119,726 research outputs found
Djeboa compressa K. Viets 1914
Djeboa compressa K. Viets, 1914 (Figs. 12 A–F, 37 E) Djeboa multidentata compressa K. Viets 1914: 220. Mideopsis (Djeboa) compressa Cook 1966: 231. Material examined: Type series: SMF, Cameroon; lectotype, here designated, K. Viets 43190, “ Kamerun, Manoka b. Duala, Bach, 11.11. 1911, Damköhler leg. 1270 ”; paralectotype female, K. Viets 43191, same data as holotype, “ 1271 ”. General features: Dorsal shield oval (L/W ratio 1.1), with medial depression; muscle scars anterior and posterior to postocularia with moderately pronounced thickenings, second pair of muscle scars located well within the dorsal depression; central portion of dorsal shield purple (Fig. 37 E); gnathosomal bay V-shaped; tips of Cx-I not reaching frontal margin; medial margins of Cx-IV reduced to a median angle, approaching to each other, posterior margin of Cx-IV extending almost directly laterally. Palp (Fig. 12 D): P- 1 with a dorsal seta; P- 2 inflated, with straight ventral and convexly bowed dorsal margin; P- 3 ventral margin concave, dorsal margin usually with a pronounced indentation slightly distal to middle of the segment; P- 4 slender, basally thickened, from here to anterior tip narrowed. Legs: I-L (Fig. 12 E) with I-L- 6 L/H ratio 2.7, distally thickened; IV-L: Fig. 12 F. Discussion: Along with D. expansipalpis and D. gledhilli sp. nov., both from Côte d’Ivoire, and D. globulipalpis sp. nov. from South Africa, D. compressa forms a species group characterized by the V-shaped gnathosomal bay and P- 4 much longer than high. Distribution: Cameroon (Viets 1914), Liberia (Cook 1966).Published as part of Pešić, Vladimir, Cook, David, Gerecke, Reinhard & Smit, Harry, 2013, The water mite family Mideopsidae (Acari: Hydrachnidia): a contribution to the diversity in the Afrotropical region and taxonomic changes above species level, pp. 1-75 in Zootaxa 3720 (1) on page 25, DOI: 10.11646/zootaxa.3720.1.1, http://zenodo.org/record/28502
Djeboa multidentata K. Viets 1911
Djeboa multidentata K. Viets, 1911 (Figs. 41 A–C) Djeboa multidentata K. Viets, 1911: 494. Material examined: Type series: SMF, Cameroon; holotype male, K. Viets 43185; “ Kamerun, Bach bei Djebo, 6.6. 1911, Damköhler leg., “ 1011 ”; paratype female, K. Viets 43187 “Cotype, Kamerun, Joh. Albrechts-Höhe, Bach am Wege Kumba Mundame, 2.10. 1911, Damköhler leg. 1171 “. Further material: SMF K. Viets 43203, “ Kamerun, Ndvab-Fluß, 6.6.1913, 5555”, 1 /0/0; K. Viets 43204, “Nyassa, Nkata-Bay, Fisch-Magen, Fryer coll. 7753 ”, 0/ 1 /0. General features: Dorsal shield oval (1.0– 1.2), with medial depression; muscle scars with little pronounced thickenings, located anterior and posterior to the postocularia; colour pattern consisting of an anterior violet patch (Figs. 41 A–C); gnathosomal bay Y-shaped, noticeably narrowing in posterior half; tips of Cx-I ending posterior to frontal margin; medial margin of Cx-IV reduced to a median angle; Cx-III and -IV with a series of longitudinal striae (four or five pairs on Cx-IV). Palp: P- 1 with a dorsal seta; P- 2 with concave ventral and convexly bowed dorsal margin; P- 3 proximally thicker than distally, ventral margin slightly concave; P- 4 slender, equally narrowing from the base to tip. Legs: I-L- 6 L/H ratio 2.6, ventral margin strongly protruding. Remarks: The original description of Djeboa multidentata was mostly based on a female from “Bach b. Djebo, 6.6.1911 ” while the male (same site and date) was described in a few lines and without figures (Viets 1911). An examination of material of Djeboa multidentata from the K. Viets collection shows that the female of slide 43204 surely does not belong to D. multidentata. The