1,965 research outputs found
(Verhoeffius) Kaplin 2019, subgen. nov.
No full textVerhoeffius Kaplin, subgen. nov. Type species: Trigoniophthalmus remyi Stach, 1939 (redescr. Wygodzinsky, 1958). Diagnosis. Urocoxites II–V with 2 + 2 exsertile vesicles. Description. Body length 6.5–15 mm. Hypodermal pigment absent or present on head and coxae. Paired ocelli pyriform. Cercus with one or two support apical spikes. Urocoxites II–V with 2 + 2 exsertile vesicles. Two described subgenera and 5 species (Table 1).Published as part of Kaplin, Vladimir, 2019, Taxonomic review of the genera Trigoniophthalmus Verhoeff and Coryphophthalmus Verhoeff (Archaeognatha, Machilidae) with descriptions of two new species of the genus Coryphophthalmus from Serbia, pp. 371-384 in Zootaxa 4661 (2) on pages 375-376, DOI: 10.11646/zootaxa.4661.2.6, http://zenodo.org/record/337952
Trigoniophthalmus kobani Kaplin 2019, sp. n.
No full text<i>Trigoniophthalmus kobani</i> Kaplin, sp. n. <p>(Figs 1–15)</p> <p> <b>Material.</b> Holotype, ♂ (in slides): Russia, North Ossetia <i>–</i> Alania, Prigorodny Distr., environs of Koban settl., 42 <i>°</i> 55ʹN <i>/</i> 44 <i>°</i> 30′E, 1100 m, 29.04.2018 (V. G. Kaplin). Paratype: 1♀ (in slides), same data as for holotype.</p> <p> <b>Description.</b> Body length: male 7 mm, female 7.5 mm. Body width: male and female 2.1–2.2 mm. General body color whitish. Antennal base, frons, clypeus, maxillae, mandibles, submentum, mentum, hypopharynx, thoracic sternites and coxae of all legs with purple hypodermal pigment of medium intensity. Color of scales on surface of body brownish. Antennae slightly shorter than body. Distal chains of flagellum divided into 7–9 articles in male and female. Clypeus of male with long thin bristles. Cercus approximately 0.37 (male) or 0.43 (female) body length, including about 25 articles. Apex of cercus with one spike (Fig. 1). Every second articles of cerci with 2 or 3 colorless supporting macrochaetae on the distal inner side.</p> <p>Compound eyes black (in ethanol). Ratio of length to width of compound eye about 1.02 in male and 1.05 in female; ratio of length of contact line to length of eye 0.52 in male and 0.48 in female. Paired ocelli submedian, pear-shaped, reddish-brown with very narrow white border, 1.8 times as wide as long in both sexes. Distance between inner margins of ocelli about 0.14 and between their outer margins 0.65–0.69 of total width of compound eyes in both sexes (Fig. 2).</p> <p> Apical article of maxillary palp 0.86–0.92 (male) or 1.01–1.02 (female) times as long as preceding one. Dorsal surface of 7 th, 6 th and 5 th articles of maxillary palp with 11 or 12, 12–14 and 2 or 3 hyaline spines, respectively, in both sexes. Ventral surface of 2–7 th articles of male maxillary palp with relatively numerous and long thin chaetae, on 6 th and 7 th articles such chaetae distributed more sparse (Figs 3, 4). Same long thin chaetae also present on dorsal surface of 2 nd articles of male labial palp (Fig. 5). Apical article of labial palp triangularly oval, 2.5 times as long as wide in both sexes. Mandibles with four distal teeth (Figs 6, 7).</p> <p> Fore femur and tibia of male and female widened. Fore femur of male without sensory field (Fig. 8). Ratios of length to width of femur, tibia and tarsus as shown in Table 1. Ratio of length of 3 rd tarsomere of tarsus to total length of tarsus about 0.35 in both sexes. Legs of male without long, thin bristles. Ventral surface of femora, tibiae and tarsi without spine-like setae. Middle and hind legs with coxal styli. Length of styli 0.5 mm (female) and 0.6 mm (male). Ratio of styli length to width of middle and hind coxae about 1.4–1.5 in female and 1.6–1.7 in male. Praetarsus with two claws and apically rounded cylindrical supporting projection between them. Ratio of length to width of projection about 1.6–1.7.