6,896 research outputs found
Micronecta (Micronecta) vietnamica Tran 2021, sp. nov.
No full textMicronecta (Micronecta) vietnamica sp. nov. urn:lsid:zoobank.org:act: FFBEFA51-B20A-43BC-A91E-7EFA22350817 Figs 5J–L, 12 Diagnosis Body length 1.5–1.8. Pronotum longer than median head length. Dorsum colour pale yellowish. Hemelytron punctate, with large dark markings, sometimes indistinct (Fig. 5J–L). Males: fore femur with two spines on proximal third ventrally, one spine in distal third dorsally, two spines distally, one pair of long setae dorsodistally; fore tibia without spine; palar claw petaloid, apex truncate (Fig. 12A). Median lobe of sternite VII with angular apex and four long setae (Fig. 12C). Free lobe long, parallel sided, mesial angle broadly rounded, lateral angle nearly square with ten long setae (Fig. 12D). Left paramere: shaft broad, flat, proximal part curved, distal part nearly straight, apex rounded; basal lobe tongue-shaped (Fig. 12F). Right paramere: shaft slender and parallel sided proximally, distal third curved, apex narrowly rounded; pars stridens processus with ca 25 ridges (Fig. 12E). Etymology This new species is named after the country of the type locality. Material examined Holotype VIETNAM • ♂; Hanoi, Ba Vi N.P., Khoang Xanh, Tien stream; 95 m a.s.l.; 17 Dec. 2011; Q.H. Ngo and A.D. Tran leg.; TAD1127; ZVNU. Paratypes VIETNAM – Hanoi • 16 ♂♂, 9 ♀♀; same locality data as for holotype; ZVNU • 3 ♂♂, 3 ♀♀; same locality data as for holotype; NHMW • 1 ♀; Ba Vi N.P., Khanh Thuong, Mit stream, forest area; 80 m a.s.l.; 17 Dec. 2011; Q.H. Ngo et al. leg.; TAD1124; ZVNU • 1 ♂, 1 ♀; Ba Vi N.P., Minh Quang, Cai stream; 65 m a.s.l.; 17 Dec. 2011; Q.H Ngo. et al. leg.; TAD1126; ZVNU • 2 ♂♂, 5 ♀♀; Ba Vi N.P., Minh Quang, stream nearby Tan Vien Pagoda; 340 m a.s.l.; 26 Apr. 2012; Q.H. Ngo et al. leg.; TAD1206; ZVNU • 1 ♀; Ba Vi N.P., Minh Quang, Cai stream; 65 m a.s.l.; 26 Apr. 2012; Q.H. Ngo et al. leg.; TAD1207; ZVNU • 3 ♂♂, 3 ♀♀; Ba Vi N.P., Khoang Xanh, Tien stream; 95 m a.s.l.; 27 Apr. 2012; Q.H. Ngo and A.D. Tran leg.; TAD1208; ZRC • 5 ♂♂; Ba Vi N.P., Yen Bai, Mo stream; ca 114 m a.s.l.; 28 Apr. 2012; Q.H Ngo and A.D. Tran leg.; TAD1211; ZVNU. – Lao Cai Prov. • 8 ♂♂, 1 ♀; Sa Pa, Seo Trung Ho stream; 22 Oct. 2012; N.H. Dinh et al. leg.; DNH12.05; ZVNU • 1 ♀; Sa Pa, Nam Sai, Seo Nam Sai stream 1; 24 Oct. 2012; N.H. Dinh et al. leg.; DNH12.09; ZVNU • 1 ♂; Sa Pa, Nam Sai, Seo Nam Sai stream 2; 24 Oct. 2012; N.H. Dinh et al. leg.; DNH12.10; ZVNU • 1 ♂; Sa Pa, Thanh Phu, Nam Cang stream; 26 Oct. 2013; A.D. Tran et al. leg.; TAD1359; ZVNU. – Cao Bang Prov. • 1 ♂; Ha Quang, road 203, Truong Ha commune, stream near Kim Dong tomb; 17 Dec. 2013; A.D. Tran et al. leg.; TAD1371; ZVNU • 6 ♀♀; Phia Oac-Phia Den N.P., road to Quang Thanh commune, Na Leng stream; 29 Oct. 2020; V.V. Nguyen et al. leg.; PO20-06; ZVNU • 1 ♂, 4 ♀♀; Phia Oac-Phia Den N.P., road to Quang Thanh commune, Khue Lan stream; 29 Oct. 2020; V.V. Nguyen et al. leg.; PO20-07; ZVNU. – Lang Son Prov. • 1 ♂, 3 ♀♀; Loc Binh, Mau Son, Khuoi Tang stream, 590 m a.s.l.; 6 Nov. 2020; A.D. Tran et al. leg.; TAD20-34; ZVNU • 12 ♂♂, 9 ♀♀; Loc Binh, Mau Son, Long Dau stream; 300 m a.s.l.; 7 Nov. 2020; A.D. Tran et al. leg.; TAD20-36; ZVNU. – Phu Tho Prov. • 2 ♂♂, 2 ♀♀; Xuan Son N.P., Ban Coi, Coi stream, site 2, near bridge; 7 Jun. 2013; A.D. Tran et al. leg.; TAD1338; ZVNU • 1 ♂, 2 ♀♀; Xuan Son N.P., Kim Thuong, Xoan stream; 28 Aug. 2013; A.D. Tran et al. leg.; TAD1343; ZVNU • 2 ♀♀; Xuan Son N.P., Ban Coi, Coi stream, site 1, water from underground; 29 Aug. 2013; A.D. Tran et al. leg.; TAD1349; ZVNU • 1 ♀; Xuan Son N.P., Ban Coi, Coi stream, site 3, by the road to Tan Son; 29 Aug. 2013; A.D. Tran et al. leg.; TAD1351; ZVNU • 1 ♂, 2 ♀♀; Xuan Son N.P., stream flows through the road, near Lang Village; 21°5.662′ N, 104°56.236′ E; 26 Aug. 2014; T.S. Nguyen leg.; TS1402; ZVNU • 1 ♂, 1 ♀; Xuan Son N.P., stream flows through the road, near Lang Village; 21°5.662′ N, 104°56.236′ E; 21 May 2015; T.S. Nguyen leg.; TS1502; ZVNU • 2 ♂♂, 2 ♀♀; Xuan Son N.P., stream flows through the road, near Lang Village; 21°5.662′ N, 104°56.236′ E; 24 Aug. 2015; T.S. Nguyen leg.; TS8502; ZVNU • 4 ♂♂, 5 ♀♀; Xuan Son N.P., Than stream, site 2; 21°11.257′ N, 104°52.382′ E; 28 Aug. 2014; T.S. Nguyen leg.; TS1415; ZVNU • 3 ♂♂, 1 ♀; Xuan Son N.P., Than stream, site 3; 21°11.393′ N, 104°52.625′ E; 28 Aug. 2014; T.S. Nguyen leg.; TS1416; ZVNU • 1 ♂; Xoan stream, near Lang Village; 21°5.453′ N, 104°56.122′ E; 21 May 2015; T.S. Nguyen leg.; TS1501; ZVNU • 4 ♂♂, 1 ♀; Xuan Son N.P., Xoan stream, near its confluence with Ha Bang stream; 21°5.303′ N, 104°57.559′ E; 22 May 2015; T.S. Nguyen leg.; TS1507; ZVNU • 3 ♀♀; Xuan Son N.P., Xoan stream, near its confluence with Ha Bang stream; 21°5.303′ N, 104°57.559′ E; T.S. Nguyen leg.; 6 May 2016, TS1607; ZVNU • 1 ♂, 8 ♀♀; Xuan Son N.P., Ha Bang stream, Ha Bang village area; 21°04.343′ N, 104°59.182′ E; 25 May 2015; T.S. Nguyen leg.; TS1510; ZVNU • 2 ♂♂; Xuan Son N.P., Lap stream, near Ngoc waterfall; 21°08.311′ N, 104°56.202′ E; 23 May 2015; T.S. Nguyen leg.; TS1511; ZVNU • 2 ♂♂, 1 ♀; Xuan Son N.P., Than stream, site 1, water flowing from the cave; 21°11.096′ N, 104°52.587′ E; 24 May 2015; T.S. Nguyen leg.; TS1514; ZVNU • 4 ♂♂; Xuan Son N.P., Than stream, site 2; 21°11.257′ N, 104°52.382′ E; 24 Aug. 2015; T.S. Nguyen leg.; TS1515; ZVNU • 1 ♂, 3 ♀♀; Xoan stream, near Lang Village; 21°5.453′ N, 104°56.122′ E; 24 Aug. 2015; T.S. Nguyen leg.; TS8501; ZVNU • 3 ♂♂, 1 ♀, Xuan Son N.P., Than stream, site 3; 21°11.393′ N, 104°52.625′ E; 27 Aug. 2015; T.S. Nguyen leg.; TS8516; ZVNU • 2 ♀♀; Xuan Son N.P., Xoan stream, Xoan village area; 21°05.270′ N, 104°57.591′ E; 23 May 2015; T.S. Nguyen leg.; TS1509; ZVNU • 2 ♀♀; Xuan Son N.P., Xoan stream, Xoan village area; 21°05.270′ N, 104°57.591′ E; 26 Aug. 2015; T.S. Nguyen leg.; TS8509; ZVNU • 4 ♂♂, 1 ♀; Xuan Son N.P., Xoan stream, Xoan village area; 21°05.270′ N, 104°57.591′ E; 7 May 2016; T.S. Nguyen leg.; TS1609; ZVNU • 3 ♂♂, 1 ♀; Xuan Son N.P., Coi stream, site 1, by the road to Tan Son; 21°09.204′ N, 104°56.927′ E; 27 Aug. 2015; T.S. Nguyen leg.; TS8517; ZVNU • 2 ♂♂, 2 ♀♀; Xuan Son N.P., Coi stream, site 1, by the road to Tan Son; 21°09.204′ N, 104°56.927′ E; 8 May 2016; T.S. Nguyen leg.; TS1617; ZVNU • 1 ♀; Xuan Son N.P., Coi stream, site 2, near concrete bridge; 21°09.130′ N, 104°56.761′ E; 29 Aug. 2014; T.S. Nguyen leg.; TS1418; ZVNU • 6 ♂♂, 2 ♀♀; Xuan Son N.P., Coi stream, site 2, near concrete bridge; 21°09.130′ N, 104°56.761′ E; 24 May 2015; T.S. Nguyen leg.; TS1518; ZVNU • 3 ♂♂, 7 ♀♀; Xuan Son N.P., Coi stream, site 3, in front of the cave; 21°01.190′ N, 104°56.624′ E; 24 May 2015; T.S. Nguyen leg.; TS1519; ZVNU • 15 ♂♂, 12 ♀♀; Xuan Son N.P., Coi stream, site 3, in front of the cave; 21°01.190′ N, 104°56.624′ E; 8 May 2016; T.S. Nguyen leg.; TS1619; ZVNU • 4 ♂♂, 3 ♀♀; Xuan Son N.P., Tan Ong stream, 2 km from Chin Tang stream; 21°04.201′ N, 104°56.906′ E; 22 May 2015; T.S. Nguyen leg.; TS1504; ZVNU • 2 ♂♂, 2 ♀♀; Xuan Son N.P., Tan Ong stream, 2 km from Chin Tang stream; 21°04.201′ N, 104°56.906′ E; 25 May 2015; T.S. Nguyen leg.; TS8504; ZVNU • 1 ♀; Xuan Son N.P., Tan Ong stream, 4 km from Chin Tang stream; 21°04.288′ N, 104°57.271′ E; 26 Aug. 2014; T.S. Nguyen leg.; TS1405; ZVNU • 2 ♀♀; Xuan Son N.P., Tan Ong stream, 4 km from Chin Tang stream; 21°04.288′ N, 104°57.271′ E; 22 May 2015; T.S. Nguyen leg.; TS1505; ZVNU • 2 ♀♀; Xuan Son N.P., Ha Bang stream, Ha Bang village area; 21°04.343′ N, 