female on slide SMF 43187 labeled as cotype of D. multidentata agrees with the original description but as the palp is in situ and many characters are not visible, it is surely not the specimen described by K. Viets. Distribution: Cameroon.Published as part of Pešić, Vladimir, Cook, David, Gerecke, Reinhard & Smit, Harry, 2013, The water mite family Mideopsidae (Acari: Hydrachnidia): a contribution to the diversity in the Afrotropical region and taxonomic changes above species level, pp. 1-75 in Zootaxa 3720 (1) on pages 49-51, DOI: 10.11646/zootaxa.3720.1.1, http://zenodo.org/record/28502
Arrenurus consanguineus K. Viets 1954
Arrenurus consanguineus K. Viets, 1954 Arrenurus gladiiferus Rosso de Ferradás, 2006b (erron., non Lundblad, 1944) (Figure 3) Material examined— Bolivia. 1/5/0, pond Hotel Flora y Fauna, 3 km SE of Buena Vista, 17°29.949 S 63°38.152 W, 10-iii-1999, leg. D. Tempelman. Description — Male: Idiosoma dorsally 616 long and 454 wide, ventrally 648 long. Dorsal shield incomplete (but dorsal furrow continues ventrally); dorsal shield 340 wide, distance dorsal shield – anterior idiosoma margin 109. Medial margin of fourth coxae longer than medial margin of third coxae. Gonopore 42 long, genital field 365 wide, rather broad. Length of P1-5: 30, 64, 40, 60, 32. P2 with 7-8 large, sword-like setae. Remarks — My male specimen matches the description of K. Viets (1954b). Viets didn’t illustrate the setae associated with Dgl-2 and -3, but these setae are very small and easily overlooked. The specimen reported from Argentina by Rosso de Ferradás (2006b) A as. gladiiferus doesn’t belong to that species but A to. consanguineus. Arrenurus gladiiferus has a convex posterior margin of the cauda without an indentation, and the setae of P2 are gradually tapering and not sword-like as in A. consanguineus. Distribution — Previously known from Brazil (K. Viets 1954b) and Argentina (Rosso de Ferradás 2006b, as A. gladiiferus).Published as part of Smit, Harry, 2020, New records of the water mite genus Arrenurus Dugès, 1834 from South America (Acari: Hydrachnidia: Arrenuridae), with the description of five new species and one new subspecies, pp. 371-389 in Acarologia 60 (2) on pages 374-377, DOI: 10.24349/acarologia/20204374, http://zenodo.org/record/448788
Mideopsellides geae K.O. Viets 1962
Mideopsellides geae K.O. Viets, 1962 (Figs. 41 D–E, 42 A–F) New records: South Africa: DC 102 84 Eastern Cape Province, Kaaimans River E of George, 5–10 m wide, sand & rocks, stained, good flow (9.5 ˚C), 10.vi. 1984 Cook 7 / 5 /0 (mounted); West Cape Province, Bloukrans River at crossing with Road R 102, 15.xi. 2004 Smit 1 /0/0. Distribution: South Africa (Eastern Cape Province—K.O. Viets 1962; present study).Published as part of Pešić, Vladimir, Cook, David, Gerecke, Reinhard & Smit, Harry, 2013, The water mite family Mideopsidae (Acari: Hydrachnidia): a contribution to the diversity in the Afrotropical region and taxonomic changes above species level, pp. 1-75 in Zootaxa 3720 (1) on page 64, DOI: 10.11646/zootaxa.3720.1.1, http://zenodo.org/record/28502
Lobohalacarus Viets 1939
Lobohalacarus Viets, 1939 Type species. Walterella weberi Romijn & Viets, 1924. Female. All ventral plates fused, AP included. Area of GP with two to seven pairs of slender pgs, mostly four or five pairs of setae present. Female GO in posterior part of GP; each genital sclerite with zero to three sgs and zero to three external gac; no acetabula outside genital sclerites. Ovipositor short, at rest anteriad not extending beyond GO. Everted ovipositor showing a basal pair of small spines and five apical pairs of sclerotized genital spines (Bartsch 1975 d: fig. 51). AE with pair of