</p> <p>Posterior margin of pronotum with a deep notch (Fig.9). In both sexes, uritesI andV–VII with 1+1 eversible vesicles, uritesII– IV with 2 + 2 such vesicles (Figs 10, 11). In male, posterior angle of urosternites II–VII and VIII approximately 76–78° and 84°, respectively; but in female, anterior angle of urosternites II–VII about 86–89°. Ratios of lengths of styli (without apical spine) and urocoxites II–IX as shown in Table 2. Inner posterior lobes of urocoxites VII between eversible vesicles of female slightly protruding (Fig. 11); ratio of length to total width of these lobes about 0.43. Thoracic tergites, urotergites I–IV, urosternites and urocoxites I–VI without macrochaetae in both sexes. Distribution of sublateral macrochaetae on urotergites V–X and spines on urocoxites VII–IX as shown in Table 3. Urocoxites IX with 2–3 and 5–6 (male) or 2 and 4 (female) outer and inner sublateral spines, respectively (Figs 12, 13).</p> <p>Ovipositor slender, elongate (2.3 mm), slightly surpassing apex of styli IX (Fig. 13). Anterior and posterior gonapophyses with approximately 40 and 41 divisions, respectively. One or two basal divisions of anterior gonapophyses and about 22 or 23 basal divisions of posterior gonapophyses glabrous.Apical macrochaetae of gonapophyses as long as three or four apical divisions combined. Distal divisions of anterior gonapophyses with 5–7, posterior gonapophyses with 3 or 4 setae (not counting sensory setae and apical macrochaetae) (Figs 14, 15). Ovarioles with 10 large eggs.</p> <p>Male genitalia with one pair of parameres on abdominal segment IX. Parameres with 1 + 6 divisions, slightly not attaining apex of penis (Fig 12). Penis and parameres not attaining of apex of urocoxites IX for 4.8 of width of aedeagus of penis. Phallobasis of penis 1.6 times as long as aedeagus.</p> <p> <b>Differential diagnosis.</b> Between species of the subgenera <i>Trigoniocellus</i> with 2 + 2 eversible vesicles on urites II–IV <i>T. kobani</i> <b>sp. n.</b> belongs to a group of small congeners with a body length of 7–9.5 mm, black or dark eyes, redish-brown paired ocelli, ratio of length to width of compound eye more than 1, the length of the contact line between compound eyes about 0.5–0.6 of eye length. This group includes 6 known species: <i>T. kobani</i> <b>sp. n.</b>, <i>T. tseyi</i> <b>sp. n.</b>, <i>T. minor</i> Kaplin, 2015, <i>T. borgustani</i> Kaplin, 2015, <i>T. subalpinus</i> Kaplin, 2017 and <i>T. abchasicus</i> Kaplin, 2017. <i>Trigoniophthalmus kobani</i> <b>sp. n.</b> differs from other species of this group by the color of paired ocelli, distribution of long thin chaetae on male labial palp, by the ratio of lengths of apical article and preceding one of maxillary palp, posterior angle of urosternites II–V, the number of outer and inner sublateral macrochaetae on urocoxites IX. The main differences between species of this group are summarized in Table 8.</p> <p> <b>Habitats.</b> Specimens of <i>Trigoniophthalmus kobani</i> <b>sp. n.</b> were collected in mountain forest (Quercus, Carpinus, shrubs) under stones, 1100 m above sea level.</p> <p> <b>Etymology.</b> The new species takes its name from the type locality: Koban, North Ossetia – Alania.</p>Published as part of <i>Kaplin, V. G., 2019, New species of bristletails of the genus Trigoniophthalmus Verhoeff, 1910 (Archaeognatha: Machilidae) from North Ossetia - Alania (Russia), pp. 25-34 in Caucasian Entomological Bulletin (Caucas. entomol. bull.) (Caucas. entomol. bull.) 15 (1)</i> on pages 25-28, DOI: 10.23885/181433262019151-2534, <a href="http://zenodo.org/record/8145596">http://zenodo.org/record/8145596</a>
Trigoniophthalmus Kaplin 2019, s. str.
No full textTrigoniophthalmus s. str. Diagnosis. With 2 + 2 exsertile vesicles on abdominal segments II–V. Spine-like setae on the ventral surface of tarsus absent. Description. Body length 8–12 mm. Hypodermal pigment on head and coxae. Compound eyes black, slightly widened. Paired ocelli black, submedian, close together, transverse, drop-shaped. Spine-like setae on the ventral surface of tarsus absent. Urocoxites II–V with 2 + 2 exsertile vesicles. Ovipositor shortened, not exceeding posterior angle of coxite IX, with 18–22 divisions. Anterior gonapophyses with long setae, divisions of posterior gonapophyses without setae (Figs 5–8). The subgenus includes one described species (Table 1).Published as part of Kaplin, Vladimir, 2019, Taxonomic review of the genera Trigoniophthalmus Verhoeff and Coryphophthalmus Verhoeff (Archaeognatha, Machilidae) with descriptions of two new species of the genus Coryphophthalmus from Serbia, pp. 371-384 in Zootaxa 4661 (2) on page 372, DOI: 10.11646/zootaxa.4661.2.6, http://zenodo.org/record/337952
Trigoniophthalmus tseyi Kaplin 2019, sp. n.