104°59.182′ E; 27 Aug. 2014; T.S. Nguyen leg.; TS1410; ZVNU • 1 ♂, 2 ♀♀; Xuan Son N.P., Ha Bang stream, Ha Bang village area; 21°04.343′ N, 104°59.182′ E; 25 May 2015; T.S. Nguyen leg.; TS1510; ZVNU • 1 ♂, 1 ♀; Xuan Son N.P., Ha Bang stream, Ha Bang village area; 21°04.343′ N, 104°59.182′ E; 7 May 2016; T.S. Nguyen leg.; TS1610; ZVNU • 1 ♂; Xuan Son N.P., Chieng stream, near ranger station; 21°06.072′ N, 104°59.433′ E; 27 Aug. 2014; T.S. Nguyen leg.; TS1408; ZVNU • 1 ♂; Xuan Son N.P., Lap stream, near the foot of Ngoc waterfall; 21°08.311′ N, 104°56.202′ E; 27 Aug. 2014; T.S. Nguyen leg.; TS1411; ZVNU • 4 ♀♀; Xuan Son N.P., Lap stream, near the foot of Ngoc waterfall; 21°08.311′ N, 104°56.202′ E; 26 Aug. 2015; T.S. Nguyen leg.; TS8511; ZVNU • 2 ♂♂, 2 ♀♀; Xuan Son N.P., Lap stream, near the foot of Ngoc waterfall; 21°08.311′ N, 104°56.202′ E; 7 May 2016; T.S. Nguyen leg.; TS1611; ZVNU • 2 ♂♂, 2 ♀♀; Xuan Son N.P., Lap stream, the first concrete bridge from Ngoc waterfall; 5 Jun. 2013; A.D. Tran et al. leg.; TAD1332; ZVNU • 2 ♀♀; Xuan Son N.P., Lap stream, the first concrete bridge from Ngoc waterfall; 21°8.398′ N, 104°56.303′ E; 26 Aug. 2015; T.S. Nguyen leg.; TS8512; ZVNU • 17 ♂♂, 14 ♀♀; Xuan Son N.P., Lap stream, the first concrete bridge from Ngoc waterfall; 21°8.398′ N, 104°56.303′ E; 7 May 2016; T.S. Nguyen leg.; TS1612; ZVNU • 3 ♂♂, 11 ♀♀; Xuan Son N.P., Lap stream, the second concrete bridge from Ngoc waterfall; 5 Jun. 2013; A.D. Tran et al. leg.; TAD1333; ZVNU • 13 ♂♂, 18 ♀♀; Xuan Son N.P., Lap stream, the second concrete bridge from Ngoc waterfall; 21°08.322′ N, 104°56.256′ E; 23 May 2015; T.S. Nguyen leg.; TS1513; ZVNU • 14 ♂♂, 22 ♀♀; Xuan Son N.P., Lap stream, the second concrete bridge from Ngoc waterfall; 21°08.322′ N, 104°56.256′ E; 7 May 2016; T.S. Nguyen leg.; TS1613; ZVNU. – Hoa Binh Prov. • 4 ♂♂, 5 ♀♀; Lac Son, Tu Do commune, Mu stream (upstream of Mu waterfall); 27 Aug. 2020; A.D. Tran et al. leg.; TAD20-01; ZVNU. – Thanh Hoa Prov. • 1 ♀; Ben En N.P., Nhu Xuan, Tan Binh, stream 300 m behind Xuan Ly ranger station; 26 Feb. 2013; T.D. Pham leg.; BE1302; ZVNU. – Nghe An Prov. • 1 ♀; Pu Mat N.P., Khe Kem waterfall and stream, site #1; 20 Dec. 2012; Q.H. Ngo leg.; NQH1203; ZVNU • 11 ♂♂, 4 ♀♀; Pu Mat N.P., Khe Kem stream, site #2, ca 300 m downstream from waterfall; 20 Dec. 2012; Q.H. Ngo leg.; NQH1204; ZVNU • 8 ♂♂, 1 ♀; Pu Mat N.P., Khe Kem stream, site #3; 20 Dec. 2012; Q.H. Ngo leg.; NQH1205; ZVNU • 2 ♂♂; Pu Mat N.P., Khe Kem stream, site #4; 20 Dec. 2012; Q.H. Ngo leg.; NQH1206; ZVNU • 5 ♂♂, 2 ♀♀; Con Cuong, Tung Huong Village, Khe Thoi stream; 21 Dec. 2012; Q.H. Ngo leg.; NQH1210; ZVNU • 2 ♀♀; Pu Mat N.P., Khe Kem waterfall and stream, site #1; 18 Apr. 2013; A.D. Tran et al. leg.; TAD1310; ZVNU • 3 ♀♀; Pu Mat N.P., Khe Kem waterfall and stream, site #2, ca 300 m downstream from waterfall; 18 Apr. 2013; A.D. Tran et al. leg.; TAD1311; ZVNU • 1 ♂, 1 ♀; Pu Mat N.P., Khe Kem stream, site #3, near visitor’s hut; 18 Apr. 2013; A.D. Tran et al. leg.; TAD1312; ZVNU • 1 ♂; Pu Mat N.P., Khe Kem stream, site #6; 18 Apr. 2013; A.D. Tran et al. leg.; TAD1315; ZVNU • 1 ♂, 1 ♀; Pu Mat N.P., Kem waterfall and stream; 30 Aug. 2020; A.D. Tran, P.L. Le and T.C. Nguyen leg.; TAD20-06; ZVNU • 2 ♀♀; Pu Mat N.P., Kem stream, site 3, ca 1.5 km from waterfall; 30 Aug. 2020; A.D. Tran et al. leg.; TAD20-08; ZVNU • 1 ♂; Pu Mat N.P., Chat stream, tributary of Choang stream; 1 Sep. 2020; A.D. Tran et al. leg.; TAD20-14; ZVNU • 1 ♂, 2 ♀♀; Pu Mat N.P., Da Mai stream, tributary of Choang stream, near ranger post; 1 Sep. 2020; A.D. Tran and T.C. Nguyen leg.; TAD20-16; ZVNU. – Ha Tinh Prov. • 1 ♀; Vu Quang N.P., a small branch of Song Con River; 23 Apr. 2003; A.D. Tran leg.; TAD0303; ZRC. Description COLOUR (Fig. 5J–L). Frons and vertex pale yellowish, vertex with a longitudinal brown mark medially, eyes blackish brown. Pronotum yellowish brown with a pair of dark brown transverse marks. Hemelytron hyaline light yellowish. In most of specimens: clavus bordered with dark brown mark, becoming thicker on anterior margin; corium with two dark brown marks on middle part; a dark brown mark extending towards membrane; membrane generally transculent, brownish. Some specimens collected from Cao Bang and Phu Tho with variations in colour pattern of hemelytra. In some specimens from Phu Tho Prov. (Fig. 5K, at localities TS1510, TS1511): markings on hemelytra distinct but brighter overall, marks in middle part more fused into one large mark, and a dark mark (resembling inverted A-shape) just before membrane; membrane light coloured. In specimens from Cao Bang Prov. (Fig. 5L): markings on hemelytra indistinct, less contrasting against background colour of hemelytra; membrane light coloured. Venter of thorax and abdomen brownish yellow and legs pale yellow. Structural characteristics MACROPTEROUS MALES. Body length 1.63–2.16 (holotype: 1.83), width 0.94–1.13 (holotype: 1.08). Head width 0.68–0.77 (holotype: 0.74). Pronotum short, about five times as wide as long (0.79: 0.16). Pronotum slightly wider than head. Synthlipsis about 1.5 times wider than posterior width of an eye (0.35: 0.24). Lengths of leg segments: fore leg: femur 0.3, tibia 0.18, pala 0.14; middle leg: femur 0.74, tibia 0.27, tarsus 0.36, claw 0.2; hind leg: femur 0.58, tibia 0.4, tarsus I 0.41, tarsus II 0.16, claw 0.04. Prestrigilar flap on segment V with a short apex (Fig. 12B). Strigil present. Median lobe of sternite VII, free lobe, and parameres as in diagnosis. MACROPTEROUS FEMALES. Body length 1.63–2.09, width 0.94–1.19. Head width 0.71–0.77. Pronotum about five times as wide as long (0.76: 0.16). Pronotum slightly wider than head. Synthlipsis about 1.3 times as wide as posterior width of an eye (0.33: 0.24). Lengths of leg segments: fore leg: femur 0.23, tibiotarsus 0.27; middle leg: femur 0.67, tibia 0.21, tarsus 0.36, claw 0.2; hind leg: femur 0.6, tibia 0.36, tarsus I 0.41, tarsus II 0.18, claw 0.09. Distribution Northern Vietnam: Lao Cai, Cao Bang, Phu Tho, Ha Noi, Thanh Hoa, Nghe An, Ha Tinh. Remarks This new species may be related to the group of M. anatolica, M. arcuata sp. nov., M. acuminata sp. nov., and M. fulvopicta sp. nov., as it also has the elongate sub-rectangular free lobe, the broad and flat shaft of the left paramere, and the slender, distally curved shaft of the right paramere. It has strong dark pattern on the dorsum like M. fulvopicta sp. nov., but its dark marks are usually slenderer, and the patterns on the corium are different, sometimes indistinct (see Fig. 5L). The left paramere of M. vietnamica sp. nov. has a rather straight shaft and a trapezoid basal lobe, while in the four species mentioned above, the left paramere has a more sinuate lateral margin of the shaft and the basal lobe is either subovate or subrectangular. The right paramere of M. vietnamica sp. nov. is slenderer and strongly curved at the distal third, while that of the four species above is bent or curved at the distal fourth. In addition, the palar claw of M. vietnamica sp. nov. is much broader and has a different outline compared to that of M. anatolica, M. arcuata sp. nov., M. acuminata sp. nov., and M. fulvopicta sp. nov.Published as part of Ha, Tuyet Ngan & Tran, Anh Duc, 2021, Taxonomy of Micronectidae (Heteroptera: Nepomorpha) from Vietnam, with descriptions of 11 new species, pp. 1-82 in European Journal of Taxonomy 756 on pages 40-44, DOI: 10.5852/ejt.2021.756.1407, http://zenodo.org/record/503301
Micronecta (Micronecta) clavata Tran 2021, sp. nov.
