102,237 research outputs found

    Stenopelmatus mineraldelmonte Weissman & Vandergast & Song & Shin & Mckenna & Ueshima 2021, n. sp.

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    Stenopelmatus mineraldelmonte Weissman, n. sp. Mineral del Monte Jerusalem Cricket Figs 81–83, Tables 1, 2Published as part of Weissman, David B., Vandergast, Amy G., Song, Hojun, Shin, Seunggwan, Mckenna, Duane D. & Ueshima, Norihiro, 2021, Generic relationships of New World Jerusalem crickets (Orthoptera: Stenopelmatoidea: Stenopelmatinae), including all known species of Stenopelmatus, pp. 1-122 in Zootaxa 4917 (1) on page 55, DOI: 10.11646/zootaxa.4917.1.1, http://zenodo.org/record/447203

    Stenopelmatus hondurasito Weissman & Vandergast & Song & Shin & Mckenna & Ueshima 2021, n. sp.

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    Stenopelmatus hondurasito Weissman, n. sp. Honduras Small Black Jerusalem Cricket Figs 66–70, Tables 1, 2Published as part of Weissman, David B., Vandergast, Amy G., Song, Hojun, Shin, Seunggwan, Mckenna, Duane D. & Ueshima, Norihiro, 2021, Generic relationships of New World Jerusalem crickets (Orthoptera: Stenopelmatoidea: Stenopelmatinae), including all known species of Stenopelmatus, pp. 1-122 in Zootaxa 4917 (1) on page 49, DOI: 10.11646/zootaxa.4917.1.1, http://zenodo.org/record/447203

    Stenopelmatus sanfelipe Weissman & Vandergast & Song & Shin & Mckenna & Ueshima 2021, n. sp.

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    Stenopelmatus sanfelipe Weissman, n. sp. San Felipe Jerusalem Cricket Figs 132–142, Tables 1, 2Published as part of Weissman, David B., Vandergast, Amy G., Song, Hojun, Shin, Seunggwan, Mckenna, Duane D. & Ueshima, Norihiro, 2021, Generic relationships of New World Jerusalem crickets (Orthoptera: Stenopelmatoidea: Stenopelmatinae), including all known species of Stenopelmatus, pp. 1-122 in Zootaxa 4917 (1) on page 81, DOI: 10.11646/zootaxa.4917.1.1, http://zenodo.org/record/447203

    Stenopelmatus ecuadorensis Weissman & Vandergast & Song & Shin & Mckenna & Ueshima 2021, n. sp.

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    Stenopelmatus ecuadorensis Weissman, n. sp. Ecuador Jerusalem Cricket Figs 41–44, Table 1, 2Published as part of Weissman, David B., Vandergast, Amy G., Song, Hojun, Shin, Seunggwan, Mckenna, Duane D. & Ueshima, Norihiro, 2021, Generic relationships of New World Jerusalem crickets (Orthoptera: Stenopelmatoidea: Stenopelmatinae), including all known species of Stenopelmatus, pp. 1-122 in Zootaxa 4917 (1) on page 37, DOI: 10.11646/zootaxa.4917.1.1, http://zenodo.org/record/447203

    Stenopelmatus saltillo Weissman & Vandergast & Song & Shin & Mckenna & Ueshima 2021, n. sp.

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    Stenopelmatus saltillo Weissman, n. sp. Saltillo Jerusalem Cricket Figs 126–131, Table 2Published as part of Weissman, David B., Vandergast, Amy G., Song, Hojun, Shin, Seunggwan, Mckenna, Duane D. & Ueshima, Norihiro, 2021, Generic relationships of New World Jerusalem crickets (Orthoptera: Stenopelmatoidea: Stenopelmatinae), including all known species of Stenopelmatus, pp. 1-122 in Zootaxa 4917 (1) on page 78, DOI: 10.11646/zootaxa.4917.1.1, http://zenodo.org/record/447203

    Gwynnagraecia Rentz, Su and Ueshima 2021, Gen. et sp. nov.