No full textTrigoniophthalmus tseyi Kaplin, sp. n. (Figs 16–28) Material. Holotype, ♂ (in slides): Russia, North Ossetia – Alania, Alagir Distr., Verkhniy Tsey settl.,environs of Recom sanctuary, 42 ° 47′31ʺN / 43 ° 54′14ʺE, 1950 m, 28.04.2018 (V. G. Kaplin). Paratypes: 1♂, 2♀ (1♀ in slides), same data as for holotype. Description. Body length: male 7–7.5 mm, female 7.3–8 mm. Body width: males and females 1.8–2.1 mm. General body color whitish. Antennal base, frons, maxillae, mandibles, hypopharynx with brown hypodermal pigment of weak and medium intensity. Color of scales on surface of body brownish. Antennae slightly shorter than body. Distal chains of flagellum divided into 9–12 articles in male and female (Fig. 16). Clypeus of male with long thin bristles. Cercus approximately 0.41–0.46 body length, including about 17 articles in both sexes. Apex of cercus with one spike (Fig.17). Articles of cerci, except apical one in female and two distal ones in male, with 1 or 2 colorless supporting macrochaetae on the inner side. Cerci of female also with 3 or 4 macrochaetae on outer side, male with only 1 macrochaeta in such position. Supporting macrochaetae also present on lateral sides of articles of caudal filament. Compound eyes black (in ethanol). Ratio of length to width of compound eye about 1.02–1.04 in males and 1.04–1.06 in females; ratio of length of contact line to length of eye 0.50–0.54 in both sexes. Paired ocelli submedian, pear-shaped, brown with a welldefined white border, 1.4–1.6 times as wide as long in both sexes. Distance between inner margins of ocelli about 0.16–0.18 and between their outer margins 0.60–0.65 total width of compound eyes in both sexes (Fig. 18). Apical article of maxillary palp 0.77 (male) or 0.82 (female) times as long as preceding one. Dorsal surface of 7 th, 6 th and 5 th articles of maxillary palp in both sexes with 11–13, 11–13 and 4 hyaline spines, respectively. Ventral surface of 2–7 th articles of male maxillary palp with relatively numerous and long thin chaetae, on sixth and especially seventh articles such chaetae distributed more sparse (Fig. 19). Similar long thin chaetae also present on dorsal surface of 2 nd and 3 rd articles of male labial palp. Apical article of labial palp triangularly oval, 2.2 (male) or 2.3 (female) times as long as wide (Fig. 20). Mandibles with 4 distal teeth (Fig. 21). Fore femur and tibia of male and female widened, without sensory field (Fig. 22). Ratios of length to width of femur, tibia and tarsus as shown in Table 4. Ratio of length of 3 rd tarsomere of tarsus to total length of tarsus 0.33–0.35 in both sexes. Coxa of male with long, thin bristles, missing from the femur, tibia and tarsus. Ventral surface of femur, tibia and tarsus with colorless spine-like chaetae (Table 5). Middle and hind legs with coxal styli. Length of styli 0.5 mm (female) or 0.6 mm (male). Ratio of length of styli to width of middle and hind coxae about 1.4–1.5 in both sexes. Praetarsus with 2 claws and support cone-shaped projection between them. Ratios of length to width of projection 1.9–2.1, widths of projection and pretarsus about 0.3. Posterior margin of pronotum with a deep notch. In both sexes, urites I and V–VII with 1 + 1 eversible vesicles, but urites II–IV with 2 + 2 eversible vesicles (Figs 24, 25). In male, posterior angle of urosternites II, III–VI, VII approximately 78°, 71–73° and 68°; but in female, 84°, 73–75° and 65°, respectively. In male, posterior angle of urosternite VIII about 84°. Ratios of lengths of styli (without apical spines) and urocoxites II–IX as shown in Table 6. Inner posterior lobes of urocoxites VII between eversible vesicles of female protruding (Fig. 25); ratio of length to total width of these lobes about 0.7. Thoracic tergites, urotergites I–IV, urosternites and urocoxites I–VI without macrochaetae in both sexes. Distribution of sublateral macrochaetae on urotergites V–X and spines on urocoxites VII–IX as shown in Table 7.Urocoxites IX, respectively, with 2 and 7–8 (male) or 1 and 7–11 (female) outer and inner sublateral spines (Figs 23, 26). Ovipositor slender, elongate (1.6–2.1 mm), slightly not surpassing apex of styli IX (Fig. 26). Anterior and posterior gonapophyses with approximately 47 and 50 divisions, respectively. Two basal divisions of anterior gonapophyses and about 27 basal divisions of posterior gonapophyses glabrous. Apical macrochaetae of gonapophyses as long as four apical divisions combined. Distal divisions of anterior gonapophyses with 4–6, posterior gonapophyses with 2–5 setae (not counting sensory setae and apical macrochaetae) (Figs 27, 28). Male genitalia with one pair of parameres on abdominal segment IX. Parameres with 1 + 7 divisions, penis slightly not attaining apex of parameres (Fig. 23). Penis and parameres not attaining of apex of coxites IX for 4.6 of width of aedeagus. Phallobasis of penis 1.3 times as long as aedeagus. Differential diagnosis. Trigoniophthalmus tseyi sp. n. belongs to the same species group as T. kobani sp. n. described above. Trigoniophthalmus tseyi sp. n. differs from other species of this group by ratio of distance between inner margins of paired ocelli to total width of eyes, presence of spine-like setae on legs, small posterior angle of urosternites II–V, large number of inner sublateral spines on urocoxite IX. Main differences between these species of this group are in Table 8. Habitats. Specimens of Trigoniophthalmus tseyi sp. n. were collected in mountain forest (Pinus, Fagus, shrubs) under stones, 1950 m above sea level. Etymology. The new species takes its name from the type locality: Verkhniy Tsey, North Ossetia – Alania.Published as part of Kaplin, V. G., 2019, New species of bristletails of the genus Trigoniophthalmus Verhoeff, 1910 (Archaeognatha: Machilidae) from North Ossetia - Alania (Russia), pp. 25-34 in Caucasian Entomological Bulletin (Caucas. entomol. bull.) (Caucas. entomol. bull.) 15 (1) on pages 28-31, DOI: 10.23885/181433262019151-2534, http://zenodo.org/record/814559
Persiatelurina daghestaniana Kaplin & Vasin 2019, sp.nov.