No full textMicronecta (Micronecta) clavata sp. nov. urn:lsid:zoobank.org:act: B4B1E7DD-177F-498A-9A98-80185920F66D Figs 13C–D, 15 Diagnosis Body length 1.4–1.7. Pronotum longer than median head length. Dorsum colour variable. Hemelytron with irregular-shaped light brown marks, sometimes indistinct (Fig. 13C–D). Males: fore femur with two spines on proximal third ventrally, one spine on distal third dorsally, one pair of spines distally, and one long seta dorsodistally; fore tibia without spine; palar claw moderately widened distally, apex rounded (Fig. 15A). Median lobe of sternite VII with angular apex and three or four long setae (Fig. 15C). Free lobe sub-rectangular, mesial angle rounded, lateral angle nearly square, with ca 10 long setae, posterior margin straight (Fig. 15D). Left paramere: shaft broad, flat, with a triangular projection in the middle of lateral margin, distal part tapering towards rounded apex; basal lobe large, sub-rectangular (Fig. 15F). Right paramere: shaft slender and parallel sided proximally, distal fourth curved, club-shaped with a truncate tip; pars stridens processus with about 20 ridges (Fig. 15E). Etymology The species epithet refers to the club-shaped apex of the right paramere. Material examined Holotype VIETNAM • ♂; Cao Bang Prov., Ha Quang, road 203, Truong Ha commune, stream near Kim Dong tomb; 17 Dec. 2013; A.D. Tran et al. leg.; TAD1371; ZVNU. Paratypes VIETNAM – Cao Bang Prov. • 4 ♂♂, 2 ♀♀; same locality data as for holotype; ZVNU. – Phu Tho Prov. • 2 ♀♀; Xuan Son N.P., Chieng stream, near ranger station; 21°06.072′ N, 104°59.433′ E; 27 Aug. 2014; T.S. Nguyen leg.; TS1408; ZVNU • 1 ♂; Xuan Son N.P., Ha Bang stream, near its confluence with Xoan stream; 21°5.318′ N, 104°58.87′ E; 6 May 2016; T.S. Nguyen leg.; TS1606ĐT; ZVNU • 3 ♂♂, 1 ♀; Xuan Son N.P., Ha Bang stream, Ha Bang village area; 21°04.343′ N, 104°59.182′ E; 7 May 2016; T.S. Nguyen leg.; TS1610; ZRC • 1 ♂; Xuan Son N.P., Xoan stream, Xoan village area; 21°05.270′ N, 104°57.591′ E; 7 May 2016; T.S. Nguyen leg.; TS1609; ZVNU • 1 ♂; Xuan Son N.P., Kim Thuong, Xoan stream; 28 Aug. 2013; A.D. Tran et al. leg.; TAD1343; ZVNU • 1 ♂; Xuan Son N.P., Ban Coi, Coi stream, site 1, water from underground; 29 Aug. 2013; A.D. Tran et al. leg.; TAD1349; ZVNU • 4 ♂♂, 5 ♀♀; Xuan Son N.P., Coi stream, site 3, in front of cave; 21°01.190′ N, 104°56.624′ E; 24 May 2015; T.S. Nguyen leg.; TS1519; ZVNU • 1 ♀; Xuan Son N.P., Coi stream, site 3, in front of cave; 21°01.190′ N, 104°56.624′ E; 27 Aug. 2015; T.S. Nguyen leg.; TS8519 ZVNU • 1 ♂; Xuan Son N.P., Than stream, site 2; 21°11.257′ N, 104°52.382′ E; 24 May 2015; T.S. Nguyen leg.; TS1515; ZVNU. – Hanoi • 2 ♂♂, 1 ♀; Ba Vi N.P., Khoang Xanh, Tien stream; 95 m a.s.l.; 17 Dec. 2011; Q.H. Ngo and A.D. Tran leg.; TAD1127; NHMW • 7 ♂♂, 12 ♀♀; Ba Vi N.P., Khoang Xanh, Tien stream; 95 m a.s.l.; 27 Apr. 2012; Q.H. Ngo and A.D. Tran leg.; TAD1208; ZVNU. – Ha Tinh Prov. • 1 ♀; Vu Quang N.P., Song Con River; 23 Apr. 2003; A.D. Tran leg.; TAD0302; ZRC. Description COLOUR (Fig. 13C–D). With two colour forms, especially colour pattern of hemelytra. Darker form, with distinct dark marks on dorsum (specimens from Phu Tho Prov.): frons and vertex pale yellowish, eyes dark reddish brown. Pronotum light brown. Clavus with a discrete dark brown stripe parallel to suture margin, and a transverse brown mark at anterior margin. Corium with three distinct longitudinal stripes, confluent posteriorly. Membrane translucent and brown. Venter of thorax and abdomen dark brown and legs brownish yellow. Paler form with dorsum generally light yellowish brown (specimens from Cao Bang Prov.): frons and vertex pale, eyes dark reddish brown. Pronotum pale yellowish brown. Hemelytron hyaline light yellowish. Clavus with brown suture margin. Corium with light brown longitudinal stripes indistinct. Membrane transculent, light yellowish. Venter of thorax and abdomen light yellowish orange, legs pale yellow. Structural characteristics MACROPTEROUS MALES. Body length 1.53–1.75 (holotype: 1.69), width 0.84–1.02 (holotype: 0.95). Head width 0.59–0.72 (holotype: 0.67). Head slightly wider than pronotum. Pronotum short, about four times as wide as long (0.61: 0.14). Synthlipsis about 1.4 times as wide as posterior width of an eye (0.27: 0.19). Lengths of leg segments: fore leg: femur 0.22, tibia 0.10, pala 0.13; middle leg: femur 0.52, tibia 0.18, tarsus 0.25, claw 0.16; hind leg: femur 0.43, tibia 0.27, tarsus I 0.30, tarsus II 0.12, claw 0.05. Prestrigilar flap on segment V tongue-shaped (Fig. 15B). Strigil present. Median lobe of sternite VII, free lobe, and parameres as in diagnosis. MACROPTEROUS FEMALES. Body length 1.37–1.81, width 0.72–1.02. Head width 0.59–0.68. Head slightly wider than pronotum. Pronotum about four times as wide as long (0.61: 0.14). Synthlipsis 1.6 times as wide as posterior width of an eye (0.27: 0.17). Lengths of leg segments: fore leg: femur 0.18, tibiotarsus 0.18; middle leg: femur 0.5, tibia 0.18, tarsus 0.25, claw 0.14; hind leg: femur 0.42, tibia 0.27, tarsus I 0.27, tarsus II 0.12, claw 0.04. Distribution Northern Vietnam (Cao Bang, Phu Tho). Remarks Among species of Micronecta occurring in Vietnam, four species, namely M. ornitheia, M. pocsi, M. tuwanoni, and M. clavata sp. nov., are most similar to each other, in having a triangular projection on the margin of the shaft of the left paramere. This structure is also present in the left paramere of M. erythra. However, the right paramere of M. erythra is evenly curved throughout the length of its shaft, while that of four species above is nearly straight or only curved near the base of the shaft. The key structure which can help to reliably separate M. ornitheia, M. pocsi, M. tuwanoni, and M. clavata sp. nov. from each other is the apical part of the right paramere. In M. ornitheia, it is widened, with a small tip making it resemble a bird head. In M. pocsi, it is flap-like, with a small notch on the upper margin before apical part. In M. tuwanoni, the shaft of the right paramere is bent at the distal fourth and the apical part is not widened but tapering towards its tip. In M. clavata sp. nov., it is club-shaped and its tip is more truncate, not modified. The left paramere of these four species also has diagnostic features. In M. ornitheia, the triangular projection is situated near to the base of the shaft, and there is a distinct notch basally to the projection; and the basal lobe is longer than wide, and sub-trapezoid. In M. pocsi, the triangular projection is also nearer to the base of shaft, but there is no notch on the lateral margin; and the basal lobe is sub-triangular. In M. tuwanoni, the triangular projection arises on the proximal third (thus nearer to the base) of the shaft, and the basal lobe is trapezoid. In M. clavata sp. nov., the triangular projection is near to the middle part of the lateral margin of the shaft, and the basal lobe is sub-rectangular.Published as part of Ha, Tuyet Ngan & Tran, Anh Duc, 2021, Taxonomy of Micronectidae (Heteroptera: Nepomorpha) from Vietnam, with descriptions of 11 new species, pp. 1-82 in European Journal of Taxonomy 756 on pages 49-51, DOI: 10.5852/ejt.2021.756.1407, http://zenodo.org/record/503301
Micronecta caperata Tran 2021, sp. nov.