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    Gwynnagraecia Rentz, Su and Ueshima Gen. et sp. nov. Type species: Gwynnagraecia viridis Rentz, Su & Ueshima gen. et sp. nov., here designated. 11 Named after our colleague Prof Darryl T Gwynne to celebrate his retirement. Common name. Gwynne’s Katydids Gwynnagraecia gen. nov. is proposed to accommodate three small micropterous species mostly from coastal Western Australia. The genus is unique in several characters. It is small as are many other Western Australian katydids in the Agraeciini as well as in the Tettigoniinae (Oligodectoides. Rentz, Nanodectes Rentz, see Rentz (1985)) and the Microtettigoniinae (Microtettigonia, see Rentz (2001)). Males have a peculiar extension of the tenth abdominal tergite (acting as pseudocerci?) and the titillators are largely unsclerotised. Females have a long, straight ovipositor and are wingless. The genus bears a superficial resemblance to Australiagraecia gen. nov., p. 8 but the cerci are very different from any of the species in that genus and no Australiagraecia gen. nov. species has modification of the tenth abdominal tergite. In addition to the odd extensions of the tenth tergite mentioned above, males have an exaggerated, ice-tong-like cercus which apparently fits into a modified pocket (Fig. 11J) during copulation (exception G. marandoo Gen. sp. nov., see p. 22). Vahed et al. (2014) commented on cerci of similar structure in other tettigoniids. They observed copulations in at least 44 species of Tettigoniidae. During mating males transfer a spermatophore consisting of two parts, the ejaculate which contains the ampulla containing the sperm) and the spermatophylax, a white sticky substance containing highly nutritious ingredients that are said to be essential in the successful development of eggs. Females feed on the spermatophylax after separating from mating. Vahed et al. concluded that tong-like cerci are associated with transfer of ejaculate during copulation. It appears that katydids with reduced spermatophylaxes bear extraordinarily adapted clasping cerci that, as a result, seem to prolong copulation and prevent the female from feeding on the spermatophylax. Unfortunately, we have not observed any mating in Gwynnagraecia gen. nov. or Australiagraecia gen. nov. that could test these hypotheses. These katydids are common in their environment that similar experiments could be conducted to determine if the hypothesis is true for members of the Agraeciini. It would be revealing to test this hypothesis with G. marandoo gen. et sp. nov. the only known species in the genus that does not have an exaggerated ice-tong-like male cercus. Gwynnagraecia gen. nov. superficially resembles Australiagraecia, gen. nov., p. 8, a taxon with many species in the southern portion of Australia, see p. 67. The species we describe here can be identified using the descriptions and comparing the morphological characters figured. Generic characters. Size small for tribe, form robust. Male macropterous, females apterous. Head not especially slanting (Fig. 10D), jaws not abnormally large; frons smooth, shining, not pitted or carinate (Fig. 10C), genae similar to frons; median ocellus minute, positioned on triangular frontal fastigium, lateral ocelli absent. Eye ovoid, very large, situated high on head (Figs 10B, C); frontal fastigium absent; fastigium of vertex short, stout, with minute median sulcus not projecting above antennal scape; scape large, produced quadrate on dorsal internal margin, pedicel very small, not modified; flagellum thread-like, slightly longer than length of body. Pronotum short, slightly raised, surface smooth, without carinae or pits; lateral lobe shallow, ventral margin feebly undulate, not swollen over thoracic auditory foramen; anterior margin of disk feebly concave, posterior margin truncate. Male tegmen extending for most of its length beyond apex of pronotum. Stridulatory file (Fig. 20E, F); tegmen without costa, subcosta arching, widely separated from media. Prothorax armed with a pair of elongate spines; meso- and metathorax each with prominent tube-like projections. All legs normal. All femora clothed with dark setae dorsally, ventral surface unarmed on both margins; fore tibia short, slightly swollen apically, dorsal surface unarmed, ventral margin with 6 spines on both margins, the distal two much smaller than the others; middle tibia cylindrical. but slightly swollen, dorsal surface armed on posterior margin with 2 spines positioned at the extremities of the middle third, ventral surface armed similarly to fore tibia (Figs.10L). Hind tibia armed apically with a pair of dorsal and ventral spurs (Figs 10J, K). Genicular lobes unarmed. Abdomen normal, without any prominent ridges or modifications except for genitalia (Figs 10E, N). Male tenth tergite extraordinary, an elongate curved tong projecting from each side, apex specifically modified; median incision absent, margin continuous; cercus massive, shaft straight, with apical ice-tong-like hook; titillators without any sclerotised portions; subgenital plate elongate, with median carina, styles elongate, median incision shallow. Female tenth tergite with narrow median incision; cercus fairly robust; subgenital plate triangular, apex acute. Ovipositor very elongate, much longer than length of body, gracefully upcurved, unarmed. Overall colour in life bright green, shining; abdomen with a pair of faint, thin, parallel stripes over most of the dorsal surface.Published as part of Rentz, Dcf, Su, You Ning & Ueshima, Norihiro, 2021, Studies in Australian Tettigoniidae: New short-winged Agraeciini from Australia (Orthoptera: Tettigoniidae; Conocephalinae; Agraeciini), pp. 1-72 in Zootaxa 5059 (2) on pages 19-20, DOI: 10.11646/zootaxa.5059.1.1, http://zenodo.org/record/560250

    Hungerfordia ngereamensis M. Yamazaki and Ueshima, sp. nov.