No full textPersiatelurina daghestaniana Kaplin, sp.nov. (Figs 1–17) Holotype. Female (in slides), Russia, Daghestan, environs of Derbent fortress, 42°03'10''N, 48°16'26''E, 22.IV.2018, V. Kaplin, V. Vasin (VIZR). Paratypes. Two females (in slides), same data as for holotype (VIZR). Description. Body length 3.2–3.8 mm; head length 0.26–0.28 mm; thorax length 1.1–1.2 mm. Head width 0.8–0.9 mm; thorax width 1.3– 1.4 mm; width of abdominal segment IX about 0.5 mm. General colouration yellowish white with golden chaetae and scales. Body small, elongate (2.2– 2.6 times as long as wide), ateluriform in shape, covered (including head and coxae) by relatively large scales which 1–2.1 times as long as wide and with 14–20 rays (antennae and cerci partially broken). Macrochaetae simple or with apical bifurcations. Head small, 1.5 times as wide as long (Fig. 1). Basal annuli of flagellum with trichobothria. Mandibles with row of macrochaetae along outer surface (last of these macrochaetae with apical bifurcation) as well as with well-developed incisor and molar regions. Incisor region with seven incisives. Maxillary palps with five palpomeres. Ultimate palpomere of maxillary palps 1.2 times as long as penultimate one; ultimate palpomere with four apical sensorial papillae of usual form (Fig.3). Galea of maxilla with two apical conules: one more rounded than other. Apical tooth of lacinia bifid; pectinate prostheca long with about 20–22 narrow hyaline projections, five bifurcated hyaline lamellae (first of these lamellae large and perpendicular, rest lamellae inclined), about five simple macrochaetae and three small chaetae along inner margin. Perpendicular lamella in apical part with five lateral projections (Fig. 4). Apical palpomere of labial palp ovoid, about 1.3–1.5 times as long as wide, with six typical sensorial papillae (Fig. 2). Undersurface of last three palpomeres of labial palps with numerous relatively shortened and thickened, slightly curved and apically bifurcated chaetae. Glossae divided into two pairs of lobes, shorter than paraglossae. Paraglossae with simple and bifid chaetae. Lateral margins of thoracic tergites with a row of setae, two of which (located in posterolateral angles of tergites) apically bifid and longest (Fig. 9). Anterior border of pronotum with numerous small chaetae. Posterior margin of pro-, meso- and metanotum with eight to ten small chaetae (Fig. 9). Legs quite elongate. Coxae and femora widened (Fig. 5). Ratio of lengths: for I, II and III coxae about 2.0, 1.6 and 1.4, respectively; for femora, 1.7, 1.6 and 1.5; for tibiae, 3.7, 4.2 and 4.5. All tibiae with two lyriform spines located at apex of dorsal part, but all femora with only one such spine (Fig. 6). Tibia also with one relatively large apical spur and four ventral simple macrochaetae. Middle part of all femora with two long macrochaetae on poorly marked ledge: distal one bifid, proximal one simple. Praetarsus with two strong claws, two small pulvilli, medial empodial claw and slightly convex small support oval platform with 3–4 transverse ribs (Fig. 7). Urotergites I–VIII infralaterally with 2+2 bifid and 2+2 additional simple macrochaetae: inner chaetae more robust and longer than other ones. Posterior margin of urotergites I–VIII with four small chaetae (Fig. 10). Urotergite IX with posterolateral corners protruding, with 1+1 apical, 5+5 outer and 1+1 inner macrochaetae; outer and inner macrochaetae shorter than apical ones (Fig. 11). Urotergite X with semicircular concave hind margin; ratio of width to depth of this notch (concavity) about 2.2; apex of posterolateral angles of urotergite X with 1+1 long and strong macrochaetae; inner and outer margins of this notch with three macrochaetae each (Fig. 8). Urosternite I with 1+1 submedian small chaetae. Urosternite II with one pair of submedian vesicles and with six setae (Fig. 12). Urosternite III with four macrochaetae and two small chaetae (Fig. 13). Urosternites IV–VI with five or six chaetae on hind margin. Two submedian macrochaetae of urosternites III–VI long, apically bifurcated (Fig. 13). Urosternite VII with one pair of pseudovesicles, 1+1 submedian long simple macrochaetae between pseudovesicles and 2+2 sublateral smaller setae (Fig. 14). Subgenital plate well developed, 1.8 times as wide as long, with rounded hind margin and single row of simple setae, apically slightly acute (Fig. 14). Urosternites VII–IX with styli; ratios of length of styli (without apical spines) on urosternite