No full textMicronecta caperata sp. nov. urn:lsid:zoobank.org:act: 826E598E-CEF2-4A77-AA7B-6403633B323C Figs 21E, 24 Diagnosis Body length 1.5–1.9. Pronotum distinctly shorter than median head length; lateral margin of pronotum very short. Dorsum colour yellowish brown. Hemelytron punctate, with scattered small light brownish spots; suture between clavus, corium, and membrane indistinct (Fig. 21E). Males: fore femur with two spines in distal third dorsally; fore tibia without spine; palar claw narrow, evenly curved (Fig. 24A). Median lobe of sternite VII with angular apex and four long setae (Fig. 24B). Free lobe absent. Left paramere: shaft broad and flat, distal part with ca five transverse wrinkles, apex rounded; basal lobe longer than wide, sub-trapezoid (Fig. 24D). Right paramere: shaft relatively slender, tapering towards apex, gently curved at distal third, apex narrowly rounded (Fig. 24C); pars stridens processus with ca 30 ridges. Etymology The species epithet refers to the wrinkles on the left paramere. Material examined All are hindwing-brachypterous. Holotype VIETNAM • ♂; Nghe An Prov., Pu Mat N.P., Khe Kem stream, site #3; 20 Dec. 2012; Q.H. Ngo leg.; NQH1205; ZVNU. Paratypes VIETNAM – Nghe An Prov. • 7 ♂♂, 1 ♀; same locality data as for holotype; ZVNU • 4 ♂♂, 1 ♀; Pu Mat N.P., Khe Kem stream, site #4; 20 Dec. 2012; Q.H. Ngo leg.; NQH1206; ZVNU • 2 ♂♂, 1 ♀; Pu Mat N.P., Khe Kem stream, site #5; 20 Dec. 2012; Q.H. Ngo leg.; NQH1207; ZVNU • 6 ♂♂, 2 ♀♀; Pu Mat N.P., Khe Kem stream, site #6; 20 Dec. 2012; Q.H. Ngo leg.; NQH1208; ZVNU • 3 ♂♂; Pu Mat N.P., Khe Kem stream, site #5; 18 Apr. 2012; Q.H. Ngo leg.; TAD1314; ZVNU • 9 ♂♂, 5 ♀♀; Pu Mat N.P., Chat stream, tributary of Choang stream; 1 Sep. 2020; A.D. Tran et al. leg.; TAD20-14; ZVNU • 4 ♂♂, 2 ♀♀; same collection data as for preceding; ZRC • 1 ♂; Pu Mat N.P., Choang stream, near Ban Bu; 1 Sep. 2020; A.D. Tran and T.C. Nguyen leg.; TAD20-17; NHMW. – Cao Bang Prov. • 1 ♂; Hoa An, stream at Ban Say bridge; 17 Dec. 2013; A.D. Tran et al. leg.; TAD1372; ZVNU • 2 ♂♂; Quang Uyen, road 207, Doc Lap commune, stream #2; 18 Dec. 2013; A.D. Tran et al. leg.; TAD1374; ZVNU. – Phu Tho Prov. • 1 ♂; Xuan Son N.P., Ban Coi, Coi stream, site 3, by the road to Tan Son; 7 Jun. 2013; A.D. Tran et al. leg.; TAD1339; ZVNU • 1 ♂; Xuan Son N.P., Xoan stream, near its confluence with Ha Bang stream; 21°5.303′ N, 104°57.559′ E; 22 May 2015; T.S. Nguyen leg.; TS1507; ZVNU • 1 ♀; Xuan Son N.P., Xoan stream, near its confluence with Ha Bang stream; 21°5.303′ N, 104°57.559′ E; 6 May 2016; T.S. Nguyen leg.; TS1607; ZVNU • 2 ♀♀; Xuan Son N.P., Ha Bang stream, near its confluence with Xoan stream; 21°5.318′ N, 104°58.87′ E; 22 May 2015; T.S. Nguyen leg.; TS1506; ZVNU • 2 ♂♂; Xuan Son N.P., Coi stream, site 1, by the road to Tan Son; 21°09.204′ N, 104°56.927′ E; 8 May 2016; T.S. Nguyen leg.; TS1617; ZVNU • 2 ♀♀; Ha Bang stream, Ha Bang village area; 21°04.343′ N, 104°59.182′ E; 7 May 2016; T.S. Nguyen leg.; TS1610; ZVNU. – Lam Dong Prov. • 1 ♂; Bao Loc, Damb’ri waterfall; 19 May 2003; A.D. Tran leg.; TAD0326; ZRC. Description COLOUR (Fig. 21E). Frons and vertex light yellowish, eyes reddish. Pronotum light brownish yellow. Hemelytron light yellow with scattered small light brownish spots. Membrane with similar colour to corium, but more translucent. Hindwings absent. Venter of thorax and abdomen, and legs pale yellow. Structural characteristics HINDWING- BRACHYPTEROUS MALES. Body length 1.57–1.88 (holotype: 1.75), width 0.81–1.03 (holotype: 1.00). Head width 0.6–0.72 (holotype: 0.68). Head slightly wider than pronotum. Pronotum very short, mostly covered by head, about 7.2 times as wide as long (0.65: 0.09). Synthlips about 1.35 times as wide as posterior width of an eye (0.27: 0.20). Lengths of leg segments: fore leg: femur 0.24, tibia 0.13, pala 0.12; middle leg: femur 0.58, tibia 0.2, tarsus 0.29, claw 0.2; hind leg: femur 0.42, tibia 0.3, tarsus I 0.3, tarsus II 0.16, claw 0.05. Prestrigilar flap on segment V absent. Strigil present. Median lobe of sternite VII, and parameres as in diagnosis. HINDWING- BRACHYPTEROUS FEMALES. Body length 1.56–1.9, width 0.86–1.03. Head width 0.6–0.74. Head slightly wider than pronotum. Pronotum very short, mostly covered by head, about 7 times as wide as long (0.67: 0.10). Synthlips about 1.4 times as wide as posterior width of an eye (0.29: 0.21). Lengths of leg segments: fore leg: femur 0.23, tibiotarsus 0.2; middle leg: femur 0.59, tibia 0.22, tarsus 0.26, claw 0.18; hind leg: femur 0.38, tibia 0.3, tarsus I 0.32, tarsus II 0.17, claw 0.05. Distribution Vietnam: Cao Bang, Phu Tho, Nghe An, Lam Dong. Remarks Micronecta caperata sp. nov. has a mixed set of characteristics that does not allow its placement in any of the currently recognised subgenera of Micronecta. It is probably the only species of Micronecta that has no free lobe of tergite VIII. Its palar claw is very narrow and constricted pre-apically, which is similar to that of the subgenus Indonectella (resepresented by only one species, M. grisea). However, M. caperata sp. nov. has a strigil on segment VI, while the strigil is absent in Indonectella. The very short pronotum of this species is similar to that of M. johorensis Fernando, 1964, M. dentifera Nieser, 2002, and M. eucosmeta Hutchinson, 1940, but the parameres of M. caperata sp. nov. are completely different from those of these three species. The left paramere is broad and flat, which is a characteristic of the nominate subgenus Micronecta. The outline of the right paramere of M. caperata sp. nov. is relatively similar to that of M. ceylonica Wrósblewski, 1968, but its left paramere is distinctly different from that of the latter. The left paramere, with wrinkles on distal part of shaft, is unique for M. caperata sp. nov. Although this new species does not fit into any of the recognised subgenus, we refrain from establishing a new subgenus for a single species, until an intrageneric phylogenetic analysis of Micronecta becomes available. We thus temporarily do not assign this species to any of the current subgenera.Published as part of Ha, Tuyet Ngan & Tran, Anh Duc, 2021, Taxonomy of Micronectidae (Heteroptera: Nepomorpha) from Vietnam, with descriptions of 11 new species, pp. 1-82 in European Journal of Taxonomy 756 on pages 70-72, DOI: 10.5852/ejt.2021.756.1407, http://zenodo.org/record/503301
Micronecta (Micronecta) undulata Tran 2021, sp. nov.
No full textMicronecta (Micronecta) undulata sp. nov. urn:lsid:zoobank.org:act: 98F8AC39-93D8-4C61-900B-B3BD1F21B676 Figs 13F, 17 Diagnosis Body length 1.9–2.2. Pronotum longer than median head length. Hemelytron with a broad, transverse, dark brown mark on greyish background (Fig. 13F). Males: fore femur with two spines on middle third, one or two spines distally, and no long seta dorsodistally; fore tibia with one small spine on proximal third dorsally, and three spines on distal third; palar claw distally moderately widened, apex rounded (Fig. 17A). Median lobe of sternite VII with narrow, pointed apex and four long setae (Fig. 17C). Free lobe short, distally widened and wavy, both angles nearly square, without long setae; posterior margin strongly sinuate (Fig. 17D). Left paramere: shaft relatively short and broad, apex truncate, with two ridges running to lateral margin, thus forming two triangular projections on lateral margin; basal lobe longer than wide, sub-rectangular (Fig. 17F). Right paramere: shaft parallel sided, slightly curved at proximal part, abruptly constricted at distal third, apical part wavy and lobate; pars stridens processus with 18–27 ridges (Fig. 17E). Etymology The species epithet refers to the wavy, undulating free lobe. Material examined Holotype VIETNAM • ♂; Phu Tho Prov., Xuan Son N.P., Coi stream, site 1, by the road to Tan Son; 21°09.204′ N, 104°56.927′ E; 27 Aug. 2015; T.S. Nguyen leg.; TS8517; ZVNU. Paratypes VIETNAM – Phu Tho Prov. • 10 ♂♂, 9 ♀♀; same locality data as for holotype; ZVNU • 4 ♂♂, 3 ♀♀; same locality data as for holotype; NHMW • 2 ♂♂, 1 ♀; Xuan Son N.P., Ban Coi, Coi stream, site 2, near bridge; 7 Jun. 2013; A.D. Tran et al. leg.; TAD1338; ZVNU • 4 ♂♂, 2 ♀♀; Xuan Son N.P., Ban Coi, Coi stream, site 3, by the road to Tan Son; 7 Jun. 2013; A.D. Tran et al. leg.; TAD1339; ZVNU • 1 ♂, 1 ♀; Xuan Son N.P., Kim Thuong, Xoan stream; 28 Aug. 2013; A.D. Tran et al. leg.; TAD1343; ZVNU • 3 ♂♂, 4 ♀♀; Xuan Son N.P., Ban Coi, Coi stream, site 3, by the road to Tan Son; 29 Aug. 2013; A.D. Tran et al. leg.; TAD1351; ZRC • 11 ♂♂, 11 ♀♀; Xuan Son N.P., Xoan stream, near its confluence with Ha Bang stream; 21°5.303′ N, 104°57.559′ E; 27 Aug. 2014; T.S. Nguyen leg.; TS1407; ZVNU • 2 ♀♀; Xuan