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    A New Thai Enhydris (Serpentes: Colubridae: Homalopsinae)

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    Murphy, John C., Voris, Harold K. (2005): A New Thai Enhydris (Serpentes: Colubridae: Homalopsinae). Raffles Bulletin of Zoology 53 (1): 143-147, DOI: http://doi.org/10.5281/zenodo.461902

    A New Species Of Enhydris (Serpentes: Colubridae: Homalopsinae) From The Kapuas River System, West Kalimantan, Indonesia

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    Murphy, John C., Voris, Harold K., Auliya, Mark (2005): A New Species Of Enhydris (Serpentes: Colubridae: Homalopsinae) From The Kapuas River System, West Kalimantan, Indonesia. Raffles Bulletin of Zoology 53 (2): 271-275, DOI: http://doi.org/10.5281/zenodo.461918

    FIGURE 4 in The dog-faced water snakes, a revision of the genus Cerberus Cuvier, (Squamata, Serpentes, Homalopsidae), with the description of a new species

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    FIGURE 4. The Australian Bockadam, Cerberus australis. Two color morphs from the Northern Territory, vicinity of Darwin. JCM.Published as part of Murphy, John C., Voris, Harold K. & Karns, Daryl R., 2012, The dog-faced water snakes, a revision of the genus Cerberus Cuvier, (Squamata, Serpentes, Homalopsidae), with the description of a new species, pp. 1-34 in Zootaxa 3484 on page 10, DOI: 10.5281/zenodo.28237

    FIGURE 4 in The masked water snakes of the genus Homalopsis Kuhl & van Hasselt, 1822 (Squamata, Serpentes, Homalopsidae), with the description of a new species

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    FIGURE 4. Box whisker graphs comparing ventral and subcaudal scale count data for Homalopsis by gender. A. Among both males and females the only significant differences in ventral scale counts (p <0.05) are between H. mereljcoxi sp. nov. and the other three species. B. Among both males and females the only significant differences in subcaudal scale counts (p <0.05) are between H. semizonata and H. mereljcoxi sp. nov..Published as part of Murphy, John C., Voris, Harold K., Traub, Joshua & Cumberbatch, Christina, 2012, The masked water snakes of the genus Homalopsis Kuhl & van Hasselt, 1822 (Squamata, Serpentes, Homalopsidae), with the description of a new species, pp. 1-26 in Zootaxa 3208 on page 8, DOI: 10.5281/zenodo.20995

    FIGURE 2 in Phylogeny and biogeography of the Enhydris clade (Serpentes: Homalopsidae)

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    FIGURE 2. Geographic distribution of the species of the Enhydris clade in South and Southeast Asia. Homalopsid snakes are generally found in lowland habitats at elevations of less than 200 m. These maps are based on a combination of verified published localities (Murphy 2007) and adjacent contiguous lowland (<200 m) topography. Note that due to the size of the maps some higher elevations in southern Myanmar, the Thai-Malay Peninsula, Sumatra, Java, and Vietnam appear shaded on the Enhydris enhydris map even though these mountains are not inhabited by E. enhydris.Published as part of Karns, Daryl R., Lukoschek, Vimoksalehi, Osterhage, Jennifer, Murphy, John C. & Voris, Harold K., 2010, Phylogeny and biogeography of the Enhydris clade (Serpentes: Homalopsidae), pp. 18-30 in Zootaxa 2452 (1) on page 24, DOI: 10.11646/zootaxa.2452.1.2, http://zenodo.org/record/530691

    The terrestrial life of sea kraits : insights from a long-term study on two Laticauda species (Reptilia: Squamata: Elapidae) in the Andaman Islands, India

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    CITATION: Tyabji, Z. et al. 2018. The terrestrial life of sea kraits : insights from a long-term study on two Laticauda species (Reptilia: Squamata: Elapidae) in the Andaman Islands, India. Journal of Threatened Taxa, 10(11):12443–12450, doi:10.11609/jott.4311.10.11.12443-12450.The original publication is available at https://threatenedtaxa.orgSea kraits forage in water and return to land to digest their prey, mate, slough, and lay their eggs. The temporal terrestrial patterns in encounter rate and behaviour of two species of sea kraits Laticauda colubrina and L. laticaudata were studied over four years at the New Wandoor beach in the southern Andaman Islands. The encounter rate of L. colubrina was found to be 20 times higher than L. laticaudata, and sea kraits were observed to prefer the natural refuge that the microhabitat of uprooted trees provide. Additionally, nesting observations are presented that emphasize the need to promote the conservation of these crucial terrestrial habitats.Andaman Nicobar Environment Teamhttps://threatenedtaxa.org/index.php/JoTT/article/view/4311Publisher's versio