VII, urocoxite VIII and urocoxite IX to length of corresponding urosternite and urocoxites about 0.74, 0.78 and 0.84, respectively; ratios of lengths of apical spines and styli (without apical spines) for urosternite VII as well as for urocoxites VIII and IX about 0.30, 0.25 and 0.18, respectively (Figs 14, 15). Ovipositor spindle-shaped, slightly shorter than styli IX. Gonapophyses VIII and IX with nine divisions and acute apices; gonapophyses VIII with numerous straight chaetae, but its outer margin also with one longer chaeta on each of 2–9th or 2–8th divisions; inner margin of gonapophysis IX with one such chaeta on each of 5–9th divisions (Figs 16, 17); gonapophyses IX with an inner apical spiny area; apical part of gonapophyses IX also with well-developed oval sensorial field. Basal part of cerci with trichobothria. Male unknown. Remarks. The new species belongs to the genus Persiatelurina, which includes two previosly described species: P. caucasica (Kaplin, 2016) from Abkhazia and P. farsiana Molero et al., 2018 from Southwestern Iran. The new species differs from these species by structure of the legs, maxillae, maxillary and labial palps, pronotum, urotergite X, urosternites II, VII, urocoxites VIII and IX, subgenital plate, ovipositor. The main differences between species are shown in Table. Etymology. The name of the new species derives from the Daghestan Republic combined with the suffix “-iana”. Habitats. All specimens of Persiatelurina daghestaniana sp. nov. were collected in mountain forest (Quércus, Fráxinus, Rosa, other shrubs), in soil under stones, with small ants.Published as part of Kaplin, V. G. & Vasin, V. G., 2019, A new species of silverfish of the genus Persiatelurina Molero et al. (Zygentoma: Nicoletiidae) from Daghestan, pp. 88-93 in Zoosystematica Rossica (Zoosyst. Rossica) (Zoosyst. Rossica) 28 (1) on pages 89-93, DOI: 10.31610/zsr/2019.28.1.8
Petrobiellus sachalinensis Kaplin 2020, sp. nov.
No full text<i>Petrobiellus sachalinensis</i> sp. nov. <p>(Figs 1–13)</p> <p> <i>Holotype.</i> Female (slide-mounted), <b>Russia</b>, <i>Sakhalin Prov</i>., western shore of northern Sakhalin I., near Aleksandrovsk-Sakhalinsky, 50°54′N 142°09′E, shoreline, under stones, 8.VIII.2019, V. Kaplin leg.</p> <p> <i>Paratypes.</i> 11 females (1 female on slide), same data as for holotype.</p> <p> <i>Description.</i> Female. Body length (not including antennae): 8.4–11.6 mm; body width: 2.9–3.1 mm; cercal length: 5.5–5.7 mm; total width of compound eyes: 0.94–1.00 mm; eye length: 0.54– 0.57 mm; paired ocelli width: 0.49–0.51 mm, length: 0.17–0.18 mm. Middle and hind coxal styli length: 0.60–0.65 mm. Ovipositor length: 3.35– 3.50 mm. General body colour whitish or slightly yellowish, almost without pigment. Antennal base, frons, lateral parts of clypeus, basal part of first and third articles of maxillary palps, mandibulae, hypopharynx, and first, third and basal half of second tarsomeres with brown or brown-violet pigment of weak or medium intensity. Maxillary and labial palps, flagellum and pedicellus of antennae, legs, coxal and abdominal styli without scales. Scapus with very sparse scales. Colour of scales whitish or light gray. Metanotum also with one pair, urotergites with three pairs of large spots of brown scales (Fig. 1). Antennae very long, 1.5– 1.9 times as long as body. Distal chains of flagellum divided into 14–23 annuli (Fig. 2). Clypeus without long thin bristles. Cercus approximately 0.48– 0.55 times as long as body, with about 21–24 divisions. Apex of cercus with one apical spike (Fig. 3). All divisions of cerci, except for apical one or two, with 1–3 colorless spines on inner side supporting cerci above substrate. Divisions in middle part of cerci also with one outer lateral spine (Fig. 4).</p> <p>Compound eyes dark, with bluish tinge (in ethanol). Ratio of length to width of compound eye about 1.13–1.15; ratio of contact line to length of eyes about 0.45. Paired ocelli sole-shaped, brown, with narrow white bordering, 2.8–3.0 times as wide as long. Distance between inner margins of paired ocelli 0.13–0.15 and between their outer margins 0.96–0.98 times total width of compound eyes (Fig. 5). Frons clearly convex between paired ocelli.</p> <p> <i>Note</i>. Length of the stylus does not include apical spines.</p> <p> <i>Note</i>. <i>Petrobiellus takunagae</i> Silvestri, 1943 is not included in the table due to insufficient data in its original description; the characters distinguishing this species from <i>P. sachalinensis</i> <b>sp. nov.