Son N.P., Xoan stream, near its confluence with Ha Bang stream; 21°5.303′ N, 104°57.559′ E; 6 May 2016; T.S. Nguyen leg.; TS1607; ZVNU • 4 ♂♂, 6 ♀♀; Xuan Son N.P., Xoan stream, Xoan village area; 21°05.270′ N, 104°57.591′ E; 23 May 2015; T.S. Nguyen leg.; TS1509; ZVNU • 1 ♀; Xuan Son N.P., Xoan stream, Xoan village area; 21°05.270′ N, 104°57.591′ E; 7 May 2016; T.S. Nguyen leg.; TS1609; ZVNU • 1 ♀; Xuan Son N.P., Than stream, site 2; 21°11.257′ N, 104°52.382′ E; 24 May 2015; T.S. Nguyen leg.; TS1515; ZVNU • 1 ♂, 1 ♀; Xuan Son N.P., Than stream, site 3; 21°11.393′ N, 104°52.625′ E; 28 Aug. 2014; T.S. Nguyen leg.; TS1416; ZVNU • 8 ♂♂, 13 ♀♀; Xuan Son N.P., Than stream, site 3; 21°11.393′ N, 104°52.625′ E; 27 Aug. 2015; T.S. Nguyen leg.; TS8516; ZVNU • 3 ♂♂, 2 ♀♀; Xuan Son N.P., Ha Bang stream, near its confluence with Xoan stream; 21°5.318′ N, 104°58.87′ E; 26 Aug. 2014; T.S. Nguyen leg.; TS1406; ZVNU • 1 ♂; Xuan Son N.P., Ha Bang stream, near its confluence with Xoan stream; 21°5.318′ N, 104°58.87′ E; 6 May 2016; T.S. Nguyen leg.; TS1606ĐT; ZVNU • 1 ♂, 2 ♀♀; Xuan Son N.P., Tan Ong stream, 4 km to Chin Tang stream; 21°04.288′ N, 104°57.271′ E; 26 Aug. 2014; T.S. Nguyen leg.; TS1405; ZVNU • 3 ♂♂, 1 ♀; Xuan Son N.P., Tan Ong stream, 4 km to Chin Tang stream; 21°04.288′ N, 104°57.271′ E; 22 May 2015; T.S. Nguyen leg.; TS1505; ZVNU • 1 ♂; Xuan Son N.P., Coi stream, site 1, by the road to Tan Son; 21°09.204′ N, 104°56.927′ E; 8 May 2016; T.S. Nguyen leg.; TS1617; ZVNU • 8 ♂♂, 2 ♀♀; Xuan Son N.P., Coi stream, site 2, near concrete bridge; 21°09.130′ N, 104°56.761′ E; 24 May 2015; T.S. Nguyen leg.; TS1518; ZVNU. Description COLOUR (Fig. 13F). Dorsum generally brownish grey. Frons and vertex pale yellowish, eyes blackish brown. Pronotum greyish brown. Hemelytron with greyish background. Clavus with brown punctures near anterior margin. Corium with a broad transverse dark brown mark in the middle. Membrane generally dark brown, more hyaline apically. Venter of thorax and abdomen greyish brown and legs pale yellow or nearly whitish. Structural characteristics MACROPTEROUS MALES. Body length 1.94–2.19 (holotype: 2.03), width 0.94–1.06 (holotype: 1.00). Head width 0.81–0.83 (holotype: 0.76). Pronotum about 2.4 times as wide as long (0.87: 0.36). Pronotum slightly wider than head. Synthlipsis about 1.3 times as wide as posterior width of an eye (0.37: 0.29). Lengths of leg segments: fore leg: femur 0.32, tibia 0.14, pala 0.14; middle leg: femur 0.67, tibia 0.25, tarsus 0.34, claw 0.21; hind leg: femur 0.56, tibia 0.40, tarsus I 0.39, tarsus II 0.16, claw 0.16. Prestrigilar flap on segment V crescent-shaped (Fig. 17B). Strigil present. Median lobe of sternite VII, free lobe, and parameres as in diagnosis. MACROPTEROUS FEMALES. Body length 1.94–2.03, width 1.0–1.03. Head width 0.81–0.82. Pronotum about 2.6 times as wide as long (0.88:0.34). Pronotum slightly wider than head. Synthlipsis about 1.4 times as wide as posterior width of an eye (0.39: 0.27). Lengths of leg segments: fore leg: femur 0.29, tibiotarsus 0.27; middle leg: femur 0.72, tibia 0.29, tarsus 0.36, claw 0.22; hind leg: femur 0.52, tibia 0.36, tarsus I 0.42, tarsus II 0.16, claw 0.16. Distribution Vietnam: Phu Tho. Remarks See Remarks under M. pingae sp. nov.Published as part of Ha, Tuyet Ngan & Tran, Anh Duc, 2021, Taxonomy of Micronectidae (Heteroptera: Nepomorpha) from Vietnam, with descriptions of 11 new species, pp. 1-82 in European Journal of Taxonomy 756 on pages 54-56, DOI: 10.5852/ejt.2021.756.1407, http://zenodo.org/record/503301
Micronecta (Micronecta) cultellata Tran 2021, sp. nov.
No full text<i>Micronecta</i> (<i>Micronecta</i>) <i>cultellata</i> sp. nov. <p>urn:lsid:zoobank.org:act: A59D915E-EEA7-4278-B2B0-4C0707189B6F</p> <p>Figs 5E, 8</p> Diagnosis <p>Body length 1.6–2.2. Pronotum slightly shorter than median head length, lateral margin of pronotum short. Hemelytron punctate, orangish (Fig. 5E).</p> <p>Males: fore femur with a pair of spines on proximal third ventrally, one spine in distal third, three spines distally, and one long seta dorsodistally; fore tibia without spine; palar claw broad, apex rounded (Fig. 8A). Median lobe of sternite VII short and broad, with an angular apex and four long setae (Fig. 8C). Free lobe long, mesial angle obtuse, lateral angle square, with ca 20 long setae (Fig. 8D). Left paramere: shaft broad, flattened, apex narrowly rounded; basal lobe sub-rectangular (Fig. 8F). Right paramere: shaft evenly curved, distal half parallel sided, apex rounded; pars stridens processus with ca 30 ridges (Fig. 8E).</p> Etymology <p>The species epithet refers to the shape of the left paramere’s shaft, which resembles the blade of a knife.</p> Material examined <p> <b>Holotype</b></p> <p> VIETNAM • ♂; Lao Cai Prov., Sa Pa, Trung Chai, km 119 Nat. Road #4D; 28 Oct. 2013; Q.H. Ngo <i>et al.</i> leg.; TAD1370; ZVNU.</p> <p> <b>Paratypes</b></p> <p> VIETNAM – <b>Lao Cai Prov.</b> • 3 ♂♂, 3 ♀♀; same locality data as for holotype; ZVNU • 2 ♂♂, 2 ♀♀; same locality data as for holotype; NHMW • 4 ♂♂; Sa Pa, Nam Cang, Nam Cang stream; 23 Oct. 2012; N.H. Dinh <i>et al.</i> leg.; DNH12.08; ZVNU • 3 ♂♂; same collection data as for preceding; ZRC • 1 ♀; Sa Pa, waterfall by roadside from Nui Xe to Sa Pa town, ca 13 km from Sa Pa; 25 Oct. 2013; A.D. Tran <i>et al.</i> leg.; TAD1358b; ZVNU • 1 ♀; Sa Pa, Thanh Phú, N ậm Cang stream; 26 Oct. 2013; A.D. Tran <i>et al.</i> leg.; TAD1359; ZVNU • 1 ♀; Sa Pa, Cat Cat, Ho stream (tributary of Muong Hoa stream); 27 Oct. 2013; A.D. Tran <i>et al.</i> leg.; TAD1366; ZVNU. – <b>Ha Giang Prov.</b> • 1 ♂; Vi Xuyen, Xin Chai commune, Da stream by the road from Thanh Thuy to Lao Chai; 9 May 2014; A.D. Tran <i>et al.</i> leg.; TAD1405; ZVNU • 1 ♂, 1 ♀; Xin Man, Nam Dan commune, Thac Tien-Deo Gio, stream 100 m below waterfall; 11 May 2014; A.D. Tran <i>et al.</i> leg.; TAD1413; ZVNU. – <b>Cao Bang Prov.</b> • 2 ♀♀; Quang Uyen, road 207, Doc Lap commune, stream #2; 18 Dec. 2013; A.D. Tran <i>et al.</i> leg.; TAD1374; ZVNU.</p> Description <p>COLOUR (Fig. 5E). Dorsum generally light orangish. Frons and vertex pale yellowish, eyes reddish brown. Pronotum light greyish brown. Hemelytron light orangish, without distinct mark. Suture between clavus and corium brownish, a band with minute red dots along anterior margin of clavus. Membrane translucent apically. Venter of thorax and abdomen, and legs brownish yellow.</p> Structural characteristics <p>MACROPTEROUS MALES. Body length 1.78–2.16 (holotype: 1.93), width 1.0–1.16 (holotype: 1.13). Head width 0.75–0.78 (holotype: 0.70), slightly wider than pronotum. Pronotum about four times as wide as long, width 0.71, length 0.18. Synthlipsis about twice the posterior width of an eye (0.35: 0.18). Lengths of leg segments: fore leg: femur 0.30, tibia 0.17, pala 0.14; middle leg: femur 0.65, tibia 0.20, tarsus 0.34, claw 0.16; hind leg: femur 0.45, tibia 0.36, tarsus I 0.36, tarsus II 0.18, claw 0.05. Prestrigilar flap on segment V elongated, with weakly acute apex (Fig. 8B). Strigil present. Median lobe of sternite VII, free lobe, and parameres as in diagnosis.</p> <p>MACROPTEROUS FEMALES. Body length 1.63–2.09, width 0.9–1.19. Head width 0.74–0.80, slightly wider than pronotum. Pronotum about four times as wide as long, width 0.75, length 0.19. Synthlipsis about twice the posterior width of an eye (0.36: 0.19). Lengths of leg segments: fore leg: femur 0.27, tibiotarsus0.21; middle leg: femur 0.76, tibia 0.27, tarsus 0.32, claw 0.20; hind leg: femur 0.41, tibia 0.32, tarsus I 0.36, tarsus II 0.16, claw 0.09.</p> Distribution <p>Vietnam: northern area (Cao Bang, Ha Giang, Lao Cai).</p> Remarks <p> <i>Micronecta cultellata</i> sp. nov. is similar to <i>M. drepani</i> and <i>M. erythra</i> in the general form of the right paramere, with the evenly curved shaft, and in the shape of the free lobe. These three species can be separated from each other by the characteristics as follows. <i>Micronecta erythra</i> has a distinct colour pattern on the hemelytra and a distinct triangular projection on the lateral side of the left paramere. <i>Micronecta drepani</i> has dark marks on the hemelytra, but they do not form a distinct pattern. The left paramere of <i>M. drepani</i> has a short ridge running onto the lateral margin of the shaft, and the apex of the shaft is broad. <i>Micronecta cultellata</i> sp. nov. has a light orangish dorsum, without dark markings, and its left paramere lacks modifications on the lateral margin, with the apex narrow. The right paramere of <i>M. drepani</i> differs from that of <i>M. erythra</i> and <i>M. cultellata</i> sp. nov. by having a thicker, swollen basal part of the shaft (on the mesial side). This structure is not swollen in <i>M. erythra</i> and <i>M. cultellata</i> sp. nov., but gradually tapering to the distal part instead.</p>Published as part of <i>Ha, Tuyet Ngan & Tran, Anh Duc, 2021, Taxonomy of Micronectidae (Heteroptera: Nepomorpha) from Vietnam, with descriptions of 11 new species, pp. 1-82 in European Journal of Taxonomy 756</i> on pages 30-32, DOI: 10.5852/ejt.2021.756.1407, <a href="http://zenodo.org/record/5033010">http://zenodo.org/record/5033010</a>
