25,882 research outputs found
Polypheretima mekongmontis Nguyen, Tran & Nguyen 2014
No full text<i>Polypheretima mekongmontis</i> Nguyen, Tran & Nguyen, 2014 <p> <i>Polypheretima mekongmontis</i> Nguyen, Tran & Nguyen, 2014: 118, Fig. 6.</p> <p> <b>Type locality.</b> Vietnam (Kien Giang: Hon Dat).</p> <p> <b>Type material.</b> CTU (EW.028), Vietnam.</p> <p> <b>Examined material.</b> 1 C (<b>CTU-EW.028.h01</b>) and 10 C (<b>CTU-EW.028.p02</b>) Kien Giang Prov., Hon Dat District, Hon Me Mt. (10º06' 12.3N; 104º52' 43.5E), mango garden, 13/11/2010, coll. Nguyen Thanh Tung.</p> <p> <b>Records from Vietnam. Kien Giang</b> (Hon Dat, Hon Tre Isl.) (Nguyen <i>et al.</i> 2014).</p> <p> <b>Distribution.</b> Only known from Vietnam.</p> <p> <b>Vietnamese name.</b> Trùn núi mêkông.</p>Published as part of <i>Nguyen, Tung T., Nguyen, Anh D., Tran, Binh T. T. & Blakemore, Robert J., 2016, A comprehensive checklist of earthworm species and subspecies from Vietnam (Annelida: Clitellata: Oligochaeta: Almidae, Eudrilidae, Glossoscolecidae, Lumbricidae, Megascolecidae, Moniligastridae, Ocnerodrilidae, Octochaetidae), pp. 1-92 in Zootaxa 4140 (1)</i> on page 74, DOI: 10.11646/zootaxa.4140.1.1, <a href="http://zenodo.org/record/256507">http://zenodo.org/record/256507</a>
Amynthas nhonmontis Nguyen & Nguyen 2015
No full text<i>Amynthas nhonmontis</i> Nguyen & Nguyen, 2015 <p> <i>Pheretima nhonmontis</i> Nguyen & Nguyen, 2015: 132, Fig. 2.</p> <p> <b>Type locality.</b> Vietnam (An Giang: Tinh Bien).</p> <p> <b>Type material.</b> CTU, Vietnam.</p> <p> <b>Examined material.</b> 1 C (<b>CTU-EW.023.h01</b>) and 6 C (<b>CTU-EW.023.p02</b>) natural forest, Nhon mountain, (104°56'09.2"E, 10°35'39.6"N), 56 m a.s.l., Tinh Bien district, An Giang province, Vietnam, 7/11/2010, coll. Nguyen Thanh Tung.</p> <p> <b>Records from Vietnam. An Giang</b> (Tinh Bien) (Nguyen & Nguyen 2015).</p> <p> <b>Distribution.</b> Only known from Vietnam.</p> <p> <b>Vietnamese name.</b> Giun núi Nhọn</p>Published as part of <i>Nguyen, Tung T., Nguyen, Anh D., Tran, Binh T. T. & Blakemore, Robert J., 2016, A comprehensive checklist of earthworm species and subspecies from Vietnam (Annelida: Clitellata: Oligochaeta: Almidae, Eudrilidae, Glossoscolecidae, Lumbricidae, Megascolecidae, Moniligastridae, Ocnerodrilidae, Octochaetidae), pp. 1-92 in Zootaxa 4140 (1)</i> on page 40, DOI: 10.11646/zootaxa.4140.1.1, <a href="http://zenodo.org/record/256507">http://zenodo.org/record/256507</a>
Metaphire chaubinhensis Do & Tran 1994
No full text<i>Metaphire chaubinhensis</i> (Do & Tran, 1994) <p> <i>Pheretima chaubinhensis</i> Do & Tran, 1994 (in: Do <i>et al.</i> 1994): 24, fig. 1B; Do 1994: 64; Nakamura 1999: 62; Thai 2000a: 308; Huynh & Nguyen 2004a: 115.</p> <p> <i>Metaphire chaubinhensis—</i> Blakemore 2007a: 22; Blakemore 2008b.</p> <p> <b>Type locality.</b> Vietnam (Nghe An: Quy Chau).</p> <p> <b>Type material.</b> SORC, Vietnam.</p> <p> <b>Records from Vietnam. Hanoi</b> (Ba Vi NP); <b>Nghe An</b> (Quy Chau) (Do <i>et al.</i> 1994; Do 1994; Huynh & Nguyen 2004a).</p> <p> <b>Distribution.</b> Only known from Vietnam.</p> <p> <b>Vietnamese name.</b> Giun châu bình.</p> <p> <b>Remarks.</b> Blakemore (2007a, 2008b) considered it as " <i>species inquirenda</i> ", further examination was not possible because all previously collected material has been lost.</p>Published as part of <i>Nguyen, Tung T., Nguyen, Anh D., Tran, Binh T. T. & Blakemore, Robert J., 2016, A comprehensive checklist of earthworm species and subspecies from Vietnam (Annelida: Clitellata: Oligochaeta: Almidae, Eudrilidae, Glossoscolecidae, Lumbricidae, Megascolecidae, Moniligastridae, Ocnerodrilidae, Octochaetidae), pp. 1-92 in Zootaxa 4140 (1)</i> on page 53, DOI: 10.11646/zootaxa.4140.1.1, <a href="http://zenodo.org/record/256507">http://zenodo.org/record/256507</a>
Amynthas minhdam Nguyen & Tran & Lam & Nguyen 2020, sp. nov.