</b> are given in the <i>Comparison</i> section after the description of the new species.</p> <p>.</p> <p>Apical article of maxillary palp 0.60–0.62 times as long as preceding one. Fifth article 1.04– 1.06 times as long as 4th article (Fig. 6). Dorsal surface of 7th, 6th and 5th articles of maxillary palp with 13–16, 10–13 and 3 hyaline spines, respectively. Apical article of labial palp fingerlike, 3.3–3.4 times as long as wide (Fig. 7). Mandibles with 2–3 distal teeth (Fig 8). Femora slightly widened. Ratios of length to width of femur, tibia and tarsus as shown in Table 1. Fore and middle legs shorter than hind legs. Fore and middle tarsus and tibia 0.77 and 0.83 times as long as those of hind legs, respectively. Ratio of length of apical hind tarsomere to total length of hind tarsus about 0.32. Ventral surface of legs without long thin bristles. Fore, middle and hind tarsi and hind tibia with slightly pigmented spines (Fig. 9). Number of spines as shown in Table 2.</p> <p>Middle and hind legs with coxal styli. Ratio of length of styli to width of middle and hind coxae 1.03–1.06 and 1.09, respectively.</p> <p>Urites I, VI and VII with 1 + 1, urites II–V with 2 + 2 eversible vesicles (Fig. 10). Posterior angle of urosternites II, III–V, VI, and VII approximately 118°, 110–112°, 107°, and 120°, respectively. Inner posterior lobes of urocoxites VII protruding. Ratio width to length of protruding lobe about 1.0. Ratios of lengths of stylus (without apical spine), urosternites and urocoxites II–IX as shown in Table 3. Ratio of lengths of urosternite and urocoxites I about 0.35. Urocoxites IX with 1 + 1 outer sublateral spines.</p> <p>Ovipositor slender, elongate, significantly protruding beyond apices of styli IX. Anterior and posterior gonapophyses with 52 or 53 divisions (Fig. 11). Four basal divisions of anterior gonapophyses and about 11 basal divisions of posterior gonapophyses glabrous. Apical spines of gonapophyses as long as three apical divisions combined. Distal divisions of anterior and posterior gonapophyses with seven or eight setae (not counting sensory setae and apical spines) (Figs 12, 13).</p> <p>Males are unknown; probably the species is parthenogenetic.</p> <p> <i>Comparison. Petrobiellus sachalinensis</i> <b>sp.nov.</b> can be distinguished from its congeners by the following characters: the general body colour whitish or slightly yellowish (<i>vs</i> yellowish or rufous); colour of scales whitish or light gray (<i>vs</i> brownish or dark brownish); body (excluding head and tarsi) almost without pigment (<i>vs</i> significantly pigmented); and paired ocelli almost sole-shaped (<i>vs</i> pistiliform, strongly narrowed in middle parts). The new species also differs from all other species of the genus in the most elongated compound eyes; relatively short coxal styli; long 4th article of maxillary palp; small number of short supporting spines in ventral part of tarsi; ratios of lengths of urosternite and urocoxite, stylus and urocoxite; posterior angle of urosternite; and number of divisions of ovipositor. The main morphological differences of females of <i>P. sachalinensis</i> <b>sp. nov.</b> from those of the other species of <i>Petrobiellus</i> are given in Table 4. According to the brief description of <i>P. takunagae</i> by Silvestri (1943), the distance between the paired ocelli in this species is about half the length of ocellus, and in <i>P. sachalinensis</i> <b>sp. nov.</b>, this distance is almost equal to the length of ocellus. The new species differs from <i>P. takunagae</i> also in femora without pigment and tibiae poorly pigmented (<i>vs</i> femora and tibiae with intensive blackish-violet pigment); and four basal divisions of anterior gonapophyses glabrous (<i>vs</i> about 12 basal divisions of anterior gonapophyses glabrous).</p> <p> <i>Etymology</i>. The species name is derived from the toponym of Sakhalin.</p> <p> <i>Habitats.</i> Western shores of the northern part of Sakhalin Island. Under stones near the line of seashore (Fig. 14; see Addenda section).</p>Published as part of <i>Kaplin, V. G., 2020, A new species of bristletails of the genus Petrobiellus (Microcoryphia: Machilidae) from Sakhalin, pp. 17-22 in Zoosystematica Rossica (Zoosyst. Rossica) (Zoosyst. Rossica) 29 (1)</i> on pages 18-22, DOI: 10.31610/zsr/2020.29.1.17, <a href="http://zenodo.org/record/10124220">http://zenodo.org/record/10124220</a>
Turquimachilis taurica Kaplin 2021, sp. nov.