Growth and nodulation of symbiotic Medicago truncatula at different levels of phosphorus availability
No full textMedicago truncatula is an important model plant for characterization of P deficiency on leguminous plants at the physiological and molecular levels. Growth optimization of this plant with regard to P supply is the first essential step for elucidation of the role of P in regulation of nodulation. Hence, a study was carried out to address the growth pattern of M. truncatula hydroponically grown at different gradual increases in P levels. The findings revealed that M. truncatula had a narrow P regime, with an optimum P level (12 M P) which is relatively close to the concentration that induces P toxicity. The accumulated P concentration (2.7 mg g(1) dry matter), which is normal for other crops and legumes, adversely affected the growth of M. truncatula plants. Under P deficiency, M. truncatula showed a higher symbiotic efficiency with Sinorhizobium meliloti 2011 in comparison with S. meliloti 102F51, partially as a result of higher electron allocation to N-2 versus H. The total composition of free amino acids in the phloem was significantly affected by P deprivation. This pattern was found to be almost exclusively the result of the increase in the asparagine level, suggesting that asparagine might be the shoot-derived signal that translocates to the nodules and exerts the down-regulation of nitrogenase activity. Additionally, P deprivation was found to have a strong influence on the contents of the nodule carbon metabolites. While levels of sucrose and succinate tended to decrease, a higher accumulation of malate was observed. These findings have provided evidence that N-2 fixation of M. truncatula is mediated through an N feedback mechanism which is closely related to nodule carbon metabolism
Micronecta (Micronecta) ornitheia Nieser, Chen & Yang 2005
No full textMicronecta (Micronecta) ornitheia Nieser, Chen & Yang, 2005 Figs 13A, 14A–D Micronecta ornitheia Nieser, Chen & Yang, 2005: 190, 197–200, figs 32–41 (type locality: Yunnan, China). Micronecta ornitheia – Chen, Nieser & Sangpradub 2006: 779 (record Thailand). Diagnosis Body length 1.7–1.8. Pronotum longer than median head length. Dorsum generally light brown. Hemelytron with small, irregular-shaped reddish marks, mostly on costal part; embolium with a longitudinal dark grey stripe and two brown patches (Fig. 13A). Males: fore femur with two spines on middle third, a spine on distal third dorsally, and a pair of spines distally; fore tibia with three large and one small spine on distal third; palar claw distally widened, apical margin rounded. Median lobe of sternite VII narrow, with pointed apex and four long setae (Fig. 14A). Free lobe slightly expanded distally, mesial angle rounded, lateral angle slightly produced with ca 20 long setae, posterior margin nearly straight (Fig. 14B). Left paramere: shaft broad, flat, with a deep notch along lateral margin near base, then followed by a triangular projection, distal part parallel sided, apex of paramere rounded; basal lobe longer than wide, sub-trapezoid (Fig. 14D). Right paramere: shaft slender proximally and club-shaped apically (Fig. 14C). Material examined VIETNAM – Lao Cai Prov. • 36 ♂♂, 31 ♀♀; Sa Pa, Thanh Phu, near junction of Nam Cang and Muong Hoa streams; 26 Oct. 2013; Q.H. Ngo leg.; TAD1360; ZVNU • 3 ♂♂, 4 ♀♀; Sa Pa, Hoang Lien N.P., Cat Cat; 1 Jun. 2003; A.D. Tran leg.; TAD0336; ZRC. – Ha Giang Prov. • 3 ♂♂, 4 ♀♀; Bac Me, road 176, Minh Son commune, stream at Ban Van bridge; 7 May 2014; A.D. Tran et al. leg.; TAD1403; ZVNU • 1 ♂, 6 ♀♀; Yen Minh, Na Khe commune, stream near Nat’ road 4C; 8 May 2014; A.D. Tran et al. leg.; TAD1404; ZVNU • 2 ♂♂; Vi Xuyen, Xin Chai commune, Da stream by the road from Thanh Thuy to Lao Chai; 9 May 2014; A.D. Tran et al. leg.; TAD1405; ZVNU • 15 ♂♂, 8 ♀♀; Vi Xuyen, Nat’ road 2, Phuong Tien commune, Suu stream, near bridge; 9 May 2014; A.D. Tran et al. leg.; TAD1406; ZVNU • 7 ♂♂, 11 ♀♀; Vi Xuyen, Nat’ road 2, Phuong Tien commune, Suu stream, near bridge; 9 May 2014; A.D. Tran et al. leg.; TAD1407; ZVNU • 1 ♀; Vi Xuyen, National road 4C, Thuan Hoa commune, Nam Dieng stream, ca 12.5 km to Ha Giang city; 9 May 2014; A.D. Tran et al. leg.; TAD1409; ZVNU • 1 ♂, Xin Man, road 178, tributary of Nam Yen River, near bridge ca 2 km from Coc Pai town; 11 May 2014; A.D. Tran et al. leg.; TAD1412; ZVNU. – Cao Bang Prov. • 5 ♂♂, 20 ♀♀; Trung Khanh, Duc Hong commune, stream; 19 Dec. 2013; A.D. Tran et al. leg.; TAD1376; ZVNU • 1 ♂, 4 ♀♀; Bao Lac, Dinh Phung commune, near Tung Suu, stream by the Nat’ road 34; 20 Dec. 2013; A.D. Tran et al. leg.; TAD1378; ZVNU • 20 ♂♂, 29 ♀♀; Bao Lac, Hung Dao commune, Nhi Ao River, near Nam Xui bridge; 20 Dec. 2013; A.D. Tran et al. leg.; TAD1380; ZVNU • 3 ♂♂, 12 ♀♀; Bao Lac, Bao Toan commune, Na Roa Village, Khuoi Rong stream, tributary of Gam River; 20 Dec. 2013; A.D. Tran et al. leg.; TAD1381; ZVNU. – Phu Tho Prov. • 1 ♂, 2 ♀♀; Xuan Son N.P., Ban Coi, Coi stream, site 3, by the road to Tan Son; 7 Jun. 2013; A.D. Tran et al. leg.; TAD1339; ZVNU • 2 ♂♂, 1 ♀; Xuan Son N.P., Ha Bang stream, near its confluence with Xoan stream; 21°5.318′ N, 104°58.87′ E; 22 May 2015; T.S. Nguyen leg.; TS1506; ZVNU • 1 ♂, 2 ♀♀; Xuan Son N.P., Xoan stream, near Lang Village; 21°5.453′ N, 104°56.122′ E; 21 May 2015; T.S. Nguyen leg.; TS1501; ZVNU • 1 ♂; Xuan Son N.P., Xoan stream, Xoan village area; 21°05.270′ N, 104°57.591′ E; 7 May 2016; T.S. Nguyen leg.; TS1609; ZVNU • 1 ♂; Xuan Son N.P., Chieng stream, near ranger station; 21°06.072′ N, 104°59.433′ E; 27 Aug. 2014; T.S. Nguyen leg.; TS1408; ZVNU • 1 ♂, 4 ♀♀; Xuan Son N.P., Chieng stream, near ranger station; 21°06.072′ N, 104°59.433′ E; 22 May 2015; T.S. Nguyen leg.; TS1508; ZVNU • 1 ♂, 5 ♀♀; Xuan Son N.P., Chieng stream, near ranger station; 21°06.072′ N, 104°59.433′ E; 25 Aug. 2015; T.S. Nguyen leg.; TS8508; ZVNU • 2 ♂♂, 3 ♀♀; Xuan Son N.P., Chieng stream, near ranger station; 21°06.072′ N, 104°59.433′ E; 6 May 2016; T.S. Nguyen leg.; TS1608; ZVNU • 2 ♂♂, 3 ♀♀; Xuan Son N.P., Coi stream, site 1, by the road to Tan Son; 21°09.204′ N, 104°56.927′ E; 8 May 2016; T.S. Nguyen leg.; TS1617; ZVNU. – Thanh Hoa Prov. • 5 ♂♂, 1 ♀; Pu Hu Nature Reserve, Muong Lat, Trung Ly, Co Luong, Cha Lan stream; 4 Apr. 2013; T.D. Pham leg.; PH1301; ZVNU • 2 ♂♂; Pu Luong Nature Reserve, Ba Thuoc, Co Lung, Ban Khuyn, Khuyn stream; 18 Mar. 2013; T.D. Pham leg.; PL1303; ZVNU. – Nghe An Prov . • 24 ♂♂, 29 ♀♀; Ky Son, Nat’ road 7, Ta Ca commune, Loi stream; 16 Apr. 2013; A.D. Tran et al. leg.; TAD1301; ZVNU • 9 ♂♂, 5 ♀♀; Ky Son, Nat’ road 7 at km 191+572, stream at Ban Na bridge; 16 Apr. 2013; A.D. Tran et al. leg.; TAD1304; ZVNU • 43 ♂♂, 34 ♀♀; Ky Son, Nat’ road 7 at km 182+950, stream at Khe Nan bridge; 16 Apr. 2013; A.D. Tran et al. leg.; TAD1305; ZVNU • 1 ♂, 1 ♀; Con Cuong, Lien Huong Village, Khe Thoi stream; 21 Dec. 2012; Q.H. Ngo leg.; NQH1209; ZVNU • 2 ♂♂, 3 ♀♀; Con Cuong, Tung Huong Village, Khe Thoi stream; 21 Dec. 2012; Q.H. Ngo leg.; NQH1210; ZVNU. – Ha Tinh Prov . • 9 ♂♂, 4 ♀♀; Vu Quang N.P., Song Con River; 23 Apr. 2003; A.D. Tran leg.; TAD0302; ZRC. Distribution China, Thailand (Nieser et al. 2005; Chen et al. 2006). First records for Vietnam. Remarks Nieser et al. (2005) already noted the similarity (right paramere) and differences (body length, dorsal colour pattern, and left paramere) between M. ornitheia and M. sanctaecatherinae Hutchinson, 1940, which is only known from India and Sri Lanka. Nieser et al. (2005) also made a comparison between the parameres of M. ornitheia, M. pocsi and related species from China and India. See also Remarks under M. clavata sp. nov. for additional differences between M. ornitheia and other similar species occurring in Vietnam.Published as part of Ha, Tuyet Ngan & Tran, Anh Duc, 2021, Taxonomy of Micronectidae (Heteroptera: Nepomorpha) from Vietnam, with descriptions of 11 new species, pp. 1-82 in European Journal of Taxonomy 756 on pages 44-48, DOI: 10.5852/ejt.2021.756.1407, http://zenodo.org/record/503301
Phallostethus cuulong Shibukawa, Tran & Tran, 2012, new species