No full text<i>Amynthas minhdam</i> Nguyen & Tran, sp. nov. <p>(Figure 5)</p> <p> <b>Examined material.</b> <i>Holotype</i>: 1 mature (complete body), (CTU-EW.168.h01), natural forest (10°24’21”N, 107°16’18”E), 33 m, the foothill of Minh Dam Mts, Long Hai town, Long Dien District, Ba Ria – Vung Tau Province, 25 October 2016, coll. Nguyen Phuc Hau. <i>Paratypes</i>: 1 mature (CTU-EW.168.p02), same data as for holotype; 3 matures (complete body) (CTU-EW.168.p03) and 2 matures (IEBR-EW.168.p03) same data as for holotype, but 17 October 2019, coll. Lam Hai Dang.</p> <p> <b>Etymology.</b> <i>minhdam</i>, a noun in apposition, to emphasize the Minh Dam Mountain where the type specimens were collected.</p> <p> <b>Diagnosis.</b> Small-medium size, length 59–65 mm, diameter 3.0–5.0 mm, segments 75–87. Prostomium 1/3 epilobous. First dorsal pore in 12/13. Setal numbers: 48–53 in viii, 63–67 in xxx, 7–9 between two male porophores in xviii. Two pairs of spermathecal pores located lateroventrally in intersegments 5/6/7. One medioventral pre-setal genital marking, often accompanied by one pair of markings on setal ring in vii. Ventral distance between two male pores about 0.3x circumference. A pair of genital markings located medially in xviii. Holandric. Testis sac connected. Intestinal caeca simple. Septa 8/9/10 absent.</p> <p> <b>Description.</b> Body cylindrical, small size, length 59–65 mm, diameter 3.0–5.0 mm, weight 0.2–0.25 gr, segments 75–87. Body transparent, uniformly pale, clitellum light brown. Prostomium 1/3 epilobous. First dorsal pore in 12/13. Setae perichaetine, pre-clitellar setae stouter and sparser than post-clitellar ones; setal numbers: 48–53 in viii, 63–67in xxx, 7–9 between male porophores in xviii; setal distance: aa>ab, zz>zy. Clitellum within xiv–xvi, without dorsal pores and setae. Female pore single, in midventral xiv.</p> <p>Two pairs of spermathecal pores located lateroventrally in intersegments 5/6/7. One genital marking in vii, presetal and medioventral, an additional pair of genital markings on the setal ring vii, behind spermathecal pores. Male porophores located on the setal ring xviii; ventral distance between two male pores about 0.3X body circumference. A pair of genital markings (same size as male porophores) located medially in the setal ring xviii.</p> <p>Septa 5/6/7/8 thick, 8/9/10 absent, 10/11/12/13 thick. Oesophageal gizzard within viii–x. Intestinal origin at xv; caeca simple, originating at xxvii and extending anteriorly to xxv. Last hearts in xiii. Pharyngeal micronephridia developed in 5/6/7. Typhlosole simple, lamelliform. Lymph glands absent.</p> <p>Spermathecae paired in vi and vii. Ampulla oval-shaped; ducts about 1/3 ampulla. Diverticula thin, shorter than ampulla, directly attached to the base of ampullar duct; seminal chamber small, sub-divided by constrictions into 4–5 parts (chain-formed). Spermathecal ducts without nephridia. Accessory glands mushroom-shaped in vii, in association with genital markings.</p> <p>Holandric. Testis sacs developed, connected in x–xi. Seminal vesicles well developed in xi–xii. Ovaries well developed in 12/13. Ovisacs invisible. Prostate glands deeply lobuled, paired in xvii–xx; ducts long, waved and directly attached to body wall. A pair of large, mushroom-shaped accessory glands in association with genital markings in xviii.</p> <p> <b>Habitat.</b> The species was found under the litter layers, but on the soil surface.</p> <p> <b>Remarks.</b> The species can be assigned to the <i>A. morrisi</i> group due to having two pairs of spermathecal pores in 5/6/7 and holandry (Sims & Easton 1972). To date, this group comprises 47 species, of which 13 have been recorded in northern and central Vietnam (Thai 1982, 1984a,b; Le 1995; Pham 1995; Do <i>et al</i>. 1994, 1995; Nguyen <i>et al.</i> 2016a).</p> <p> Within the <i>A. morrisi</i> group, the new species is fairly similar to <i>A. sapinianus</i> (Chen, 1946) and <i>A. morrisi</i> (Beddard, 1892) in having round genital markings paired in xviii, medially of the male pores, spermathecal pores located lateroventrally and caeca simple. <i>A. minhdam</i> <b>sp. nov.</b> differs from <i>A. sapinianus</i> in the presence of genital markings in the spermathecal region and the chain-shaped seminal chamber of the spermathecal diverticulum (vs. absence of genital markings, clavate-shaped seminal chamber) (Chen 1946).</p> <p> <i>A. minhdam</i> <b>sp. nov.</b> is also differentiated from <i>A. morrisi</i> in having one pair of genital markings located ventrally next to the male pores, on the setal ring of xviii, first dorsal pore in 12/13, and a chain-shaped seminal chamber. Instead, <i>A. morrisi</i> has the first dorsal pore in 10/11 or 11/12 and a clavate-shaped seminal chamber (Blakemore 2002, Chang <i>et al</i>. 