No full text<i>Turquimachilis taurica</i> sp. nov. <p>(Figs 1–14)</p> <p> <i>Holotype</i>. Female (slide-mounted); <i>Republic of Crimea</i>, Grand Canyon of Crimea, 44°31 <b>′</b> 40 <b>″</b> N, 34°01 <b>′</b> 00 <b>″</b> E, 500–600 m, under stones, 7.IX.2020, V. Kaplin leg.</p> <p> <i>Description.</i> Body length: 9.6 mm; body width: 2.0 mm; length of antenna: about 7.5 mm; length of cerci: 3.4 mm; total width of eyes: 1.00 mm; length of eye: 0.44 mm; width of paired ocellus: 0.36 mm; length of paired ocellus: about 0.13 mm; length of coxal styli of legs: about 0.4 mm; length of ovipositor: 1.45 mm.</p> <p>General body colour (in ethanol) light brownish, with dark brown scales on upper and lower sides of body. Frons, clypeus, occiput, mandibles, maxillae, maxillary and labial palps, and legs with reddish brown or reddish violet pigment. Head, maxillary palps and coxae most pigmented. Scapus, pedicellus and flagellum of antennae without pigment.</p> <p>Antennae shorter than body. Distal chains of flagellum divided into 9–11 annuli (Fig. 1). Cercus approximately 0.36 times the body length. Apex of cercus (Fig. 2) with two lateral spikes. Articles of cerci, except for apical two, with colourless supporting macrochaetae on inner side.</p> <p>Compound eyes dark brown (in ethanol). Ratio of compound eye length to its width 0.89; ratio of contact line length to eye length 0.61 (Fig. 3). Paired ocelli black with narrow white rim, shoeshaped, subinferior to compound eyes. Frons slightly swollen between paired ocelli. Distance between inner margins of ocelli about 0.21 times and between their outer margins 0.92 times the total width of compound eyes.</p> <p>Maxillary palp (Fig. 4): apical palpomere 0.84 times as long as preceding one; ratio of lengths of 5th and 4th palpomeres about 1.64; dorsal surface of 7th, 6th and 5th palpomeres with 17–18, 16–18 and 5–6 hyaline spines, respectively. Labial palp (Fig. 5): apical palpomere triangular oval, 2.1–2.2 times as long as wide, with about 21–25 sensorial cones. Mandible with four well-developed distal teeth (Fig. 6).</p> <p>Fore and middle femur and tibia widened (Fig. 7). Fore and hind legs 1.09 and 1.16 times as long as middle legs, respectively. Ratios of length to width of femur, tibia and tarsus as in Table 1. Ratio of length of 3rd tarsomere of hind tarsus to its total length about 0.39. Ventral surfaces of coxae, femora, tibiae and tarsi without hyaline spine-like chaetae. Ratio of coxal stylus length to width of middle and hind coxa about 1.7 (Fig. 8).</p> <p>Urocoxites I and VI–VII with 1 + 1 eversible vesicles; urocoxites II–V with 2 + 2 eversible vesicles. Posterior angle of urosternites II–VI approximately 82–86°, that of urosternite VII about 74°. Ratios of lengths of urosternites and urocoxites II–VI 0.67–0.69. Ratios of lengths of urostyli (without apical spine) and urocoxites II–VII 0.50–0.54. Ratio of length of urostylus and urocoxite VIII 0.68, IX 0.45. Ratios of lengths of apical spines and urostyli II–VIII 0.55–0.62 (Figs 9–11). Urocoxite VII (Fig. 10) with well-developed lobes protruding between styli; ratio of length to width of one lobe about 0.75. Thoracic tergites, urosternites, urocoxites I–VIII, and urotergites I–VI and X without macrochaetae. Urocoxites IX (Fig. 12) with 3–5 inner sublateral spines (spiniform chaetae), without outer sublateral spines. Urotergites VII with 1 + 1, urotergites VIII and IX with 2 + 2 sublateral spines.</p> <p> Ovipositor weakly sclerotised, thickened, covered by urocoxites IX, stout, similar to that of <i>T. mendesi</i>.</p> <p>Gonapophysis VIII (Fig. 13) with 17 divisions; its apex with two small spine-like lobules and a subterminal chaeta between them; chaeta being a little longer than apical division; apical division with about 15 sensory spines and 2–3 relatively long thin chaetae in preapical part and with two curved chaetae in basal part. Remaining divisions of gonapophysis VIII with a row of setae, some of which, mainly those on inner and outer sides of gonapophysis, long and ciliary. Basal division without chaetae. Gonapophysis VIII without digging teeth.