No full textPhallostethus cuulong, new species New Vietnamese name: Cá bụng đầu (Figures 1–4) Holotype. ZRC 53233, male, 24.2 mm SL, a branch of Hau River (a distributary of Mekong), Cu Lao Dung, Soc Trang Province, Vietnam (9 ° 30.8 ’ N, 106 ° 13.7 ’ E), 0–0.5 m depth, 31 July 2009, collected by K. Shibukawa. Paratypes. Total eight specimens (five males and three females), 20.0– 24.5 mm SL: CTU-P 2327, 1 specimen (female), 23.7 mm SL, Duyen Hai, Tra Vinh Province, Vietnam (9 ° 40.9 ’ N, 106 ° 30.7 ’ E), 0.3–0.8 m depths, 5 April 2009, collected by K. Shibukawa, V.V. Tran and L.X. Tran; CTU-P 2494, 1 specimen (male), 22.5 mm SL, My Thanh River (a distributary of Mekong), Vinh Chau, Soc Trang Province, Vietnam (9 ° 22.7 ’ N, 106 ° 0.7 ’ E), 0.5–1.2 m depths, 1 August 2009, collected by H.P. Ha, V.V. Tran and L.X. Tran; CTU-P 5020, 1 specimen (male, cleared and stained), 23.5 mm SL, Cho Lach, Ben Tre, Vietnam (10 ° 10.5 ’ N, 106 °. 8.9 ’ E), 0.5 m depth, 3 February 2010, collected by L.X. Tran; NSMT-P 106664, 1 specimen (male), 20.0 mm SL, collected with CTU-P 2494; NSMT-P 106665, 1 specimen (female), 22.0 mm SL, collected with CTU-P 2494; USNM 404477, 1 specimen (female), 23.8 mm SL, collected with CTU-P 2494; USNM 404478, 1 specimen (male), 20.3 mm SL, Cau Ke, Tra Vinh Province, Vietnam (9 ° 57.1 ’ N, 106 ° 1.8 ’ E), 0.5–3.5 m depths, 28 May 2010, collected by L.X. Tran; USNM 404479, 1 specimen (male, cleared and stained), 24.5 mm SL, collected with CTU-P 5020. Diagnosis. Phallostethus cuulong is distinguished from congeners in having following characters: seven serrae on the second ctenactinium in adult males (vs. five and eight in P. dunckeri and P. l e h i, respectively); 11–13 pectoral-fin rays (vs. 9–10 and 12 in P. dunckeri and P. l e h i, respectively); 11–14 + 25-26 = 37–40 vertebrae (vs. 13 + 27 = 40 and 12 + 28 = 40 in P. dunckeri and P. l e h i); approximately 5–19 teeth on paradentary (vs. 15–20 and 28 or more in P. dunckeri and P. le h i, respectively). All six examined males are dextral (vs. one and two known males of P. dunckeri are sinistral and dextral respectively, and all four know males of P. l e h i are sinistral). Description. Counts of the holotype are asterisked, and the frequency of each count is given in parentheses following the relevant count. Dorsal-fin rays 7 (2), 8 * (6) or 9 (1); anal-fin rays 24 * (4), 25 (1), 26 (3) or 27 (1); pectoral-fin rays 11 (2), 12 * (7) or 13 * (9); scales in lateral series 34 (5), 35 * (10) or 36 (3); predorsal scales 1 + 25 (1), 2 + 25 (1), 2 + 26 (4) or 2 + 27 * (3); transverse scales 6 (3), 7 * (12) or 8 * (3); circumpeduncular scales 12 * (8) or 13 (1); paradentary teeth approximately 5 (1), 6 (3), 7 (3), 8 (3), 10 (1), 13 * (1), 14 (2), 15 * (1), 18 (2) or 19 (1). The following measurements are % of SL: head length 22.1–24.1; snout length 7.0– 8.5; eye diameter 6.7–7.3; interorbital width 3.3–5.2; length of jaw 8.0– 9.4; predorsal length 78.0– 82.6; preanal length 46.4–48.7; maximum body depth 15.0– 18.7; body depth at anal-fin origin 12.4 –15.0; body width 9.4 –14.0; caudal-peduncle length 18.6–20.7; caudal-peduncle depth 5.6–7.6; length of dorsal-fin base 9.1–10.5; length of anal-fin base 32.3–36.7; pectoral-fin length 14.6–19.1; caudal-fin length 20.1–22.4. Head depressed anteriorly, with flat or barely concave interorbital space. Dorsal surface of head with membranous dome when alive or freshly collected (which can be seen in the cleared and stained specimens, e.g., Fig. 4), but shrunken and not apparent in alcohol specimen. Snout rather pointed. Eyes lateral on head, large, diameter slightly less than snout length. Mouth terminal or subterminal; jaws small, barely extend to a level of anterior margin of eye; upper jaws highly protractile. Body compressed, moderately deep. Anus and urogenital openings anterior, ventral to pectoral-fin base. A slightly frayed, fleshy membranous mid-ventral keel between urogenital opening and anal-fin origin. In males, a distinct mid-ventral groove, deepened and widened anteriorly, supports the priapium and mid-ventral keel. Pectoral fin falcate, the uppermost branched ray longest in most specimens; pectoral-fin rays branched, except for the uppermost 1 (uppermost nubby ossicle not included here; see “Materials and Methods” above) and lowermost 1–2 rays unbranched. Pelvic fin absent in males, present but rudimentary in females (Fig. 3). First dorsal fin absent; origin of second dorsal fin at, or slightly before, a level of posterior end of anal-fin base; anterior 2–3 rays simple, whereas the other rays branched. Anal fin with a long base, commencing well before midlength; anterior 2–4 rays simple, whereas the other rays branched. Caudal fin emarginate, symmetrical dorsoventrally. Male bilaterally asymmetric, dextral; namely, seminal papilla offset to right side of body (= aproctal side), and anus offset to left side of body (= proctal side); a long rod-like toxactinium curved from left to right; a large fleshy pad, the pulvinulus, covers articulation point of toxactinium and aproctal axial bone (Fig. 3). Scales on body cycloid, moderately large and deciduous; scales on abdomen largest; body entirely scaled, except for pectoral-fin base, mid-ventral groove before anal-fin origin, and mid-predorsal narrow naked space slightly behind occipital region; head and fins naked, except for posterior part of occipital region and basal part of caudal fin with some scales. Teeth on premaxilla and dentary unicuspid, slightly curved inward. Paradentary slender (as in Phallostethus dunckeri illustrated by Parenti, 1984: 4, fig. 1), with 5–19 minute teeth laterally; teeth on paradentary form a uniserial row or, in some larger specimens, biserial rows; teeth on inner row, if present, much smaller than those on outer row. Cephalic sensory canals reduced, comprising: two short infraorbital canals (each with terminal pores only) anteriorly and anteroventrally to eye; preopercular canal (with 6–7 pores). Main external bones in males including a long, curved toxactinium and a short stout ctenactinium with seven serrae dorsally (not including a hook-like distal tip); two smallest males examined (CTU-P 2493 and USNM 404478, 20.0– 20.3 mm SL) bearing 5–6 serrae on ctenactinium, assumued to represent the immature condition (and not included in the diagnosis, above). First pleural rib attached to fifth vertebra in males, fourth in females; first pleural rib in female much shorter than in male. Branchiostegal rays 4. Color when alive or freshly collected. Body subtranslucent in life, but whitish immediately after death (Fig. 1); a bright white blotch over brain when alive (assumed to fade just after death); iris silvery; minute melanophores scattered on snout, cheek and jaws; a melanophore at angle of lower jaw; a large reddish yellow blotch, slightly smaller than eye, at mid-lateral caudal fin base; male priapium with several large and small melanophores, particularly on the aproctal side (= right side in the new species) just anterior to the base of second ctenactinium; a series of minute black dots along mid-lateral septum of body musculature at least on caudal part of body; inner side of pectoral fin with many melanophores at least dorsally (the area with melanophores much more broader in females than in males); a series of mid-ventral black dots from anal-fin origin to caudal-fin base; other fins transparent. Color in alcohol. Head and body pale straw-colored; a series of irregular-sized melanophores (several of them dash-like) along midlateral septum of body musculature (at least on caudal part); paired patches of melanophores on anterior part of snout dorsally; many melanophores scattered on head above neurocranium, those posterior distinctly larger than those anterior; a melanophore at angle of lower jaw; a patch of minute gular melanophores; two patches of melanophores at throat and just behind urogenital opening in females; male priapium with several large and small melanophores, particularly on the aproctal side (= right side in this species) just anterior to the base of second ctenactinium; a mid-ventral series of black dots from anal- to caudal-fin bases, each along anal-fin base on interspace between fin rays (continuous and forming a irregular blackish gray line in some specimens); inner side of pectoral fin with many melanophores dorsally (the area with melanophores much broader in females than in males); caudal fin covered by numerous minute melanophores; other fins transparent. Distribution, habitat and the other notes. Phallostethus cuulong is known from nine specimens, six males and three females, collected from shallow waters (<1.2m depth) around banks of slow-flowing turbid canals and rivers with soft muddy bottoms in