2016). The genital markings are very variable in <i>Amynthas morrisi</i> (Chang <i>et al.</i> 2016), but these variations do not appear to include the pattern characteristic of the new species. <i>A. morrisi</i> has pre-clitellar genital markings paired and/or unpaired small discs varying in numbers and positions; paired on some of vi–ix, median to spermathecal pore; unpaired on some of v–ix, midventral, pre-setal; and post-clitellar markings of two immediately median to each male porophore, one pre-setal and one post-setal, one of the two sometimes absent, slightly larger than the porophore, surrounded by several concentric furrows.Additional presetal markings on xviii and xix in some specimens, mid-ventral or median to male pores, numbers variable (Chang <i>et al</i>. 2016).</p> <p> The new species, <i>A. minhdam</i> <b>sp. nov.</b>, is also similar to <i>A. brevicapitatus</i> (Thai, 1984), <i>A. deonganganus</i> (Thai, 1984) and <i>A. duplicoecus</i> (Thai, 1982) in having one pair of genital markings in xviii, median to the male pore. However, the new species differs from these three new species in having two pairs of spermathecal pores in 5/6/7 (vs. two pairs in 7/8/ 9 in <i>A. brevicapitatus</i>, three pairs in 5/6/7/ 8 in <i>A. deonganganus</i>, four pairs in 5/6/7/8/ 9 in <i>A. duplicoecus</i>). <i>A. minhdam</i> <b>sp. nov.</b> also differs from <i>A. deonganganus</i> and <i>A. duplicoecus</i> in having first dorsal pore in 12/13 (vs. 11/12 and 5/ 6 in <i>A. deonganganus</i> and <i>A. duplicoecus</i>, respectively) and simple intestinal caeca (vs. manicate caeca). Two species, <i>A. minhdam</i> <b>sp. nov.</b> and <i>A. brevicapitatus</i> also share the character state of simple intestinal caeca, but they are clearly distinguished by the following: <i>A. minhdam</i> <b>sp. nov.</b> has spermathecal pores in lateroventrally 5/6/7, first dorsal pore in 12/13, a pair of genital markings in the male region, chain-shaped seminal chamber; while <i>A. brevicapitatus</i> has spermathecal pores in laterally 7/8/9, first dorsal pore in 11/12, more than one pair of genital markings in the male region and clavate-shaped seminal chamber.</p>Published as part of <i>Nguyen, Tung T., Tran, Binh T. T., Lam, Dang H. & Nguyen, Anh D., 2020, Four new species of Amynthas earthworms in southeastern Vietnam (Annelida, Oligochaeta, Megascolecidae), pp. 277-290 in Zootaxa 4790 (2)</i> on pages 286-288, DOI: 10.11646/zootaxa.4790.2.5, <a href="http://zenodo.org/record/3889527">http://zenodo.org/record/3889527</a>
Rhyacobates angustus Tran & Nguyen, 2016, sp.n.
No full text<i>Rhyacobates angustus</i> sp.n. <p>(Figs. 1–10)</p> <p> <b>Material examined.</b> Holotype (apterous female) and allotype (apterous male), VIETNAM, Lao Cai Prov., Sa Pa, Ban Ho, Ban Den, Nam Pu stream (feeder stream of Muong Hoa stream), 22°15.709’N 103°58.054’E, 416 m asl., coll. Tran A.D. <i>et al.</i>, 29 May 2013, TAD1316 (ZMHU).</p> <p>Paratypes: 2 males (apterous), same locality data as holotype (ZMHU).</p> <p> <b>Description.</b> Size: apterous morph, males, length 7.7–8.1 (allotype 8.0), width 2.26–2.40 (allotype 2.40); female (holotype), length 11.3, width 2.95.</p> <p>Colour: Dorsum of body mainly black, covered with silvery pubescence, especially on thoracic pleura, metacoxa and abdomen. Dorsum of head yellow with median, longitudinal brown mark on anterior three quarters.</p> <p>Antennae mainly black, segment 4 with whitish area in distal two-fifths. Pronotum mainly black with median yellow mark (in female, width of mark broad, about one third of pronotum mark; in male, yellow mark narrow). Mesonotum black with median yellow mark from anterior fifth to posterior margin; median mark of female broader than in male. Metanotum black, in female with median yellow marking, in male without yellow mark. Abdominal terga of male chiefly black or brown. Abdominal terga of female mainly black or brown, with yellow connexiva. Venter of body covered with silvery and golden pubescence. Venter of female mainly light yellow, except mesosternum and mesopleuron black on anterior part, with sub-triangular yellow on posterior part of mesosternum. Venter of male similar to female, but triangular yellow mark on mesosternum narrower, abdominal venter dark yellow. All acetabulae, coxae and trochanters yellow. Fore femur yellow with light brown dorsal stripe on distal part. Middle and hind femora yellow-brown at basal part, dark brown or black at distal part. Fore tibia yellowbrown and basal part, dark brown at distal part; other tibiae and all tarsi dark brown or black.