</p> <p>Gonapophysis IX (Fig. 14) with 19 divisions, a somewhat curved horn at apex and a subterminal chaeta being slightly longer than apical division. Apical division also with a group of 11–12 tiny sensory spines; these spines present also on preceding 5–6 divisions, but number of spines decreasing towards base. Each of these divisions also with about 2–4 chaetae; chaetae on outer side longer. Basal division without chaetae.</p> <p> <i>Comparison.</i> According to the structure of the female gonapophyses and 2 + 2 eversible vesicles on urocoxites II–V, the new species belongs to the genus <i>Turquimachilis</i> (Bach de Roca et al., 2013), which was monotypic, with the only species <i>T. mendesi</i>. The main morphological differences between <i>T. taurica</i> <b>sp.nov.</b> and <i>T. mendesi</i> are given in Table 2. The new species differs from <i>T. mendesi</i> in the structure of the compound eyes, maxillary palp, urocoxites, and the number of divisions of gonapophyses. The ratio of lengths of contact line and compound eye in female of <i>T. taurica</i> <b>sp. nov.</b> is about 0.6, while in female of <i>T. mendesi</i> it is about 0.5. The numbers of hyaline spines on the dorsal surface of the 6th and 7th palpomeres of maxillary palps are 16–18 and 9–12, respectively. The ratios of length to width of hind tibia are about 2.8 and about 2.6, respectively; the ratios of lengths of urostyli and urocoxites II–V are 0.50–0.52 and 0.38–0.43; the number of divisions of gonapophyses are 17–19 and 15–16, respectively. The urocoxites of <i>T. mendesi</i> have no spiniform chaetae, whereas urocoxites IX of <i>T. taurica</i> <b>sp. nov.</b> bear 3–5 inner sublateral spines.</p> <p> <i>Etymology</i>. The species name is a Latin adjective referring to Chersonesus Taurica, an ancient name of the Crimea, where the holotype was collected.</p> <p> <i>Habitat</i>. The holotype of <i>T. taurica</i> <b>sp. nov.</b> was collected in a forest (<i>Carpinus</i>, <i>Fagus</i>, <i>Fraxinus excelsior</i>, <i>Quercus robur</i>) among the stones.</p>Published as part of <i>Kaplin, V. G., 2021, A new species of bristletails of the genus Turquimachilis (Microcoryphia: Machilidae) from the Crimea, pp. 40-45 in Zoosystematica Rossica (Zoosyst. Rossica) (Zoosyst. Rossica) 30 (1)</i> on pages 41-45, DOI: 10.31610/zsr/2021.30.1.40, <a href="http://zenodo.org/record/10124241">http://zenodo.org/record/10124241</a>
Figs 18–24 in NEW SPECIES OF BRISTRLETAILS OF THE GENUS CORYPHOPHTHALMUS VERHOEFF, 1910 (ARCHAEOGNATHA: MACHILIDAE) FROM NORTH AND SOUTH OSSETIA
No full textFigs 18–24. Coryphophthalmus tskhinvalicus Kaplin, sp. n., holotype (18–20, 23, 24) ♂,Published as part of Kaplin, V. G., Kiseleva, L. V. & Kozhevnikova, O. P., 2020, NEW SPECIES OF BRISTRLETAILS OF THE GENUS CORYPHOPHTHALMUS VERHOEFF, 1910 (ARCHAEOGNATHA: MACHILIDAE) FROM NORTH AND SOUTH OSSETIA, pp. 1-13 in Far Eastern Entomologist 406 on page 8, DOI: 10.25221/fee.406.1, http://zenodo.org/record/716547
Figs 10–17 in NEW SPECIES OF BRISTRLETAILS OF THE GENUS CORYPHOPHTHALMUS VERHOEFF, 1910 (ARCHAEOGNATHA: MACHILIDAE) FROM NORTH AND SOUTH OSSETIA
No full textFigs 10–17. Coryphophthalmus morulus Kaplin, sp. n., holotype ♂ (10, 15–17), paratypePublished as part of Kaplin, V. G., Kiseleva, L. V. & Kozhevnikova, O. P., 2020, NEW SPECIES OF BRISTRLETAILS OF THE GENUS CORYPHOPHTHALMUS VERHOEFF, 1910 (ARCHAEOGNATHA: MACHILIDAE) FROM NORTH AND SOUTH OSSETIA, pp. 1-13 in Far Eastern Entomologist 406 on page 5, DOI: 10.25221/fee.406.1, http://zenodo.org/record/716547
Charimachilis WYGODZINSKY 1939
No full textGenus Charimachilis Wygodzinsky, 1939 Type species: Praemachilis orientalis Silvestri, 1908.Published as part of Kaplin, V. G. & Martynov, V. V., 2022, A new species of the bristletail genus Charimachilis Wygodzinsky, 1939 (Microcorypha: Machilidae) from Eastern Ukraine, pp. 1-8 in Far Eastern Entomologist 449 on page 2, DOI: 10.25221/fee.449.1, http://zenodo.org/record/716684
- …