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    Hungerfordia ngereamensis M. Yamazaki and Ueshima sp. nov. (Figs. 2, 12 O) Material examined. Holotype. Chongelungel, south of Ngeream (Loc. BA 4 - 2), Airai state, Babeldaob island, Palau. Collected by R. U. and M. Y. on July 7, 2006. UMUTZ-MG-B0293T. Paratypes. Babeldaob: Loc. BA 3 -1, 10 specimens, 27 /VI/ 2005, UMUTZ-MG-B0172; Loc. BA 4 -2, 26 specimens, 7 /VII/ 2006, UMUTZ-MG-B0293. Diagnosis. Shell sinistral, large to medium-size for Hungerfordia species, slender conical. Shell color white, upper whorls frequently yellow or yellowish brown. Apical septum absent. Axial rib on most whorls highly protruded as spines, very widely spaced, strongly folded at the peripheries of the whorls; spiny projections of the ribs moderately high, rather narrow, arising at some distance below the suture; ribs above the aperture abruptly becoming closely spaced, with greatly reduced spiny projections, less folded. Interspace between the spiny ribs with some very weak and very short growth wrinkles below the suture on the lower whorls. Spiral sculptures present throughout teleoconch. Aperture weakly protruded. Columellar tooth very weakly emerged within the aperture in front view, but strongly developed deeply inward. Peristome triple, with doubled inner peristomes, stout. Outer peristome strongly developed, widely expanded. Inner peristome double, also expanded; distal inner peristome strongly developed just behind the proximal inner peristome, gradually narrowed toward the aperture; proximal inner peristome always narrower than the distal one in front view. Interspace between the outer and inner peristomes wide; outer surface with a series of fine and very low riblets at the abapertural half (near the outer peristome), almost smooth on the adapertural half (just behind the inner peristomes). Description. Shell (Figs. 2 A–F, 12 O). Shell sinistral, large to medium-size for Hungerfordia species, conical or fusiform, penultimate whorl widest (slightly wider than the last whorl) or last 2 whorls almost equally widest when removing the expanded ribs and peristomes; sides almost straight. Shell color white, upper whorls occasionally yellow. Whorls 6–7, slightly convex; last 1 / 8 whorls (including the interspace between the outer and inner peristomes) expanded upward and ascending onto the penultimate whorl in left lateral view. Suture weakly impressed. Constriction located above the parietal/columellar junction of the inner peristome, associated with abrupt changes of the ribbing pattern, internally without palatal plica except a low axial-crest caused by the constriction. Tuba 7 / 8 whorls. Protoconch 1 + 1 / 8 – 1 + 1 / 4 whorls, almost smooth, very weakly and finely punctated. Apical septum absent. Axial ribs strongly developed on the teleoconch, oblique to the coiling axis, highly protruded as spines on most whorls: ribs on the first 1 / 3 – 1 / 2 whorls of the initial teleoconch weak, almost straight (not folded at the peripheries of the whorls), initially closely spaced and very low, gradually becoming widely spaced and stronger; ribs on the subsequent whorls except above the aperture (ribs on most whorls except above the aperture) strong, highly protruded as spines, very widely spaced (9–11 ribs in a whorl), not synchronized with those on the previous whorls, strongly folded at the peripheries of the whorls, strongly elevated by axial protrusions of shell wall (along the ribs) on the last 3 / 4 whorls; upper margin of the ribs above the spiny projections (between the suture and spines) very low, never protruded, indistinct; spiny projections of the ribs high, pointed, folded at the peripheries of the whorls, trough-shaped, rather narrow, arising at some distance below the suture (arising near the peripheries of the whorls), upper margin folded over the lower margin, lower margin gradually decreased in height and extended below to the suture of the next whorl, with fine spiral sculptures on the outer surface, protruded upward on most whorls, becoming protruded laterally on the last 5 / 8 whorls; ribs above the aperture (ribs on the last 1 to last 3 / 4 whorls around the constriction) abruptly becoming low and closely spaced, with greatly reduced spiny projections (but still retaining very low spiny projections), less folded than those on the preceding whorl, becoming much less folded toward the aperture, partially hidden by the inner peristome. Interspace between the spiny ribs smooth on the upper whorls, with some growth wrinkles below the suture on the last 1.25–2.25 whorls; growth wrinkles developed below the suture on the lower whorls, closely spaced, short, very low, initially very weak, becoming rather prominent on the last 3 / 4 whorls. Spiral sculptures present throughout the teleoconch, fine, rather prominent, sinuous in high magnification, extending on the outer surface of the spiny ribs and outer peristome. Base rather convex, without basal fold; interspace between the ribs with a series of fine and short growth wrinkles around the columella. Axial protrusions of shell wall (along the ribs) on the last 3 / 4 whorls strongly developed, hidden by the spiny ribs in fresh specimens (becoming obvious after removal of the spiny projections of the ribs), associated with prominent axial depressions between them; abapertural side slightly or hardly convex in umbilical view; adapertural side well convex, wider than the abapertural side in umbilical view. Umbilicus closed in adult. Aperture hardly or slightly tilted against the coiling axis (in left lateral view), slightly protruded from the previous whorl; columellar axis within the aperture almost vertical or slightly oblique in front view. Columellar tooth present, very weakly appearing within the aperture in front view, but strongly developed deeply inside the aperture, never extended onto the inner peristome. Peristome triple, with an outer peristome and doubled inner peristomes. Outer peristome strongly developed, widely expanded, but hardly or only slightly expanded beyond the inner peristome in front view, obtusely angulate at the palatal side, frequently with a rounded baso-columellar edge, reflected backward, abruptly expanded from the preceding whorl, with concentric growth riblets or growth lines on the inner surface, with fine spiral sculptures on the outer surface. Inner peristome strongly protruded (separated) from the outer one, double with distal and proximal inner peristomes; distal inner peristome strongly developed just behind the proximal one, widely expanded, shortly expanded beyond the proximal one in front view; proximal inner peristome squarish circular, always narrower and weaker than the distal one, rather widely expanded, parietal margin almost straight and located near or below the middle level of the upper body whorl. Interspace between the outer and inner peristomes wide, with a series of fine riblets on the outer surface near the outer peristome, rather smooth near the inner peristome; uppermost margin (= suture) ascending onto the penultimate whorl toward the aperture in left lateral view; base extended almost horizontally in left lateral view. Dimensions. Shell height 4.5–5.3 mm, diameter 2.6–3.6 mm, suture width 1.9–2.1 mm, aperture height 1.8– 2.2 mm. Operculum (Figs. 2 G 1–4). Operculum corneous, multispiral, circular, transparent, amber colored, thin, flat; outer surface smooth; inner surface with a very weak arcuate ridge near the columellar margin. Penis. Penis absent. Radula. Not examined. Distribution and ecology. Endemic to Palau: Southern limestone coast of Babeldaob island. This species inhabits limestone rocks. Remarks. H. ngereamensis is most similar to H. triplochilus Yamazaki & Ueshima, 2013 (Figs. 12 O, P), but differs in the following points: the axial protrusions of shell wall along the ribs are strongly developed on the last 3 / 4 whorls; the riblets between the inner and outer peristomes are less developed, much lower than the latter species; the distal inner peristome is always wider and more strongly developed than the proximal inner peristome; the shell is broader and less slender; the distribution is endemic to Babeldaob island. H. ngereamensis differs from a sympatric congener, H. echinata tubulispina Yamazaki & Ueshima, 2013, in the following points: the spiny projections of the ribs are wider, never tubular; the inner peristome is double; the interspace between the outer and inner peristomes is wider; the axial ribs above the aperture have very short spiny projections. H. ngereamensis also differs from H. expansilabris Yamazaki & Ueshima, 2013, which is distributed in the same area of Babeldaob island, in the following points: the outer and inner peristomes are not so widely expanded as the latter species, more robust; the columellar tooth is present; the distal inner peristome is always wider and more strongly developed than the proximal inner peristome; the axial ribs above the aperture have short spiny projections. Etymology. The specific name is derived from the type locality. FIGURE 2. Hungerfordia ngereamensis sp. nov. (A 1–6) A topotypic paratype, Loc. BA 4 - 2, B0293a; (B 1–2) a topotypic paratype after removal of the spiny projections of the axial ribs, B0293b: (B 1) umbilical view, only the last whorl shown, note that the axial protrusions of the shell wall are strongly developed along the ribs; (B 2) oblique dorsal view, showing the axial protrusions of the shell wall on the last 3 / 4 whorls; (C 1–2) a yellowish and subfusiform specimen, a topotypic paratype, B0293c: (C 2) sculptures on the upper body whorl, showing the reduced axial ribs above the aperture; (D) columellar tooth, dorsal view of a partially opened shell, a topotypic paratype, B0293d; (E 1–6) holotype, B0293T; (F) a worn specimen, only the last whorl shown, note the axial protrusions of the shell wall which are usually hidden by spiny ribs in fresh specimens (see Fig. E 6), a topotypic paratype, B0293e; (G 1–2) outer surface, and (G 3–4) inner surface of operculum, a topotypic paratype, B0293. Scale bars, 1mm (A–F), 0.5 mm (G). All specimens, UMUTZ-MG.Published as part of Yamazaki, Midori, Yamazaki, Kazunori, Rundell, J. & Ueshima, Rei, 2015, Systematic review of diplommatinid land snails (Caenogastropoda, Diplommatinidae) endemic to the Palau Islands. (3) Description of eight new species and two new subspecies of Hungerfordia, pp. 511-538 in Zootaxa 4057 (4) on pages 514-516, DOI: 10.11646/zootaxa.4057.4.3, http://zenodo.org/record/24168