epimeral pores. Male. Not known. Juveniles. With deutonymph, protonymph and larva. In both deutonymph and protonymph genital and anal plate fused to a genitoanal plate, in larvae genital plate lacking. Deutonymph with two pairs of gac; single pair of pgs in anterior part of GA; sgs lacking (Bartsch 1975 d: fig. 50). Protonymph with single pair of gac; without pgs and sgs. All juveniles with pair of epimeral pores. Remarks. Two Lobohalacarus species are known, one with several subspecies (Bartsch 2008 a). Lobohalacarus is a cosmopolitan genus, it lives in freshwater, is regularly found in groundwater-fed habitats, but also in surface waters, even in coastal brackish water.Published as part of Bartsch, Ilse, 2015, The genital area of Halacaridae (Acari), life stages and development of morphological characters and implication on the classification, pp. 201-259 in Zootaxa 3919 (2) on page 222, DOI: 10.11646/zootaxa.3919.2.1, http://zenodo.org/record/24524
Revisional studies on<i>Thoracophoracarus</i>K. Viets, 1914 (Arachnida, Acari, Hydrachnidia, Arrenuridae)
FIG. 3. — Thoracophoracarus kuehnei K. Viets, 1916: A-C, paratype ♂ (SMF 55029); A, ventral idiosoma; B, dorsal idiosoma; C, palp; D, paratype ♀ (SMF 47466), ventral idiosoma. Scale bars: 100 μm.Published as part of Gerecke, Reinhard, 2009, Revisional studies on Thoracophoracarus K. Viets, 1914 (Arachnida, Acari, Hydrachnidia, Arrenuridae), pp. 127-145 in Zoosystema 31 (1) on page 132, DOI: 10.5252/z2009n1a7, http://zenodo.org/record/539793
Albia (Anchistalbia) tenuipalpis Viets 1911
Albia (Anchistalbia) tenuipalpis Viets, 1911 (Figs. 1 B–D) Material examined. Ghana. 1/0/0, Stream Asukuma, Kakum NP, 5º 26.995 N 1º 25.071 W, alt. 131 m a.s.l., 12-ii- 2013. Description. Male: Idiosoma yellowish with in the centre a large purple spot, dorsally 640 long and 470 wide, ventrally 559 long. Dorsal and ventral shields present. Dorsal shield 429 wide, with five pairs of glandularia. First coxae not extending to anterior idiosoma margin. Suture lines between Cx-I and Cx-II and midline distinct, suture line between Cx-III and Cx-IV indistinct, suture lines of Cx-II and Cx-III obliterated. Suture lines of Cx-III/IV fused with midline anterior to fusion point of CX-I/II. Cxgl–4 close to suture line between Cx-III and Cx-IV. Genital field with numerous acetabula, gonopore 56 long. Length of P1–5: 38, 64, 47, 86, 36. P4 bowed. Length of I-leg-4–6: 72, 82, 70. Length of IV-leg-4–6: 98, 104, 90. Second, third and fourth legs with numerous swimming setae. Remarks. Cook (1966) reported the species from Liberia, but his specimen has a different configuration of the dorsal glandularia compared to the male from Ghana, and has an anterior and posterior blue spot. The male described by Viets (1916) has a blue spot in the centre, but is larger (735 µm) than the male from Ghana. Unfortunately Viets (1916) didn’t illustrate the dorsum, and his illustration of the venter is sketchy. The palp of Viets’ his male agrees well with the palp of the male from Ghana, which is less elongated compared to the female. Albia tenuipalpis is known from Cameroon, South Africa, possibly from Liberia and reported here for the first time from Ghana.Published as part of Smit, Harry, 2016, The water mite family Aturidae Thor (Acari: Hydrachnidia) from Ghana, with the description of twelve new species, pp. 523-543 in Zootaxa 4158 (4) on page 525, DOI: 10.11646/zootaxa.4158.4.5, http://zenodo.org/record/25747
Halacarellus Viets 1927