</p> <p>Apterous female (holotype): Body long and relatively slender. Head width 1.60, interocular width 0.65, eye length (dorsal view) 0.65. Lengths of antennal segments 1–4: 4.95: 1.39: 1.54: 1.16. Pronotum broader than long, length 0.80, width 1.71; mesonotum length 2.72, width 2.72; metanotum length 0.79. Thoracic pleura with numerous long erect bristle-like black hairs, especially on mesopleuron. All coxae with rings of dark bristle-like hairs at apical margins. Fore trochanter with long hairs on ventral side. Lengths of leg segments (femur: tibia: tarsus 1: tarsus 2): fore leg: 5.10: 4.75: 2.38: 1.10; middle leg: 13.80: 7.90: 3.30: 0.47; hind leg: 13.60: 5.69: 0.18: 0.27. Fore femur slender (width 0.51), venter of femur with row of dark bristle-like hairs and scattered soft long hairs, sub-apical part slightly constricted with small tooth-like elevation (Fig. 1). Abdomen prolonged, narrower than thorax, ventral length 5.38, gradually tapering towards apex (Figs. 7, 8). Tergum 1 posteromedially swollen, forming a semi-circular tumescence (Fig. 8). Connexival margin of segments 1–6 thickened. Abdominal segment 7 prolonged, almost completely enclosing genital segments, tergum 7 medially produced on posterior margin; sternum 7 (length 1.99) about 1.8 times as long as tergum 7 (length 1.12); connexival margin of sternum 7 about two thirds ventral length of sternum 7; posterior margin of sternum 7 broadly convex and thickened; each posterolateral corner of sternum 7 with broad, sub-triangular process (Figs. 8, 10).</p> <p>Apterous male (allotype): Head width 1.44, interocular width 0.64, eye length 0.61. Lengths of antennal segments 1–4: 4.80: 1.33: 1.48: 1.10. Pronotum broader than long, length 0.77, width 1.64; mesonotum length 2.16; metanotum length 0.68. Thoracic pleura with numerous long erect bristle-like black hairs, especially on mesopleuron. Lengths of leg segments: fore leg: 4.95: 4.35: 1.30: 0.81; middle leg: 13.50: 7.20: 3.00: 0.47; hind leg: 13.25: 4.60: 0.16: 0.23. Fore femur slender (width 0.50) with few scattered soft long hairs in venter, sub-apical part with small tooth-like elevation. Middle trochanter without spines, middle femur with many spines, those at basal fifth of femur not arranged in distinct row and those at distal four fifths arranged in distinct row (Fig. 2). Other leg structures similar to those of female. Abdomen relatively short, ventral length 3.13. Length of sternum 7: 0.31, posterior margin slightly concave. Genitalia: directed slightly downwards; venter of abdominal segment 8 length (in-situ) 0.20; proctiger laterally widened forming a rounded lobe on each side (Fig. 3); pygophore of moderate size, apical margin rounded; paramere relatively long, curved at basal third, slightly thickened at middle, apex narrow and directed inwards, setae not distinct (Figs. 4, 5); endosomal sclerites as in Fig. 6.</p> <p>Macropterous morph unknown.</p> <p> <b>Etymology.</b> The species epithet refers to the slender appearance of the female body.</p> <p> <b>Remarks.</b> <i>Rhyacobates angustus</i> <b>sp.n.</b> differs from all of its congeners in the combination of following characters: the body in dorsal view is generally slender, especially in the female; the abdomen of the female is straight (Figs. 7, 8); abdominal segment 7 of the female is prolonged and in lateral view, tapers apically (Figs. 8– 10), each posterolateral corner of sternum 7 is produced into a broad, sub-triangular process (Figs. 8, 10), the posterior margin of sternum 7 is broadly convex and thickened, without a median process (Fig. 9); the apex of the male paramere is curved and directed inwards (Fig. 5). For comparison with its congeners, see Remarks for <i>Rhyacobates zetteli</i> <b>sp.n.</b></p> <p> <b>Habitats.</b> Specimens of <i>R. angustus</i> <b>sp.n.</b> were found in a open mountain stream at an elevation of above ca. 400 m. They were found living sympatrically with <i>R. gongvo</i>, a more common species in the area (Fig. 40). After several searches and additional field trips, we could not find further specimens of this new species. Therefore, we are unable to determine whether this new species is uncommon in the area or if this was an atypical habitat and it is more common elsewhere.</p> <p> <b>Distribution.</b> Vietnam: Lao Cai.</p>Published as part of <i>Tran, A. D. & Nguyen, X. Q., 2016, Three new species of the water strider genus Rhyacobates Esaki, 1923 (Hemiptera: Gerridae) from Vietnam, pp. 501-516 in Zootaxa 4121 (5)</i> on pages 502-503, DOI: 10.11646/zootaxa.4121.5.1, <a href="http://zenodo.org/record/271684">http://zenodo.org/record/271684</a>
Amynthas longiprostaticus Nguyen & Tran & Lam & Nguyen 2020, sp. nov.