    Acauloplacella (A.) queenslandica Rentz, Su & Ueshima

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    Acauloplacella (A.) queenslandica Rentz, Su & Ueshima ANIC Number Acauloplacella sp. 1 Chirping Leaf-mimic Figs. 1; 4 A; 5 A, C; 6 A; 7; 11; Tables 1, 2; Map 1 Holotype male. 1. “ 15.50S 145.20E Gap Ck. 5 km ESE. of Mt Finnigan nr. Cooktown, Qld. 13–16.v. 1981 D. C. F. Rentz, Stop 37 ”. 2. “D. C. F. Rentz, Cytol. prep. 81-158 ” 3. “Song recorded S- 85 ”. Holotype in Australian National Insect Collection, CSIRO Entomology, Canberra. Type locality. The type locality is dense rainforest with an understorey of scattered palms, tree ferns and seedlings of a number of rainforest trees. The type was collected from a tangle of Lawyer Palm (Calamus sp.) approximately 2.5 m from the ground. Differential diagnosis. Size small for genus, form robust. Carina present on occiput; antenna scape with tooth; antennal flagellum not annulate; only scattered tubercles on pronotal disk; no tubercles on femora; male titillator simple, tongue-like; tip of ovipositor with carinae and tubercles. Male. Head. Head with prominent median carina on occiput continuous to fastigium of vertex which projects as a minute spine; frons flat, smooth, somewhat concave; genae in region of eye, sparsely tuberculate. Antenna with scape projecting well in front of fastigium of vertex, internal margin carinate and ending in a prominent tooth; pedicel 1 / 3 to 1 / 4 length of scape, with a small prominent tubercle on outer margin. Thorax. Pronotum evenly but sparsely tuberculate, median carina comprising coalesced tubercles; surface of disk traversed by a single, deep transverse sulcus positioned near the middle of the disk and continuous onto lateral lobes; ventral margin of lateral lobe beaded, the line of beads extending onto adjacent gena. Prosternum unarmed; mesosternum with margins smooth, the transverse ridge (cephalic margin) serrate only on cephalic humeral angles; meso- and metasterna each bearing a pair of prominent pits. Auditory tympanum minute, directed posteriorly. Wings. Tegmina (Fig. 4 A) short, broad, well bowed on anterior margin; stridulatory vein very thick, much broader basally than distally; mirror of left tegmen strongly coriaceous, with an arching diagonal vein, mirror of right tegmen ill-defined, but coriaceous and with a pair of parallel veins running obliquely caudad from diagonal vein. Legs. Fore leg with femur feebly serrate on dorsal margin, tibia unspined. Middle leg with femur dorsally carinate on both margins, ventral surface feebly serrate on both margins, posterior surface lacking tubercles; tibia dorsally tuberculate on both margins, without spines, ventral surface serrate only on anterior margin. Hind femur with external pagina irregular but not sulcate, dorsal surface carinate, ventral surface evenly serrate on external margin, internal margin not serrate; tibia quadrate, dorsal surface feebly serrate, the serrations more concentrated apically, ventral surface smooth, apex armed with a minute spine on each side of ventral surface. Abdomen. Tenth tergite with apex irregular, poorly defined; supra-anal plate elongate, tongue-like extending well beyond apex of abdomen, with median carina, apically acute, surface sparsely hirsute; subgenital plate (Fig. 5 A) prominently scoop-shaped, lacking a shaft, styles elongate, inflated in life. Cercus flimsy, very elongate, shrivelled in all dry material, appearing to have a stout basal tooth directed upwards, however, this tooth is not a part of the cercus but probably a paraproct. Titillator (Fig. 5 C), represented as a minutely shagreened quadrate pad, longer than broad, in some specimens appearing divided. Female. Differs from male in following: supra-anal plate ovoid, with median carina, covering cerci and base of ovipositor; subgenital plate small, broadly triangular, sides undulating, with an internal ridge, apex indented. Ovipositor with base broadly expanded, lateral margins concave; valves heavily serrate on dorsal margin (Fig. 6 A), ventral margin minutely serrate; sides of ovipositor valves with 4 minute carinae and tubercles. Colour. Pale green in life (Fig. 1), thorax of some specimens with a darker green circular mark on metepisternum. Cells of tegmina each with a prominent darker green spot or not. Legs light green. Beaded area of ventral lobe of pronotum and occiputal carina and adjacent gena yellow. Specimens examined. Paratypes: Queensland: 12 o 43 ’S. 143 o 18 ’E. 1 km ENE. of Mt Tozer, nr. Iron Range Nat. Pk., 11.vii. 1986 (D. C. F. Rentz, stop I- 20, 1 female from pyrethrin fog, ANIC). 12 o 43 ’S 141 o 17 ’E. 5km ENE. of Mt. Tozer, Iron Range Nat. Park, Qld. 10.vii. 1986 (D. C. F. Rentz, Stop I- 12, 1 male, collected as nymph, matured 14.vii. 1986, Cytol. Prep. 86-101, Song recorded S- 408, 1 female nymph, ANIC). 12 o 43 ’S 143 o 17 ’E. 9 km ENE. of Mt Tozer, Iron Range Nat. Pk, 7.vii. 1986 (D. C. F. Rentz, 1 male, 1 female, ANIC). Iron Range, 26–31.v. 1971 G. B. Monteith, 1 female, QUEN). Claudie Creek area, Claudie River Dist., 22–23.vi. 1982 (M. A. Schneider, G. Daniels, 1 male, UQEN). Packers Creek, via Portland Rds, 6.xii. 1985 (G. B. Monteith, D. Cook, 1 female, pyrethrin fog, QMUS). 