    Homalopsis Kuhl & Hasselt 1822

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    Homalopsis Kuhl & Hasselt, 1822 Coluber Linnaeus, 1758: 217. Vipera Daudin 1803 a: 220. Homalopsis Kuhl & Hasslet 1822: 101 Pythonia Blyth 1859: 279 Pythonella Theobald 1868: 66 (lapsus for Pythonia fide Williams & Wallach, 1989) Type species. Coluber horridus Daudin, 1803 a (synonym of Coluber buccatus Linnaeus, 1758) by original designation. Content. Five species: Homalopsis buccata, H. hardwickii, H. nigroventralis, H. mereljcoxi sp. nov., and H. semizonata. Diagnosis. Homalopsis can be distinguished from all other Southeast Asian snakes by their keeled and striated scales in 33–49 rows at midbody; crescent-shaped valvular nostrils; lower labials posterior to the eye are horizontally divided; and enlarged plates (frontal and parietals) on the crown. Snakes of the genus Cerberus are the species most likely to be confused with Homalopsis; however, Cerberus have fewer than 30 scales rows at midbody and the frontal and parietals are ‘fragmented’ into small irregular scales. Distribution. Homalopsis ranges from Nepal (based on an anecdotal record, Zug & Mitchell 1995) and probably northeast India, eastward to Indochina and southward into the Malayan peninsula and the Indonesian Archipelago as far east as Borneo. One specimen has been reported from Makassar, Sulawesi (Rooji 1917) but De Lang &Vogel (2005) could not validate its presence on the island. Figure 5 shows the distribution of the five species recognized here. Regional works on India and Nepal rarely mention the presence of Homalopsis. Neither Ahmed et al. (2009) nor Whitaker & Captain (2004) mention the species in India and only Schleich & Kastle (2002) report it from Nepal, and their account is based solely upon the Zug & Mitchell (1995) comment.Published as part of Murphy, John C., Voris, Harold K., Traub, Joshua & Cumberbatch, Christina, 2012, The masked water snakes of the genus Homalopsis Kuhl & van Hasselt, 1822 (Squamata, Serpentes, Homalopsidae), with the description of a new species, pp. 1-26 in Zootaxa 3208 on page 4, DOI: 10.5281/zenodo.20995

    Homalopsis hardwickii Gray 1842

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    Homalopsis hardwickii Gray, 1842 Fig. 1 b Homalopsis buccata— Günther 1864: 285; Boulenger 1890: 374; 1896: 14; Smith 1943: 390. Wall 1903: 94; Wall 1923: 38; Gyi 1970: 136; Murphy 2007: 193 Type locality. “ India.” Holotype. BMNH 1946.1.7.26.” Distribution. Possibly restricted to northeast India, no known extant population. The holotype of Homalopsis hardwickii is reported to be from India, but the datum is questionable. Two other specimens (BMNH 111.18. 1 b, g) that may be H. hardwickii have the locality of “Bengal” but do not agree well with the holotype and the locality information is followed by a question mark in the BMNH catalog. Thus, this species remains to be re-discovered and its distribution to be determined. Diagnosis. Homalopsis hardwickii has a divided loreal contacting upper labials 1–4; two prefrontal scales; 39 scale rows at midbody reduced to 28 posteriorly; one postocular scale and no presubocular scale; 13 or 14 upper labials; 159 ventrals. Homalopsis buccata has a single loreal contacting upper labials 1–4; one postocular scale plus a postsubocular scale; H. nigroventralis has upper labials 1–3 contacting the loreal; one or two postocular scales plus one postsubocular scale; and a reverse color pattern on the venter (dark olive-gray with white spots). H. semizonata has three prefrontals; one postocular and one postsubocular. The new species, H. mereljcoxi, has a single loreal contacting upper labials 1–4; 40–49 scale rows at midbody, reduced to 30 or more posteriorly; and two postocular scales plus a postsubocular. Redescription of the Holotype. A male, 511 mm SVL, 172 mm tail; dorsal scale rows 41 – 39 – 28; 159 ventrals; 84 subcaudals; 1 / 1 preocular, 1 / 1 postocular, 1 / 1 postsubocular, no subocular; internasal small and divided; 2 / 2 prefrontals; temporal formula 1 / 1; upper labials 14 / 13, first six not horizontally divided; upper labials under orbit 5 / 6; upper labials in contact with loreal 1-4 / 1-4; loreal divided on both sides. It has a typical Homalopsis pattern with alternating dark brown separated by white saddle-like blotches, the light blotches number 44, of which 19 do not extend to the vertebral line. Material examined. “ INDIA ” BMNH 1946.1. 7.26 (type).Published as part of Murphy, John C., Voris, Harold K., Traub, Joshua & Cumberbatch, Christina, 2012, The masked water snakes of the genus Homalopsis Kuhl & van Hasselt, 1822 (Squamata, Serpentes, Homalopsidae), with the description of a new species, pp. 1-26 in Zootaxa 3208 on pages 12-13, DOI: 10.5281/zenodo.20995