No full text<i>Amynthas longiprostaticus</i> Nguyen & Lam, sp. nov. <p>(Figure 2)</p> <p> <b>Material examined.</b> <i>Holotype:</i> 1 mature (complete body), (CTU-EW.088.h01), natural forests (11°25’31”N; 107°25’42”E), Cat Tien National Park, Tan Phu District, Dong Nai Province, 13 October 2013, coll. Le Van Nhan. <i>Paratypes:</i> 5 matures (complete body) (CTU-EW.088.p02), 4 matures (IEBR-EW.088.p02), same data as for holotype. <i>Non-types:</i> 30 matures (complete body), 1 mature (two fragments and two section slides), 4 immatures (CTU-EW.088.03) same data as for holotype; 17 matures (complete body) (IEBR-EW.088.03), same data as for holotype.</p> <p> <b>Etymology.</b> “ <i>longiprostaticus</i> ”, an adjective, to emphasize the long prostate gland.</p> <p> <b>Diagnosis.</b> Small size, length 51–71 mm, diameter 2.4–2.7 mm, segments 81–112. Prostomium epilobous. First dorsal pore in 11/12. Setal numbers: 42–57 in viii, 49–66 in xxx, 9–12 between male porophores in xviii. A single pair of spermathecal pores, lateroventral, in intersegment 5/6. One pair of genital markings present ventrally in xviii. Septa 8/9/10 absent. Intestinal caeca simple. Holandric. Testis sacs separated. Prostate glands well developed, moderately lobuled, paired in xviii–xxviii; ducts long, waved and directly attached to body wall.</p> <p> <b>Description.</b> Body cylindrical, small size; length 51–71 mm, diameter 2.4–2.7 mm, weight 0.2–0.3 gr, segments 81–112. Body light grey, ventrum paler than dorsum, clitellum darkish brown. Prostomium 1/3 epilobous. First dorsal pore in 11/12. Setae perichaetine, pre-clitellar setae stouter and sparser than post-clitellar ones; setal numbers: 42–57 in viii, 49–66 in xxx, 6–8 between male porophores in xviii; setal distance: aa>ab, zz>zy. Clitellum within xiv–xvi, without dorsal pores and setae. Female pore single, in midventral xiv.</p> <p>A single pair of spermathecal pores located lateroventrally in intersegments 5/6. Genital markings single, located ventrally in front of setal rings of x or xi. Male porophores located on setal ring of xviii; ventral distance between two male pores about 0.35X body circumference. A single conial genital marking located ventrally, next to male pore.</p> <p>Septa 5/6/7/8 thick, 8/9/10 absent, 10/11/12/13 thin. Oesophageal gizzard within viii–x. Intestinal origin at xv; caeca simple, originating at xxvii and extending anteriorly to xxii. Last hearts in xiii. Pharyngeal micronephridia developed 5/6/7. Typhlosole simple, lamelliform. Lymph glands absent.</p> <p>A pair of spermathecae in vi. Ampulla oval-shaped; ducts about 1/2 ampulla, robust, but getting thinner basally. Diverticula thin, shorter than ampulla, strongly twisted, directly attached to the middle of ampullar duct; seminal chamber small, oval or bean-shaped. Spermathecal ducts without nephridia. A large accessory gland located next to each ampullar duct, sometimes covering the duct. Additional accessory gland with duct in x or xi, in association with external genital marking.</p> <p>Holandric. Testis sacs developed, separated in x–xi. Seminal vesicles well developed in xi–xii. Ovaries well developed in 12/13. Ovisacs invisible. Prostate glands deeply lobuled, paired in xviii–xxviii; ducts long, waved and directly attached to body wall. A pair of large accessory glands in association with genital markings in xviii.</p> <p> <b>Habitat.</b> The species was found in natural forests of Cat Tien National Park, where the forests are among the best-conserved in Vietnam. It was collected in the 0–10 cm mineral soil below a thick litter layer.</p> <p> <b>Remarks.</b> The new species can be classified into the <i>Amynthas minimus</i> group characterized by one pair of spermathecal pores in 5/6 (Sims & Easton 1972). This group is known to comprise 25 species (Sims & Easton 1972; Thai 1982; Thai 1984a; Do <i>et al.</i> 1995; Pham 1995; Bantaowong <i>et al.</i> 2015), ten of which are known in northern and central Vietnam (Nguyen <i>et al.</i> 2016a).</p> <p> Within the <i>A. minimus</i> group, it is the only species with prostate glands that extend over up to 11 segments; in all other species the glands are shorter. Apart from that character, the new species is somewhat similar to <i>A. papilio</i> (Gates, 1930) and <i>A. khaohayod</i> Bantaowong & Panha, 2015 (in Bantaowong et <i>al.</i> 2015), both also in the <i>A. minimus</i> group, by presence of a pair of genital markings in xviii and simple intestinal caeca. <i>A. longiprostaticus</i> <b>sp. nov.