16 o 48 ’S. 145 o 38 ’E. (GPS) Kuranda, 335 m elev., Top of the Range, 19 Butler Dr, 15–30.iv. 2010 (D. C. F. Rentz, 1 male, ANIC). Kuranda, Black Mtn Rd, 12.iv. 1989 (J. Hasenpusch, 1 male, QMUS). Cape Tribulation, 29.xii. 1982 – 8.i. 1983 (G. B. Monteith, 1 male, QMUS). Song. The song of this species is a continuous chirp uttered after dark. Each chirp consists of two pulses (Fig. 7). Comments. This is a small species showing variation in the degree of tuberculation of the pronotum. About half of the specimens have no trace of the tubercles on the genae that are possessed by the holotype. The male titillator appears to have a thin median line which suggests it is divided, however, this is indicated solely by colour. Acauloplacella (A.) hasenpuschae 1 Rentz, Su & Ueshima ANIC Number Acauloplacella sp. 2 Sue’s Leaf-mimic Figs. 2 A; 4 B, C; 5 B; 6 C, 8 A–E; Tables 1, 2; Map 1 Holotype male. 1. “ 17 o 28 ’S 146 o0’E. (GPS) Qld. Garradunga, Polly Creek, nr Innisfail 18.xi. 1997 J. Hasenpusch”. Holotype in Australian National Insect Collection, CSIRO, Canberra. Type locality. The type locality is mixed coastal rainforest with an abundance of vines. Differential diagnosis. Size moderate for genus, form elongate; carina absent on occiput but with sulcus; no tooth on antennal scape; tubercles absent on head; antennal flagellum with a few spots beyond 5 th flagellar segment; tubercles and pattern characteristic on pronotal disk; two sulci on pronotal disk; tubercles present on fore femur; titillators poorly, if at all, sclerotised; ovipositor without tubercles or ridges laterally. Male. General. This is a moderately large species (Table 1) easily recognised by the relatively broad, flat central portion of the pronotum which is emphasised by yellowish colour (Fig. 2 A) and the cylindrical male cerci with a subapical hook (Fig. 5 B). Females have the ovipositor with minute serrations on the dorsal and ventral margins at the tip and no tubercles or ridges on the sides (Fig. 6 C). Head. Head smooth, without tubercles; fastigium of vertex projecting, surface deeply and broadly sulcate; occiput without any trace of carina; frons smooth, not concave. Antenna with scape quadrate, projecting well in front of fastigium of vertex, internal margin carinate only apically, surface near apex concave and lacking an apical tooth; pedicel quadrate, about 1 / 3 length of scape, with a prominent tubercle at base on internal margin. 1. Named in honour of Mrs Sue Hasenpusch, a friend and collector of many orthopteroids in the Innisfail region of north Queensland. Thorax. Pronotum with surface sparsely tuberculate, the tubercles not arranged in any pattern; surface of disk faintly flattened and lacking a median carina but with a single deep transverse sulcus on cephalic boundary of metanotum, this preceded by a much shallower and less defined and broader sulcus; lateral lobes feebly irregularly tuberculate. Prosternum unarmed; meso- and metasternum each more poorly indicated than in A. queenslandica Rentz, Su & Ueshima sp. nov; frontal ridge serrate across its entire margin. Auditory tympanum relatively prominent, circular, directed posteriorly. Wings. Tegmina (Fig. 4 B,C) short, broad, with prominent bow on anterior margin. Stridulatory vein short, thick, not raised; mirror weakly coriaceous and with a strong central Y-shaped longitudinal vein; right tegmen with mirror smooth and without any central venation, caudal portion with a small amount of coriaceous venation not extending much beyond border. Legs. Fore leg with femur with sharp dorsal carina, not serrate, ventral surface with carinae lacking serrations, posterior surface bearing 5 prominent tubercles of equal size; tibia unspined. Middle leg with femur not dorsally carinate on either margin, ventral surface with only a suggestion of serrations apically on anterior margin. Hind leg with external pagina smooth and with a longitudinal sulcus over its entire length; tibia quadrate, somewhat swollen in proximal half, apex on ventral surface with a minute spine on each side. Abdomen. Tenth tergite (Fig. 8 A) with apical margin feebly irregular; supra-anal plate (Fig. 5 B, 8 B) very elongate, at least 2.5x as long as wide, base raised, with a short feeble median carina, apex slightly concave, margins of plate bearing long hairs; paraproct with a minute, short, slender spine, barely discernable. Cercus (Fig. 5 B) prominent, rigid, short stout, cylindrical, with a minute tooth subapically, surface hirsute. Titillators represented as divided, elongate, extremely feebly, if at all, shagreened fleshy process, somewhat twisted at apex and ovoid at base; apices of adjacent fleshy lobes clothed with long hairs. Subgenital plate (Figs. 8 A, B) extending upwards to or near the level of the abdomen, but not beyond; ventral surface of plate with a feeble subapical carina (Fig. 8 C), stalk short, styles strongly cup-shaped, densely hirsute. Female. Differs from male in following: supra-anal plate extending as an elongate flange covering most of genitalia including base of ovipositor; subgenital plate broadly triangulate, sides with distinct ridges. Ovipositor (Fig. 6 C) minutely serrate on both margins, external surface without any trace of tubercles or carinae. Eggs. The eggs are laid in wood, preferably bark cracks (Fig. 8 D, E). They are flat and ovoid with a peculiar vesiculated area that protrudes from the surface once the egg has been laid. Colour. In life pale green, with yellow as indicated (Fig. 2 A); legs greenish yellow; tegmen with most veins yellow, central portion of larger cells with a darker green spot; head with fastigium of vertex and adjacent head and pronotum yellow and brown. Specimens examined. Paratypes: Queensland: 17 o 28 ’S. 146 o00’E. (GPS) Garradunga, Polly Creek, nr Innisfail, 10.xii. 1998, 27.xi. 1993, 10.ii. 1994, 26.x. 1994, 10.xii. 1994, 12.xii. 1994, 3.ii. 1995, 10.xii. 1995, 15.xi. 1996, 18.xii. 1997, 14.iii. 1999, 29.ii. 2000 (J. Hasenpusch, 4 males, 8 females, ANIC). Bellenden Ker Range, Cableway base station, 100 m elev., 17–24.x. 1981 (Earthwatch group, 1 female, QMUS). 