Halacarellus Viets, 1927 (Figs 44–46) Type species. Halacarus balticus Lohmann, 1889. Adults. Female and male with GA. Female GA with two to four (rarely seven) pairs of pgs, genital sclerites with zero to five pairs of sgs. Three pairs of internal acetabula always distinct, in freshwater inhabiting species enlarged (Viets 1928 a: fig. 49; Bartsch & Gerecke 2011: fig. 1 H). Acetabula arranged in line along genital slit, anterior pair anterior or posterior to level of mid-GO (Fig. 44; Otto 1994: fig. 27). Ovipositor at rest often extending beyond GO. Ovipositor with two basal and eight or ten apical genital spines. Basal pair of genital spines papilli- or setiform. Apical genital spines dissimilar in shape, robust and with tines, slender and spiniform, or reduced to small papillae (Figs 45, 46; Womersley 1937: pl. 6, fig. 12; Bartsch 1998 b: figs 31, 61, 2007 b: fig. 6 G). Male GA with 10–50 pairs of pgs and in general five pairs of sgs (rarely less). In the majority of species all three pairs of acetabula internal, arranged along genital slit; in a few species (of the H. capuzinus group) three pairs of acetabula arranged in posterior part of GO, within a fovea that opens to the exterior (Bartsch 1998 b: fig. 48). In freshwater species two pairs of gac posterior to GO, enlarged and external, one pair of gac small, near middle of GO (Viets 1928 a: fig. 48; Bartsch 1998 b: fig. 15). AE without epimeral pores. Juveniles. With larva and two nymphal stages. Nymphs of northern hemisphere species with genital and anal plate fused to a GA (e.g. in the type species H. balticus), most southern hemisphere species with GP and AP separated. Deutonymphal GA with two pairs of pgs, zero to two pairs of sgs and two pairs of gac (Newell 1951: figs 3, 19; Bartsch 1972: figs 25 B, 28 B, 1995 a: fig. 19, 1998 b: figs 24, 41, 63, 82, 101, 1990 c: fig. 137, 1999 c: fig. 3 F). Acetabula of nymphs small and internal, large and internal or large and external (e.g. in H. hyrcanus (Viets, 1928) and H. fontinalis Bartsch & Gerecke 2011 (Bartsch 1998 b: fig. 24; Bartsch & Gerecke 2011: fig. 2 G). Protonymphal GA generally with pair of internal gac; pgs and sgs lacking (Bartsch 1972: figs 25 D, 28 F, 1995 a: fig. 11, 1998 b: figs 44, 85, 1999 c: fig. 3 G). Larva with pair of epimeral pores on AE (Bartsch & Panesar 2000: fig. 3). Larval anal plate often with large ventral part (Bartsch 1972: fig. 25 F, 1998 b: figs 67, 88). Remarks. The genus, with about 50 species, is spread world-wide (Bartsch 2009 a). Several of the species can be assigned to distinct species groups (Bartsch 1997 a). Representatives of the northern Atlantic Ocean groups are not found on the southern hemisphere and vice versa (Bartsch 1997 a).Published as part of Bartsch, Ilse, 2015, The genital area of Halacaridae (Acari), life stages and development of morphological characters and implication on the classification, pp. 201-259 in Zootaxa 3919 (2) on page 218, DOI: 10.11646/zootaxa.3919.2.1, http://zenodo.org/record/24524
Atractides damkoehleri K. Viets 1916
Atractides damkoehleri (K. Viets, 1916) (Figs. 1–5) Megapus damkoehleri K. Viets 1916: 271. Syn. A. ugandensis (Lundblad, 1927), A. duricorius (Walter & Bader, 1952): K.O. Viets 1964 a. Material examined. South Africa, Western Cape province: SA 1 0/ 3 / 1; SA 3 0/ 1 /0; SA 4 0/ 1 /0; SA 5 1 /0/0; SA 6 1 / 1 /0; SA 8 1 /0/0; SA 10 0/ 2 /0; DC 107 84 2 / 1 /0; DC 109 84 1 /0/0. KwaZulu-Natal province: SA 12 0/ 3 /0 (0/ 1 /0 mounted); SA 13 6 / 9 /0; SA 15 1 / 6 /0; SA 16 7 / 4 /0; SA 17 1 /0/0; SA 18 13 / 11 / 1; SA 19 1 / 4 /0; SA 20 8 / 10 /0 (1 / 1 /0 mounted); SA 21 0/ 2 /0; SA 22 1 / 7 /0; SA 23 0/ 1 /0; SA 24 1 / 2 /0; DC 89 84 11 / 11 /0; DC 90 84 1 /0/0; DC 91 84 0/ 1 / 0; DC 92 84 8 / 4 /0; DC 93 84 1 /0/0. Limpopo province: SA 25 7 / 2 /0. Mpumalanga province: DC 76 84 0/ 1 /0; DC 78 84 5 / 13 / 1 (0/ 1 / 1 mounted); DC 83 84 0/ 2 /0; DC 84 84 13 / 27 /0; DC 