogmann, Lars & Rajaei, Hossein, 2020, Integrative taxonomic review of the genus Synopsia Hübner, 1825 in the Middle East (Lepidoptera: Geometridae: Ennominae), pp. 27-50 in Zootaxa 4885 (1) on pages 32-35, DOI: 10.11646/zootaxa.4885.1.2, http://zenodo.org/record/429633

    Phaselia smettboi Werner & Hausmann & Kostjuk & Wanke & Rajaei 2023, sp. nov.

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    P. smettboi sp. nov. vs. (P. joestleinae) External characters (27–32; 56–58). Forewing ground colour speckled with tiny light to dark grey-brown dots; antemedial line clearly zigzagged, all three teeth of similar length; subterminal line continuous, serrate with well-pronounced triangular spikes (ground colour with smooth dusty pattern; antemedial line irregularly zigzagged, costal tooth reduced to a barely visible curve; subterminal line interrupted in the middle, serrate with weakly pronounced triangular spikes). Male genitalia (figs 80–85; 107–108). Sacculus oval with many well-pronounced spines on the whole surface; aedeagus with a small claw-like cornutus (sacculus slightly oval, smoothly transitioning to valvula, without or with very small, fine spines on the sacculus; aedeagus with a strong, claw-like cornutus). Female genitalia (figs 125–127; 142–143). No diagnostic characters. Phenology. Possibly univoltine. Investigated specimens have been collected from early May to late July. Biology. Unknown. Habitat. Investigated specimens have been collected at altitudes from 150 m in Turkey (Bilecik) up to 1575 m in Turkey (Agri). Distribution. From Northwest Turkey over Southwest Russia and South Ural Mountains to West Kazakhstan (fig. 144). DNA barcoding. Genetic distances from morphologically most similar species: Phaselia serrularia (10.78%), P. pithana bona sp. (6.34%), P. phaeoleucaria stat. rev. (4.76%). Genetically closest species: P. phaeoleucaria stat. rev. (4.76%) and P. deliciosaria (4.93%) (fig. 145, Tab. 1). Etymology. The name smettboi originates from the famous 90s series ‘Pokémon’, which strongly influenced the childhood of the first author (MW) and that of her generation. Smettbo is the german name of a Pokémon (which is a small ‘pocket’ monster), that looks like a butterfly. With the slogan “Gotta Catch 'Em All”, the principle of Pokémon for finding all existing creatures, is a similar target in taxonomy.Published as part of Werner, Maria Johanna, Hausmann, Axel, Kostjuk, Igor, Wanke, Dominic & Rajaei, Hossein, 2023, Integrative taxonomic revision of the genus Phaselia Guenée, [1858] (Geometridae: Ennominae) in the Middle East and Central Asia, pp. 1-66 in Zootaxa 5326 (1) on pages 16-17, DOI: 10.11646/zootaxa.5326.1.1, http://zenodo.org/record/824381
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