19 o 14 ’S. 146 o 29 ’E. Bluewater Creek, nr Townsville, 11.iv. 1990 (DCF Rentz, stop 14, 1 female, ANIC). 16 o 36 ’S. 145 o 20 ’E. “Kingfisher Park”, 1 km N. of Julatten, 22.iv. 1997 (C. J. Burwell, 1 female, QMUS). Comments. The tubercles described on the fore femur of the type are present on fewer than half of the males seen in this study. The ventral surface of fore femora may be serrate on one or both margins. This species is very distinctive in both colour and morphology and there is little chance in getting it confused with the others. Records suggest that Acauloplacella reaches its southern limits in the coastal forests north of Townsville. Acauloplacella (A.) incisa 2 Rentz, Su & Ueshima Etched Leaf Mimic Figs. 2 B; 4 D; 6 B; 9 A–G; Tables 1, 2; Map 1 Holotype male. “ 16 o 49 ’S 145 o 41 ’E. QLD. Smithfield, James Cook University, nr Bldg E 2 30 APR 2009 DCF Rentz, G Wilson.” Holotype in Australian National Insect Collection, CSIRO Entomology, Canberra. Type locality. The type locality is described as a simple to complex mesophyll to notophyll vine forest on moderately to poorly drained metamorphics (excluding amphibolites) of moderate fertility of the moist and wet lowlands, foothills and uplands. Differential diagnosis. Size moderate for genus, largest of known Australian species (Table 1). Most specimens have a bright white median stripe on the pronotum, extending onto the head. The colour is deep green in this species with the central portion of most cells with a dark spot (Fig. 2 B). Carina on occiput either present or absent; antennal scape without tooth; head without tubercles; antennal flagellum without 2. Named with reference to the median incision of the male tenth tergite. annulations; tubercles on pronotal disk not forming a median carina; disk of pronotum with only median sulcus; titillators very complex; ovipositor apically with tubercles and carinae as in a. A. queenslandica. Male. General. Head. Head smooth except for a few minute tubercles beneath eye (Fig. 9 B); fastigium of vertex projecting, surface sulcate only at tip (Fig. 9 A); occiput with median carina indicated mostly by colour. Antennae with scape quadrate, projecting well in front of fastigium of vertex, internal margin feebly carinate and usually lacking an apical tooth (Fig. 9 A, with tooth). (See Table 2); pedicel quadrate, small, about 1 / 5 length of scape, with a small swelling at base on internal margin. Character queenslandica Incisa hasenpuschae mecyna 1. Size Small (Table 1) Large (Table 1) Moderate (Table 1) Large (Table 1) 2. Carina on occiput Present (Fig. 1) Absent or poorly Absent, with sulcus Indicated by colour, with developed, with sulcus sulcus (Fig. 10 A) (Fig. 9 A) Thorax. Pronotum with surface sparsely tuberculate, the tubercles much more reduced in size and number than in any other Australian species, the median carina topped by a series of tubercles; lateral lobes with a row to tubercles or serrations along ventral margin (Fig. 9 B); cephalic and caudal margins of disk projecting and lacking tubercles; surface of disk with a thin transverse sulcus. Prosternum unarmed; meso- and metasternum each prominent, frontal ridge lacking any serrations. Auditory tympanum barely discernable. Wings. Tegmina very elongate (Fig. 4 D, Table 1), evenly bowed on anterior margin. Stridulatory vein extremely thickened, especially at base, mirror and remainder of stridulatory area heavily coriaceous and not really distinct from adjacent venation; right tegmen with mirror poorly defined and not coriaceous. Legs. Fore leg with femur with a sharp median dorsal carina, minutely serrate or not, ventral surface lacking carinae, but minutely serrate or not on anterior or posterior margins. Middle leg with femur with a low median carina dorsally, surface irregular, ventral surface with anterior margin carinate and minutely serrate, posterior surface without tubercles. Hind leg with outer pagina of femur with a deep, oblique longitudinal sulcus, dorsal surface with an elongate median carina, ventral surface with out margin serrate for entire distance. Abdomen. Tenth tergite with apical margin with a prominent U-shaped median incision (Fig. 9 C); supraanal plate triangular, about 3 x longer than broad, apically acute and feebly directed upwards, surface with a longitudinal carina for its entire length; paraprocts a minute spine directed dorsally and completely hidden by cercus. Cercus elongate, very similar to that described for A. queenslandica. Titillator (Figs. 9 F, G) a complex well sclerotised structure comprising a broad “tongue” followed by a “crest” (seen as a dark brown line in Figs. 9 F), surrounding fleshy lobes apically bearing long hairs. Subgenital plate (Fig. 9 D) with ventral surface without any trace of median carina, in other respects similar to that described for A. hasenpuschae. Female. Differs from male in following: supra-anal plate chordate, with median carina, extending