85 84 1 / 5 /0 (1 / 1 /0 mounted); DC 86 84 5 / 4 / 0. General features. Dorsal integument striated; muscle attachments unsclerotized; glandularia enlarged by secondary sclerotization. Coxal field: posterior margin Cx-I strongly convex, apodemes of Cx-II directed laterally. Genital field: Ac in triangular position. Excretory pore smooth; Vgl- 1 fused to Vgl- 2. Palp without sexual dimorphism, P- 2 and - 3 ventral margin straight, P- 4 slender, sword seta slender, halfway between ventral hairs. I-L- 5 proximally subrectangular, distally protruding near insertion S- 1, S- 1 and S- 2 interspaced, S- 1 obtusely truncate, not thickened distally, S- 2 basally enlarged, bluntly pointed; I-L- 6 stout, curved, inflated proximally, from the centre to the claw furrow with parallel dorsal and ventral margins; leg claw without dorsal clawlets. Morphology. Male (SA 20, in parentheses DC 85 84): Idiosoma L/W 509 (444)/ 428 (322); glandularia maximum diameter 37 (34); coxal field L 288 (286); Cx-III W 317 (307); Cx-I+II mL 123 (102), lL 200 (191). Genital field (Fig. 1 A, 3 A): anterior margin with a border of secondary sclerotization convexely protruding, posteriorly indented; L/W 94 (88)/ 116 (101); L Ac 1-3: 35 (27), 39 (29), 42 (30). Palp (Figs. 1 C, 4 B): total L 255 (254), dL: P-1, 20 (25); P-2, 55 (52); P-3, 66 (65); P-4, 85 (84); P-5, 29 (28); L ratio P- 2 /P- 4, 0.65 (0.62); capitulum vL 104 (86); chelicera total L (163). I-L (Figs. 1 B, 4 A): I-L- 5 dL 153 (152), vL 112 (106), dL/vL ratio 1.37 (1.43), HB 48 (46), dL/HB 3.2 (3.3), S- 1 L 79 (77), L/ W 9.2 (8.4), S- 2 L 69 (71), L/ W 6.0 (5.6), distance S- 1 -2, 15 (10), L ratio S- 1 / 2, 1.15 (1.09); I-L- 6 L 106 (107), HB 22 (22), L/HB ratio 4.75 (5.0); L ratio I-L- 5 / 6, 1.44 (1.42). Female (SA 20, in parentheses SA 12, in square parentheses DC 85 84). Idiosoma L/W 669 (756)/ 544 (641); glandularia maximum diameter 42 (43) [29]; coxal field L 356 (369) [311]; Cx-III W 408 (464) [367]; Cx-I+II mL 122 [119], lL 228 [231]. Genital field (Figs. 2 B, 3 B): L/W 171 (199) [140]/ 162 (188) [147]; pregen prominent, curved, W 110 (115) [55]; genital plate L 122–123 (130) [109]; L Ac 1–3: 48–49 (54–57) [39–40], 57–60 (52–54) [43–45], 34–45 (49) [35–39]. Palp (Figs. 1 D, 4 D): total L 333 (358) [347], dL: P-1, 30 (34) [34]; P-2, 69 (77) [77]; P-3, 93 (99) [95]; P- 4, 106 (109) [103]; P-5, 35 (39) [38]; L ratio P- 2 /P- 4, 0.65 (0.71) [0.75]; capitulum vL 128 (141); chelicera total L 219 (250). I-L (Figs. 2 A, 4 C): I-L- 5 dL 215 (237) [201], vL 149 (172) [148], dL/vL ratio 1.44 (1.38) [1.36], HB 62 (72) [58], dL/HB 3.5 (3.3) [3.5], S- 1 L 111 (109) [96], L/ W 9.0 (8.9) [7.8], S- 2 L 91 (91) [80], L/ W 5.4 (6.0) [5.8], distance S- 1 -2, 22 (22) [14], L ratio S- 1 / 2, 1.22 (1.2) [1.2]; I-L- 6 L 149 (163) [137], HB 22 (22) [22], L/HB ratio 6.8 (7.4) [6.2]; L ratio I-L- 5 / 6, 1.44 (1.45) [1.47]. Deutonymph (collected together with adults of A. damkoehleri at DC 78 84 and suspected to represent this species.): Integument dorsally striated; muscle attachments unsclerotized; glandularia maximum diameter 20; Cx-III W 192; Cx-I+II mL 57, lL 114; provisional genital field with two pairs of Ac, L/W 58 / 75; excretory pore smooth; Vgl- 1 fused to Vgl- 2 (Fig. 5 A). Palp: total L 154, dL: P-1, 15; P-2, 32; P-3, 39; P-4, 46; P-5, 22; L ratio P- 2 /P- 4, 0.7. I-L (Fig. 5 B): I-L- 5 dL 89, vL 63, dL/vL ratio 1.4, HB 30, dL/HB 3.0, S- 1 L 52, L/ W 11.3, S- 2 L 45, L/ W 7.3, distance S- 1-2, 8.0, L ratio S- 1 / 2, 1.16; I-L- 6 L 69, HB 17, L/HB ratio 4.1; L ratio I-L- 5 / 6, 1.3. Remarks. The specimens examined from South Africa fit the descriptions of Atractides ugandensis (Lundblad, 1927) and A. duricorius (Walter & Bader, 