</b> differs from <i>A. khaohayod</i> in ventral distance between the two male porophores (0.35X vs. 0.18X), presence of genital markings in the spermathecal and male regions (vs. absence of genital markings), first dorsal pore in 11/12 (vs. in 12/13), and body size (length 51–71mm, diameter 2.4–2.7mm, vs. length 230mm, diameter 6.8–7.5mm), the shape of the male genital markings (conical-shaped vs. glandular-shaped) and the shape of spermathecae (ovalshaped ampulla small, oval or bean-shaped seminal chamber vs. spatulate-shaped ampulla, roughly elliptic seminal chamber). Similarly, the new species is also differentiated from <i>A. papilio</i> in the presence of genital markings in the spermathecal region (vs. absence of genital markings), first dorsal pore in 11/12 (vs. in 5/6), and size of genital markings in the male region (same size as male porophore vs. larger than male porophore, extending beyond xviii). In both species compared, the male genital markings are much larger than the ones in the new species, and the prostates are shorter.</p> <p> The character state of exceptionally long prostate glands is, within <i>Amynthas</i>, shared by <i>A. osmastoni</i> (Michaelsen, 1907) (xv–xxiv) and <i>A. taiwumontis</i> Shen, Chang, Li, Chih & Chen, 2013 (xvi–xxiv). However, <i>A. osmastoni</i> is longer than 250 mm and has three pairs of spermathecal pores in 6/7/8/9, first dorsal pore in 12/13, no genital markings in the male region. <i>A. taiwumontis</i> is about 130–150 mm long, has four pairs of spermathecal pores in 5/6/7/8/9, first dorsal pore in 11/12, and no markings in the male region. The new species differs from these two species in one pair of spermathecal pores in 5/6 and presence of genital markings in the male region, and has smaller size (55–71 mm long).</p> <p> Regarding the morphology of the male region, the new species is fairly similar to <i>A. brevicapitatus</i> (Thai, 1984), <i>A. deonganganus</i> (Thai, 1984), <i>A. corticis</i> (Kinberg, 1867) and to <i>A. carnosus</i> (Goto & Hatai, 1899), species with one pair of genital markings in immediate vicinity to the male pores. These three species differ from the new one in three or four pairs of spermathecae, connected testis sacs, and in a more anterior location of genital markings in the spermathecal region (when present), among other characters.</p>Published as part of <i>Nguyen, Tung T., Tran, Binh T. T., Lam, Dang H. & Nguyen, Anh D., 2020, Four new species of Amynthas earthworms in southeastern Vietnam (Annelida, Oligochaeta, Megascolecidae), pp. 277-290 in Zootaxa 4790 (2)</i> on pages 279-281, DOI: 10.11646/zootaxa.4790.2.5, <a href="http://zenodo.org/record/3889527">http://zenodo.org/record/3889527</a>
Drawida beddardi Rosa 1890
No full text<i>Drawida beddardi</i> (Rosa, 1890) <p> <i>Moniligaster beddardii</i> Rosa, 1890a: 379, fig. 12.</p> <p> <i>Drawida beddardii—</i> Gates 1972: 246; Thai 1983: 121; Tran 1985: 57; Nguyen 1994: 73; Le 1995a: 75; Pham 1995c: 108; Thai 2000a: 311; Huynh & Nguyen 2004b: 70; Huynh 2005a: 149; Tran & Ngo 2008: 133; Nguyen V.T. & Tran 2008: 185; Tran & Tran 2010: 8; Pham 2010: 65; Nguyen & Nguyen 2010: 123.</p> <p> <i>Drawida barwelli—</i> Blakemore 2008; Nguyen <i>et al.</i> 2012: 146; Nguyen 2013: 98; Nguyen 2014: 113.</p> <p> <b>Type locality.</b> Philippines <b>(</b> Manila).</p> <p> <b>Type material.</b> NHM London, U.K.</p> <p> <b>Records from Vietnam. Bac Giang</b> (Hiep Hoa; Luc Ngan; Son Dong: Khe Ro); <b>Vinh Phuc</b> (Me Linh; Tam Dao); <b>Hanoi</b> (Soc Son); <b>Quang Binh</b> (Quang Trach); <b>Da Nang</b> (Hoa Vang; Ba Na NP); <b>Quang Nam</b> (Hiep Duc); <b>Gia Lai</b> (Buon Luoi); <b>Phu Yen</b> (Tuy Hoa); <b>Binh Dinh</b>; <b>Hau Giang</b> (Phung Hiep: Tan Binh, Tan Phuoc Hung, Long Thanh); <b>Ben Tre</b> (Cho Lach); <b>Vinh Long</b> (Vinh Long city, Vung Liem, Long Ho); <b>Tien Giang</b> (Cai Be), <b>Can Tho</b>; <b>An Giang</b> (Cho Moi, Tinh Bien); <b>Kien Giang</b> (Kien Luong, Ha Tien) (Tran 1985; Nguyen 1994; Le 1995a; Pham 1995c, 2010; Huynh & Nguyen 2004b; Huynh 2005a; Tran & Ngo 2008; Nguyen V.T. & Tran 2008; Tran & Tran 2010; Nguyen 2014).</p> <p> <b>Distribution.</b> Myanmar (Gates 1972).</p> <p> <b>Vietnamese name.</b> Giun drawid beddard.</p> <p> <b>Remarks.</b> The species was synonymized with <i>Drawida barwelli</i> (Beddard, 1886) (Blakemore 2002), but it is a valid species distinguishable specifically from <i>D. barwelli</i> as has been shown by Shen <i>et al.</i> (2015: 434–435). Material from Vietnam shows the untwisted and short sperm ducts characteristic of <i>D. beddardi</i> (Nguyen 2013).</p>Published as part of <i>Nguyen, Tung T., Nguyen, Anh D., Tran, Binh T. T. & Blakemore, Robert J., 2016, A comprehensive checklist of earthworm species and subspecies from Vietnam (Annelida: Clitellata: Oligochaeta: Almidae, Eudrilidae, Glossoscolecidae, Lumbricidae, Megascolecidae, Moniligastridae, Ocnerodrilidae, Octochaetidae), pp. 1-92 in Zootaxa 4140 (1)</i> on page 77, DOI: 10.11646/zootaxa.4140.1.1, <a href="http://zenodo.org/record/256507">http://zenodo.org/record/256507</a>