to base of ovipositor; cercus elongate, flimsy; subgenital plate (Fig. 9 E) short, quadrate, with lateral ridges, sides parallel in most specimens. Ovipositor (Fig. 6 B) with relatively large dorsal serrations, minute ventral serrations concentrated at apex; sides of ovipositor with 2 dorso-ventral carinae and 1–3 tubercles near apex. Colour. Dark green in life with linear sequence of darker green spots on tegmina in most specimens. Median carina of head and pronotum ivory white on majority of specimens. Abdomen yellow with green rings. Ovipositor dark brown on apical dorsal and ventral margins and on ridges and tubercles, remainder straw brown. Specimens examined. Paratypes: Queensland: 12 o 43 ’S. 143 o 18 ’E. 11 km ENE. of Mt Tozer, nr Iron Range Nat. Pk, 11.vii. 1986 (DCF Rentz, Stop I- 10, 1 male, ANIC). Daintree, 6.vi. 1960 (T. G. Campbell, 1 male, ANIC). 16 o 48 ’S. 145 o 38 ’E. (GPS) Kuranda, 335 m elev., Top of the Range, 19 Butler Dr, 15– 30.iv. 2003; 15–31.iii. 2004, 1– 15.vi. 2004, 15– 31.i. 2005, 1– 15.ii. 2006, 1– 15.xi. 2007, 16– 31.vii. 2008, 1– 15.xii. 2009 (DCF Rentz, 1 male, 7 females, ANIC). 16 o 48 ’S. 145 o 37 ’E. (GPS) Kuranda (Caravan Park) 11.xii. 2004, 7.v. 2007 (DCF Rentz, 2 males, ANIC). Kuranda, 12.ii. 2000 (J. Hasenpusch, 1 male, ANIC). 16 o 49 ’S. 145 o 37 ’E. (GPS) 16 o 49 ’S. 145 o 37 ’E. Kuranda, (Village Service Station), 26.ii. 2004 (DCF Rentz, 1 female, ANIC). Acauloplacella (P.) mecyna 3 Rentz, Su & Ueshima Iron Range Leaf Mimic Figs. 3; 4 E; 10 A–F; Tables 1, 2; Map 1 Holotype male. 1. “ 12.49S 143.20E 4km S. of Lockhart River (settlement) nr. Iron Range Nat. Park, QLD. 10 July 1986 D. C. F. Rentz. Stop I- 17 ” 2. “Collected as nymph, matured in laboratory 28.vii. 1986 ”. Holotype in Australian National Insect Collect, CSIRO Entomology, Canberra. Type locality. The type locality is in an area of mixed vegetation consisting of mangroves along the river with Blady Grass (Imperata cylindrica) and eucalypts beyond. The type was collected on a low shrub of undetermined status. Differential diagnosis. Size large for known Australian Acauloplacella species (Table 1). Unique colour and pattern but readily placed in the subgenus Papuaprium on the basis of the characters noted in the introduction. Known only from male type. Sulcus on occiput but lacking carina, indicated only by colour; no tooth on antennal scape; tubercles absent on head and femora; antenna annulate; colour on pronotum and tegmina distinct, tubercles forming median carina; pronotum with single deep median sulcus; titillators a curving tongue-like structure. Male. General. Head. Head smooth, without tubercles; fastigium of vertex projecting, surface depressed and shallowly sulcate; occiput with a thin, somewhat irregular carina; frons smooth, not concave. Antenna with scape quadrate, projecting only slightly in front of fastigium of vertex, internal margin bearing a short 3. Named with reference to the very elongate “stalk” of the male subgenital plate. tooth, cephalic margin irregular, raised and carinate; pedicel cylindrical, not modified, and half the length of scape. Flagellum of antenna faintly annulate. Thorax. Pronotum with surface very sparsely tuberculate, the tubercles minute and not forming any pattern; median carina (Fig. 10 A) formed by a series of tubercles (Fig. 10 D); lateral lobes smooth along cephalic and caudal margins, ventral margin heavily serrate. Surface of disk with metasternum defined by a deep transverse sulcus, lateral lobe with a sulcus bordering pro- and mesosterna. Prosternum unarmed, meso- and metasterna each with a pair of pits; frontal ridge irregular but not serrate. Auditory tympanum relatively large, ovate, directed posteriorly. Wings. Tegmina elongate, with a prominent bow on anterior margin (Fig. 4 E). Stridulatory vein short, thick, with an accessory vein preceding; mirror weakly coriaceous and with 7 cells each with a black spot formed by a coalescence of minute dark brown to black spots; right tegmen with mirror with a weak diagonal vein and with a number of more or less parallel veins extending caudad, dark spots completely lacking. Legs. Fore leg with femur bearing a prominent dorsal carina, ventral surface weakly serrate on anterior margin, posterior pagina with a few tubercles at base; tibia serrate on anterior margin of ventral surface, auditory structure swollen. Middle leg with femur smooth on dorsal surface, ventral surface serrate on anterior margin, posterior margin smooth, pagina smooth; tibia distinctly flattened and flanged, anterior surface sulcate, ventral margin serrate, dorsal surface with tubercles on both margins. Hind leg with outer surface of femur smooth and with a median longitudinal sulcus over its entire length, ventral surface serrate along entire outer margin; tibia quadrate and flanged on outer ventral margin at base, internal margin with a few minute serrations. Abdomen. Tenth tergite (Fig. 10 B, C) with apical margin irregular, not indented; supra-anal plate (Fig. 10 B) apex indented. Cercus very elongate, cylindrical, tapering to apex; paraprocts short, stout, directed laterally, appearing to emanate from base of cercus. Titillator (Fig. 10 E) with central sclerotised portion a single curing tongue, well sclerotised and carinate dorsally (Fig. 10 F). Subgenital plate (Fig. 10 C) indented at base, the corners rounded; stalk extraordinarily elongate, with very narrow med