1952), both species described from Eastern Africa and later synonymized with A. damkoehleri (see: K. O. Viets, 1964 a). The measurements of the specimens examined lie within the variability range of A. damkoehleri given by K.O.Viets (1964 a) for populations from Eastern Transvaal. Atractides damkoehleri closely resembles two poorly known species from South Africa, A. sudafricanus K. Viets, 1956 and A. coriacellus K. Viets, 1956. K. Viets (1956) described Atractides sudafricanus based on a single juvenile female specimen from Transvaal (without exact locality), providing only figures of the venter and palp, and without discussion of its similarity with other species. Later on, K.O. Viets (1964 a) reported one female and one deutonymph of this species from Keurbooms River in the Eastern Cape and noted that additional material was needed to clarify the taxonomy of this species. He mentioned similarities in the shape of I-L- 5 and - 6 (S- 1 obtusely truncate) with A. damkoehleri, but A. sudafricanus differs from the latter species in the shape of the coxae (Cx-IV medial margin angled, probably due to secondary sclerotization), the genital field (Ac in a weakly curved line) and palp (P- 4 sword seta large, nearer to proximoventral hair). The measurements given in the original description of A. sudafricanus lie within the variability range of A. damkoehleri. K.O. Viets (1964 a) noted much variation (confirmed by our examination of the populations from different parts of South Africa) in many characters: position of P- 4 sword seta (halfway between ventral hairs, approaching to disto- or proximoventral hair, see Figs. 4 B,D), the shape of seta S- 1 (regularly more or less apically obtusely truncated, but sometimes this character is not clear when the seta is slightly bowed, often as a result of mounting), the fusion of Vgl- 3 (separate or fused with Cx-IV) and arrangement of Ac (from triangular to weakly curved). Furthermore, our examination of juvenile females of A. damkoehleri from the Mpumalanga province (material from Houtbosloop) shows much variation in the shape of Cx-IV, diameter of glandularia and fusion of Vgl- 1 and - 2 (Figs. 3 A-B) indicating that these characters are not suitable for species separation at a juvenile age. Atractides coriacellus was described by K. Viets (1956) from the Western Cape province (Great Berg River), without discussion of its similarity with other species. Due to glandularia enlarged by secondary sclerotization, P- 2 and - 3 ventral margin straight, a smooth excretory pore and fused Vgl- 1 and - 2, A. coriacellus resembles A. damkoehleri from which it differs in a longer I-L- 6 (I-L- 5 / 6 L ratio 1.26) and slender (L/HB 6.3, calculated from K. Viets 1956) and seta S- 1 apically slightly bowed and bluntly pointed. The other measurements given in the original description of Atractides coriacellus lie within the variability range of A. damkoehleri. Knowledge on the variability of additional specimens from the locus typicus is necessary before we can assess the taxonomic status of this species. Distribution. Cameroon: K. Viets (1916), Liberia: Cook (1966), South Africa: Mpumalanga province (Eastern Transvaal): K. O. Viets (1964 a), present study; Western Cape province: present study; East Africa: “ A. ugandensis ”, Lundblad (1952); “ A. duricorius ”, Walter & Bader (1952).Published as part of Pešić, Vladimir, Smit, Harry & Gerecke, Reinhard, 2011, New records of water mites of the genus Atractides Koch, 1837 (Acari: Hydrachnidia, Hygrobatidae) from South Africa, with descriptions of five new species, pp. 1-54 in Zootaxa 2986 on pages 2-8, DOI: 10.5281/zenodo.20065
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