Glyphidrilus papillatus Rosa 1890
No full text<i>Glyphidrilus papillatus</i> (Rosa, 1890) <p> <i>Bilimba papillata</i> Rosa, 1890a: 386, plate 12, fig. 1; Beddard 1895a: 687</p> <p> <i>Glyphidrilus papillatus—</i> Michaelsen 1900: 459; Chen 1938: 426; Gates 1972: 236; Thai 1983: 121; Tran 1985: 62; Do 1994: 115; Nguyen 1994: 72; Thai 2000a: 310; Huynh & Nguyen 2004b: 70; Huynh 2005a: 147; Nguyen V.T. & Tran 2008: 185; Blakemore 2008b; Tran & Tran 2010: 8; Chanabun <i>et al.</i> 2013: 17; Nguyen & Nguyen 2010: 123; Nguyen & Huynh 2011: 1018; Nguyen <i>et al.</i> 2012: 146; Nguyen 2013: 97; Nguyen 2014: 113.</p> <p> <b>Type locality.</b> Myanmar (Cobapo).</p> <p> <b>Type material.</b> Genoa Museum (44025), Italy.</p> <p> <b>Examined material.</b> 5 C, 15 A (<b>CTU-EW.030.01</b>) Au Isl., Cai Rang, Can Tho, 13/9/2011, coll. Nguyen Thanh Tung.</p> <p> <b>Records from Vietnam.</b> Widely distributed in the Red River Delta, and <b>Bac Giang</b> (Hiep Hoa; Luc Ngan); <b>Hoa Binh</b> (Mai Chau); <b>Quang Tri</b> (Gio Linh; Quang Tri); <b>Binh Dinh</b> (Quy Nhon); <b>Dak Lak</b> (Buon Don); <b>Khanh Hoa</b> (Van Ninh); <b>Binh Thuan</b> (Tanh Linh); <b>Tien Giang</b> (Tan Hoa Dong; Cai Lay); <b>Tra Vinh</b> (Duyen Hai); <b>Ca Mau</b> (Ca Mau City); <b>Can Tho</b> (Cai Rang); <b>An Giang</b> (Tinh Bien; Tri Ton) (Tran 1985; Do 1994; Nguyen 1994; Huynh & Nguyen 2004b; Huynh 2005a; Nguyen V.T. & Tran 2008; Tran & Tran 2010; Nguyen 2014).</p> <p> <b>Distribution.</b> Myanmar (Gates 1972; Chanabun <i>et al.</i> 2013).</p> <p> <b>Vietnamese name.</b> Giun khảm nhú.</p> <p> <b>Remark.</b> The species is common in rice fields in the Red River and can also be found near streams in northwestern Vietnam (Do 1994).</p>Published as part of <i>Nguyen, Tung T., Nguyen, Anh D., Tran, Binh T. T. & Blakemore, Robert J., 2016, A comprehensive checklist of earthworm species and subspecies from Vietnam (Annelida: Clitellata: Oligochaeta: Almidae, Eudrilidae, Glossoscolecidae, Lumbricidae, Megascolecidae, Moniligastridae, Ocnerodrilidae, Octochaetidae), pp. 1-92 in Zootaxa 4140 (1)</i> on page 6, DOI: 10.11646/zootaxa.4140.1.1, <a href="http://zenodo.org/record/256507">http://zenodo.org/record/256507</a>
Metaphire dorsomultitheca Nguyen & Nguyen 2015
No full text<i>Metaphire dorsomultitheca</i> Nguyen & Nguyen, 2015 <p>(Figure 11, Table 3)</p> <p> <i>Metaphire dorsomultitheca</i> Nguyen <i>et al.</i> 2015: 465, fig. 2; Nguyen <i>et al.</i> 2016: 56, 2021c: 105.</p> <p> <i>Pheretima</i> sp. 15 <i>–</i> Nguyen 2013: 87, 2014: 112.</p> <p> <b>Type locality.</b> Vietnam (An Giang: Sam mountain) (Nguyen <i>et al.</i> 2015)</p> <p> <b>Type material.</b> Laboratory of Zoology, Department of Biology, Can Tho University (EW.025.h01, EW.025. p02), Vietnam.</p> <p> <b>Diagnosis.</b> Body cylindrical, small-medium size, length 76–111 mm, diameter 2.9–3.6 mm, segments 88–120. Prostomium epilobous. First dorsal pore at 12/13. Spermathecal pores numerous in dorsal intersegments 7/8/9, polythecate. Male pores located inside copulatory pouches in xviii. Genital markings invisible outside, but two pairs opened on the front and back walls of the copulatory pouches, respectively. Septum 10/11 present only ventrally. Intestinal caeca simple. Holandric, testis sacs separated. Accessory glands coelomic, strongly covered by muscularwalled bursae in 17/18 and 18/19.</p> <p> <b>Habitat.</b> The species was usually found in the depth of 0–5 cm in the soils under small trees and shrubs on mountain peaks (Nguyen <i>et al.</i> 2015).</p> <p> <b>Distribution.</b> Known only in Vietnam (An Giang: Sam, Cam, and Co To mountains) (Nguyen <i>et al.</i> 2015) (Fig. 3).</p> <p> <b>Remarks.</b> There are no significant differences between individuals except the number of spermathecae. The K2P intraspecific genetic distance was 0.7%±0.3%. The genetic distance between <i>M. dorsomultitheca</i> and other <i>Metaphire</i> species was from 14.9%±1.6% (with <i>M. bahli</i> (I)) to 21.8%±2.0 (with <i>M. grandiverticulata</i>) (Table 4).</p>Published as part of <i>Nguyen, Tung T., Lam, Dang H. & Nguyen, Anh D., 2023, A revision of the Metaphire peguana species-group (Oligochaeta: Megascolecidae) from Vietnam, pp. 113-135 in Zootaxa 5255 (1)</i> on pages 129-130, DOI: 10.11646/zootaxa.5255.1.15, <a href="http://zenodo.org/record/7744555">http://zenodo.org/record/7744555</a>