    Cerberus Cuvier 1829

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    Cerberus Cuvier 1829 Hydrus Schneider 1799 1: 233246 (in part). Enhydris Sonnini and Latreille 1801 (in part), 4: 202. Elaps Schneider, 1801 (in part), 2: 301. Coluber (non-Linneaus, 1766) Daudin, 7: 167. Python Merrem, 1820: 89. Daudin used the combination Python rhynchops. Cerberus Cuvier, 1829, 2, 2: 81. Type species Coluber cerberus = Hydrus rynchops Schneider. Note that Swainson (1839) spelled this Cerebus, a lapsus. Homalopsis Schlegel, 1837 (in part), 2: 332. Identification. Distinguishing Cerberus species on the basis of morphology alone can be difficult which in part accounts for the fact that C. rynchops has been described 11 separate times since 1799. Table 2 summarizes some of the morphological data discussed here that can be used to separate these taxa. The variable dorsal scale row counts within and between species, the fragmentation of head shields, and the relatively similar coloration and pattern, make these species somewhat cryptic. This can be highlighted by Loveridge’s (1948) confusion when he named the New Guinea population Cerberus rynchops novaeguineae instead of recognizing its morphology as more similar to australis; and Cogger et al. (1983) synonymizing Cerberus australis and Cerberus rynchops, two species that differ genetically by 6.4–9.5% (Alfaro et al. 2004). The five species recognized here are morphologically distinct, with the differences often subtle but consistent. Four of these species have been previously compared using mitochondrial and nuclear genes (Alfaro et al. 2004, 2008). Diagnosis. Cerberus can be distinguished from all other homalopsid snakes with nasal scales in contact by: keeled, striated dorsal scales; absence of rostral appendages; and parietal scales that are partially or completely fragmented. It differs from its sister, Homalopsis, in having fewer scale rows (21–31 at midbody in Cerberus, 33– 49 rows in Homalopsis) and fragmented parietal scales. Most specimens are less than one meter, and the maximum size for most species is less than 1.2 m. The tail is 16–29 % of the snout-vent length (SVL). Females are larger than males in the species where both sexes could be examined. Head plates (prefrontal, frontal, and parietals) show a strong tendency to fragment into smaller scales. One or more large posterior upper labials are horizontally divided. Subocular scales may or may not be present, even within a single population. Temporal scales are small and not plate-like. Dorsal scale rows at midbody range from 21–31; however most specimens of most species have 23, 25, or 29 rows at midbody. Dorsal scale ornamentation consists of a strongly striated surface with central keel when viewed with a binocular light microscope with 10 x magnification, and some individuals have tubercles on head scales. Ventral scales range from 134–172 and subcaudal scales range from 42–75. Anterior maxillary teeth number 12–19 and are followed by a diastema and a pair of deeply grooved fangs. Three pairs of chin shields are present and the first pair is usually the largest. Content: Here we recognize 5 species: C. australis (Gray, 1849); C. dunsoni sp. n.; C. microlepis Boulenger, 1896; C. rynchops (Schneider, 1799), and C. schneiderii (Schlegel, 1837) new combination.Published as part of Murphy, John C., Voris, Harold K. & Karns, Daryl R., 2012, The dog-faced water snakes, a revision of the genus Cerberus Cuvier, (Squamata, Serpentes, Homalopsidae), with the description of a new species, pp. 1-34 in Zootaxa 3484 on pages 7-8, DOI: 10.5281/zenodo.28237
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