    Ganiagraecia karwinia Rentz, Su & Ueshima, 2021, Gen. et sp. nov.

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    Ganiagraecia karwinia 0 Rentz, Su and Ueshima Gen. et sp. nov. ANIC Number, Gen. Nov. 9, sp. 9 Figs 9 A-K; Figs 20C, D; Fig. 21A; Fig. 22b A; Table 2, Map 4 Common name. Grass Tree Katydid Holotype male. (In alcohol) 1. “ 25.26S 152.56E Mt. Tibrogargan (parking area), Glass House Mts. Qld. 15 August 1985 D. C. F. Rentz, R. Chiang, Stop 12”. 2. “collected as nymph, matured in laboratory 16.x.85”. 3. “ D. C. F. Rentz, Cytol. prep. 85-29”. 3. “Song recorded S-363”. 4. “ ANIC database #14008749”. Holotype in alcohol in Australian National Insect Collection, Canberra. Paratypes. Queensland. 25 o 26’S 152 o 56’E Glass House Mts, Mt Tibrogargan, (parking area), 15.viii.1985 (DCF Rentz, R. Chiang, Stop 12, 10 males, collected as nymphs, matured in laboratory 16.x, 18.x, 20.x, 25.x.1985; DCF Rentz, Cytol. Prep. 85-18, -19, -20 -21, -27, -28; 5 females, same locality data and maturation dates, ANIC). 26 o 54’S 152 o 56’E Glass House Mts, Mt Ngungun (parking area), 9.viii.1985 (DCF Rentz, R. Chiang, Stop 4, 2 males, collected as nymphs, matured in laboratory 18.x, 20.x.1985; 13 females, collected as nymphs, matured in laboratory 18.x, 20.x.1985; Cytol prep. 85-17, ANIC). Type locality. The type locality is a rather open woodland with many Grass Trees, Xanthorrhoea sp., tall eucalypts and a dense undercover of herbs and grasses. The katydids were abundant in the lush weedy vegetation. Measurements. Table 2 Differential diagnosis. Male. Antenna thin, about 5 times the length of body, annulate. Eye prominent, positioned high on head (Figs 9A, B). Cephalic margin of pronotum straight, caudal margin feebly concave. Prothorax with a prominent pair of thin spines; meso- and metathorax each bearing prominent tubercles (Figs 9D, G). Hind tibia armed dorsally on each side with many spines, packed close together, ventral surface with 6 or 7 very thin spines on each side. Cercus (Figs 9I, J) short, stout, with single internal tooth, apex acute, suggesting a minute tooth; subgenital plate with styles more robust than normal (Fig. 9M), median incision shallow, V-shaped. Female. Differs from male in following. Thorax with a depression (glandular opening?) on the posterior portion of the mesothorax (Fig. 9D). Cercus distinctly curved inwards (Fig. 9N); subgenital plate about as broad as long, acutely tapering to apex, apex shallowly incised (Fig. 9O). Ovipositor longer than length of body, weakly upcurved, unarmed (Table 2). Colouration. Overall colouration straw brown with minute greyish brown speckles over entire dorsal portion of body; ventral surface of body creamish white (Figs 9 A-B, D, G), subgenital plate light brown. Males with fastigium of vertex and median portion of pronotum with a black stripe (Fig. 9A), caudal half of pronotum black. Tegmina thin, colourless except for internal apical portion abdomen immediately beneath apex of tegmina dark (Fig. 9A, K). Female with head with brown stripe and with a thin white stripe in the middle (Fig. 9C); dark brown stripe extending from cephalic margin of pronotum to almost apex of abdomen (Fig. 9B), but not including tenth tergite. Distribution. Known only from the Glasshouse Mts., Queensland (Map 4). Habitat. Common in open woodland vegetated by tall eucalypts and Xanthorrhoea. Seasonal occurrence. Adults found in mid spring. Stridulatory file. File very distinctive (Figs 20C, D). Proximal portion with approximately 20 large teeth, decreasing in size and approximately 24-26 minute teeth on a raised swelling. 10 Karwin is an aboriginal word for Grass Trees, Xanthorrhoea species. Karyotype. 2n male= 31 (30t +Xt) Fig. 22b A Recognition. Very easily recognised by the combination of the yellow brown body colour with dark brown median dorsal stripe which contrasts with the distinct colour of the pronotum (Fig. 9L). Males have a simple cercus (Figs 9I, J) with a prominent internal tooth. Females bear a depression (glandular opening?) on the posterior portion of the mesothorax (Fig. 9D). The female cercus has a distinctive incurved tooth-like apex (Fig. 9N). Discussion. This species appears to be endemic to the Glasshouse Mts. Its appearance and habitat preference is similar to that of Australiagraecia species but morphologically very different.Published as part of Rentz, Dcf, Su, You Ning & Ueshima, Norihiro, 2021, Studies in Australian Tettigoniidae: New short-winged Agraeciini from Australia (Orthoptera: Tettigoniidae; Conocephalinae; Agraeciini), pp. 1-72 in Zootaxa 5059 (2) on pages 18-19, DOI: 10.11646/zootaxa.5059.1.1, http://zenodo.org/record/560250

    FIGURE 8 in Studies in Australian Tettigoniidae: Australian agraeciine katydids, two new genera from northern Australia (Tettigoniidae; Conocephalinae; Agraeciini)

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    FIGURE 8. Male cerci of Armadillagraecia and Kapalgagraecia species Rentz, Su, Ueshima, Robinson gen. et sp. nov. A, A. mataranka, dorsal view tip illustrating curvature; B, same species internal view. C, A. yerilla Rentz, Su, Ueshima, Robinson sp. nov., dorsal view illustrating curvature at tip; D, same species internal view. E, same species ventral view. F, A. triodiae Rentz, Su, Ueshima, Robinson sp. nov. dorsal view tip illustrating curvature. G, same species illustrating internal tooth. H, K. brayi Rentz, Su, Ueshima, Robinson sp. nov., dorsal view holotype male. I, K. nauma Rentz, Su, Ueshima, Robinson sp. nov., Pt Keates, NT, dorsal view. J, same internal. K, same ventral view.Published as part of <i>Rentz, D.C.F., Su, You Ning, Ueshima, Norihiro & Robinson, Martyn, 2010, Studies in Australian Tettigoniidae: Australian agraeciine katydids, two new genera from northern Australia (Tettigoniidae; Conocephalinae; Agraeciini), pp. 1-39 in Zootaxa 2417 (1)</i> on page 25, DOI: 10.11646/zootaxa.2417.1.1, <a href="http://zenodo.org/record/5307038">http://zenodo.org/record/5307038</a&gt
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