Annamocalamus kontumensis H. N. Nguyen, N., spec. nova
No full text<i>Annamocalamus kontumensis</i> H. N. Nguyen, N.-H. Xia & V. T. Tran, spec. nova (Fig. 1, 2). <p> <b>Typus: Vietnam, Kon Tum Province:</b> Ngoc Linh Mountain, Ngoc Linh village, c. 1200 m, 23.II.1995, fl. & fr., <i>Averyanov & al. VH020</i> (holo-: P [P00451097]).</p> <p> <i>Rhizomatis sympodialibus, rhizomemate brevi collo, culmi erectus, vaginis auriculis conspicuis, dorso dense adpresseque atro-strigosis, laminis triangularis, erectus, basis producens, deciduus, inflorrescentilis iterauctantilus, pseudospiculae ad convivia in foliis nodorum distal ramis florentibus, glumis 3, flosculis hermaphroditus 1, lodicula 3, stylus longis, stigmatibus 3, brevis, frutus carnosus.</i></p> <p> <i>Densely tufted</i> with rhizomes short necked, pachymorph. <i>Culms</i> erect, 4-8 m tall; internodes 60-80 cm long and 3- 4.5 cm in diameter, when young densely covered with appressed white hairs, becoming rough when old due to silica; culms walls 2-3 mm thick; nodes slight swollen. <i>Branches</i> a cluster of slender subequal branches, none dominant, these branching further. <i>Culm</i> sheaths persistent, greenish brown, rough, when young covered with densely appressed purplebrown bristle on the abaxial surface; 20-25 cm long and 12- 16 cm wide at the base; apex 8-10 cm wide, margins dense purple-brown hairs; blade tardily deciduous, triangular, erected, base black-brown when young and produced upwards into rounded, 10-15 × 6-8 cm, abaxial dense white powder, margins dense cilia, adaxilly dense hairs at the base; one auricle standing, 2-2.5 × 0.4-0.6 cm, other one margin curved downwards, 1.5-2 × 0.4-0.5 cm, adaxilly dense purple-brown bristle, ca. 0.9 cm long; ligule ca. 0.3 cm long with dense paleciliate, ca. 0.3 cm long. <i>Leaf</i> blades oblong-obovate, base broadly cuneate, 30-34 × 3.5-4 cm, veins in 13-14 pairs, margins and adaxilly surface dense white cilia; leaf sheaths dense white cilia, auricles ca. 0.3 ×c a. 0.2 cm with slender bristles, 5-8 mm long; ligule low rim, ca. 1mm; petiole 4-5 × 2-3 mm. <i>Pseudospikelets</i> in groups at the distal nodes on leafless flowering branches, ca. 8 mm long; 1 fertile floret, glumes 3, upper most obovate, 4-5 × 2-3 mm, apex acute, mucronate, mucro ca. 1 mm long, veins in 5, margins short white-hispid, abaxial surface sparse white cilia; lemma oblong-lanceolate, 5-6 × 3-4 mm, veins in 9-11, apex acute, mucronate, mucro 1.5- 2 mm long, convolute and covering most of the palea, margins dense white cilia; palea oblong-lanceolate, 7-8 × 2.5-3 mm, veins in 9-11, strongly involutes, sparse white cilia, apex bifid ca. 1 mm long, with a narrow groove on the back (at the base rachilla is joined); lodicules 3, obovate-shaped, 0.7-1 × ca. 0.5 mm margins long cilia; stamens 6, filaments free, 4-5 × 0.4-0.5 mm; style long; stigmas 3, plumose; ovary glabrous, short stalked. <i>Pericarp</i> thick, apple shaped, fleshy, globose, top horizontal or slightly truncated in the middle, 12-15 × 12- 15 mm.</p> <p> <i>Distribution and habitat.</i> – The new species grows in the degraded natural forest in valleys and mountain gorges, but is common along river or valleys, between 500 to 1200 m in the western highland of Vietnam.</p> <p> <i>Phenology. – Annamocalamus kontumensis</i> flowered in December 1946 <i>(Poilane 35696)</i> and January 1995 <i>(Averyanov & al.VH020).</i> New shoots seems to develop between June to August.</p> <p> <i>Local uses.</i> – This species is of considerable importance to the local people. Its culms are used for making handicrafts, household tools.</p> <p> <i>Etymology.</i> – The specific epithet refers to the type locality, Kon Tum Province, Vietnam.</p> <p> <i>Paratypi.</i> – <b>VIETNAM. Kon Tum Province:</b> Dak To District, Lo Xo Pass, 15°13’338’’N 107°44’052’’E, 1071 m, 28.VIII.2005, st., <i>H. N. Nguyen</i>, V. T. <i>Tran</i> 2820050540 (FSIV); Dak To and Dak Blan District, 1.IX.1946, fl. & fr., <i>Poilane 35696</i> (P).</p>Published as part of <i>Tran, Van Tien, Nguyen, Hoang Nghia & Xia, Nian-He, 2013, Annamocalamus H. N. Nguyen, N. - H. Xia & V. T. Tran, a new genus of bamboo (Poaceae) from Vietnam, pp. 159-165 in Candollea 68 (1)</i> on page 161, DOI: 10.15553/c2013v681a23, <a href="http://zenodo.org/record/5747054">http://zenodo.org/record/5747054</a>
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