Soc Trang and Tra Vinh Provinces, Vietnam. The first author (KS) observed that a fish, latter designated as one of the paratypes of Phallostethus cuulong (NSMT-P 106665), swam slowly at the water surface around a bank of the slow-flowing tidal canal with dense semi-aquatic vegetation. A bright white blotch on dorsal surface of head was clearly confirmed in the field, but less vivid than that of the sympatric aplocheilid, Aplocheilus panchax (well-known for its reflective “pineal” spot on the top of the head). When the fish was disturbed, it quickly swam a short distance away from the original position; it was subsequently scooped up carefully using a hand net by KS. The species was usually solitary, and collected by hand nets or seine nets. Like the other phallostethids in the Vietnamese Mekong, this species has never been seen in the fish markets. As far as we aware, all fishes of the family Phallostethidae have no vernacular names in the Vietnamese Mekong (except for the new species herein named), since they are usually overlooked. Etymology. The specific name, cuulong, is the Vietnamese name of the Mekong delta (Cưu Long), where the type series of the new species was collected. The name, here applied as a noun in apposition, means “nine dragons,” in reference to nine distributaries of the Mekong basin in Vietnam. Remarks. Following the key to genera of phallostethid fishes by Parenti (1989), the new species is clearly assigned to Phallostethus by having the combination of, e.g., shield-like pulvinulus, large seminal papilla, long toxactinium, membranous dome on dorsal surface of head, 24–27 anal-fin rays, 37–40 vertebrae, serrated ctenactinium, non-projecting lower jaw beyond upper jaw, no first dorsal fin, and 7–9 second dorsal-fin rays. In particular, no other phallostethid genera are known that bear 24 or more anal-fin rays (vs. 22 or less anal-fin rays in the other phallostethids). Within the genus, the new species resembles the Bornean species Phallostethus lehi in sharing 11–13 pectoral-fin rays, but differs in having seven serrae on second ctenactinium in adult (vs. eight in P. l e h i), 25–26 caudal vertebrae (vs. 28), and 6–19 paradentary teeth (vs. 28 or more). All six examined males of the new species are dextral, immediately distinguishing them from sinistral males in P. l e h i. The new species is also distinguished from Phallostethus dunckeri, known only from Johor, Malay Peninsula but presumed to be extinct (Parenti, 1996), by having seven serrae on second ctenactinium in adults (vs. five in P. d u n c k e r i), 11–13 pectoral-fin rays (vs. 9–10), and 25–26 caudal vertebrae (vs. 27). Sexual dimorphism in the pleural ribs was reported from Phallostethus lehi by Parenti (1996); according to her, the species has the first pair of pleural ribs on the fifth vertebrae in males, the fourth vertebrae in females. This dimorphism is also found in Phallostethus cuulong. Furthermore, P. cuulong appears to show sexual dimorphism in the number of precaudal vertebrae: all six males examined have 13–14 precaudal vertebrae, as against 11–12 in all three females examined. Although Parenti & Louie (1998) reported similar sexual dimorphism in vertebral counts from four species of Neostethus, hitherto it has never been known from the other species of Phallostethus.Published as part of Shibukawa, Koichi, Tran, Dinh Dac & Tran, Loi Xuan, 2012, Phallostethus cuulong, a new species of priapiumfish (Actinopterygii: Atheriniformes: Phallostethidae) from the Vietnamese Mekong, pp. 45-51 in Zootaxa 3363 on pages 46-51, DOI: 10.5281/zenodo.28164
Micronecta (Micronecta) drepani Nieser 2000
No full textMicronecta (Micronecta) drepani Nieser, 2000 Figs 5C, 7A–C Micronecta drepani Nieser, 2000: 278–281, 287, figs 1–10 (type locality: northern Thailand). Micronecta drepani – Nieser et al. 2005: 190, 191 (notes, checklist). — Zettel et al. 2017: 40 (record Cambodia). Diagnosis Body length 1.9–2.1. Pronotum slightly longer than median head length. Hemelytron punctate, corium with patchy, indistinct brown markings. Males: fore femur with a pair of spines on proximal third, two spines distally; fore tibia with three spines on distal third ventrally; palar claw large, sub-triangular with straight apical margin. Median lobe of sternite VII short and broad with four long setae and an angular apex. Free lobe slightly expanded posteriorly, mesial angle rounded, lateral angle produced with ca 20 long setae (Fig. 7A). Left paramere: shaft broad, flattened, basal part of shaft with a short ridge running to lateral margin, apex rounded; basal lobe long, tongue-shaped (Fig. 7C). Right paramere: shaft evenly curved, parallel sided, apex rounded (Fig. 7B). Material examined VIETNAM – Ha Giang Prov. • 1 ♂, 2 ♀♀; Vi Xuyen, Xin Chai commune, Da stream by the road from Thanh Thuy to Lao Chai; 9 May 2014; A.D. Tran et al. leg.; TAD1405; ZVNU • 1 ♂; Xin Man, road 178, tributary of Nam Yen River, near bridge ca 2 km from Coc Pai town; 11 May 2014; A.D. Tran et al. leg.; TAD1412; ZVNU • 1 ♂, 1 ♀; Xin Man, Nam Dan commune, Thac Tien-Deo Gio, waterfall and stream; 11 May 2014; A.D. Tran et al. leg.; TAD1414; ZVNU. – Cao Bang Prov. • 12 ♂♂, 8 ♀♀; Phia Oac- Phia Den N.P., road to Quang Thanh commune, Khue Lan stream; 29 Oct. 2020; V.V. Nguyen et al. leg.; PO20-07; ZVNU. – Bac Kan Prov. • 8 ♂♂, 9 ♀♀; Ba Be, Hoang Tri commune, Hoang Tri waterfall; 22 Aug. 2012; A.D. Tran et al. leg.; TAD1218; ZVNU. – Lang Son Prov. • 3 ♂♂, 9 ♀♀; Loc Binh, Mau Son, Khuoi Tang stream; 590 m a.s.l.; 6 Nov. 2020; A.D. Tran et al. leg.; TAD20-34; ZVNU. – Phu Tho Prov. • 2 ♀♀; Xuan Son N.P., Ban Coi, Coi stream, site 3, by the road to Tan Son; 7 Jun. 2013; A.D. Tran et al. leg.; TAD1339; ZVNU • 3 ♂♂, 2 ♀♀; Xuan Son N.P., Dong Son, Than stream, site 2; 29 Aug. 2013; A.D. Tran et al. leg.; TAD1347; ZVNU • 3 ♂♂, 10 ♀♀; Xuan Son N.P., Xoan stream, near its confluence with Ha Bang stream; 21°5.303′ N, 104°57.559′ E; 27 Aug. 2014; T.S. Nguyen leg.; TS1407; ZVNU • 4 ♂♂, 4 ♀♀; Xuan Son N.P., Lap stream, the second concrete bridge from Ngoc waterfall; 21°08.322′ N, 104°56.256′ E; 28 Aug. 2010; T.S. Nguyen leg.; TS1413; ZVNU • 8 ♂♂, 5 ♀♀; Xuan Son N.P., Than stream, site 2; 21°11.257′ N, 104°52.382′ E; 28 Aug. 2014; T.S. Nguyen leg.; TS1415; ZVNU • 29 ♂♂, 32 ♀♀; Xuan Son N.P., Than stream, site 2; 21°11.257′ N, 104°52.382′ E; 24 May 2015; T.S. Nguyen leg.; TS1515; ZVNU • 2 ♂♂, 1 ♀; Xuan Son N.P., Than stream, site 2; 21°11.257′ N, 104°52.382′ E; 8 May 2016; T.S. Nguyen leg.; TS1615; ZVNU • 3 ♂♂, 7 ♀♀; Xuan Son N.P., Coi stream, site 2, near concrete bridge; 21°09.130′ N, 104°56.761′ E; 29 Apr. 2014; T.S. Nguyen leg.; TS1418; ZVNU • 2 ♂♂, 1 ♀; Xuan Son N.P., Coi stream, site 2, near concrete bridge; 21°09.130′ N, 104°56.761′ E; 24 May 2015; T.S. Nguyen leg.; TS1518; ZVNU • 9 ♂♂, 13 ♀♀; Xuan Son N.P., Xoan stream, Xoan village area; 21°05.270′ N, 104°57.591′ E; 23 May 2015; T.S. Nguyen leg.; TS1509; ZVNU • 1 ♂, 8 ♀♀; Xuan Son N.P., Xoan stream, Xoan village area; 21°05.270′ N, 104°57.591′ E; 26 Aug. 2015; T.S. Nguyen leg.; TS8509; ZVNU • 4 ♂♂, 4 ♀♀; Xuan Son N.P., Xoan stream, Xoan village area; 21°05.270′ N, 104°57.591′ E; 7 May 2016; T.S. Nguyen leg.; TS1609; ZVNU • 3 ♂♂, 7 ♀♀; Xuan Son N.P., Than stream, site 3; 21°11.393′ N, 104°52.625′ E; 27 Aug. 2015; T.S. Nguyen leg.; TS8516; ZVNU • 5 ♂♂, 1 ♀; Xuan Son N.P., Lap stream, the first concrete bridge from Ngoc waterfall; 21°8.398′ N, 104°56.303′ E; 7 May 2016; T.S. Nguyen leg.; TS1612; ZVNU. – Hanoi • 5 ♂♂, 4 ♀♀; Ba Vi N.P., Minh Quang, Cai stream; 65 m a.s.l.; 17 Dec. 2011; Q.H. Ngo et al. leg.; TAD1126; ZVNU • 2 ♂♂, 1 ♀; Ba Vi N.P., Khoang Xanh, Tien stream; 95 m a.s.l.; 17 Dec. 2011; Q.H. Ngo and A.D. Tran leg.; TAD1127; ZVNU • 4 ♂♂, 4 ♀♀; Ba Vi N.P., Minh Quang, Cai stream; 65 m a.s.l.; 26 Apr. 2012; Q.H. Ngo et al. leg.; TAD1207; ZVNU • 1 ♂; Ba Vi N.P., Khoang Xanh, Tien stream; 95 m a.s.l.; 27 Apr. 2012; Q.H. Ngo and A.D. Tran leg.; TAD1208; ZVNU. – Vinh Phuc Prov. • 2 ♂♂, 1 ♀; Tam Dao N.P., Suoi Mo (upstream of Thac Bac waterfall); 16 Jun. 2003; A.D. Tran leg.; TAD0350; ZRC • 5 ♂♂, 5 ♀♀; Tam Dao N.P., Suoi Mo; 9 and 15 Jul. 2019; A.D. Tran leg.; ZVNU. – Quang Binh Prov. • 2 ♀♀; Phong Nha, Chay stream; 17 Jul. 2004; A.D. Tran leg.; DY0410; ZRC. – Quang Nam Prov. • 1 ♂, 2 ♀♀; Duy Xuyen District, My Son temple complex; 15°45.439′ N, 108°07.577′ E; Aug. 2016; T.D. Chu leg.; MS1614; ZVNU • 1 ♀; Duy Xuyen District, My Son temple complex; 15°45.482′ N, 108°07.606′ E; Apr. 2017; T.D. Chu leg.; MS1715; ZVNU. – Lam Dong Prov. • 7 ♂♂, 4 ♀♀; Bao Loc, Damb’ri waterfall; 19 May 2003; A.D. Tran leg.; TAD0326; ZRC. Distribution China (Yunnan), Thailand (Nieser et al. 2005), Cambodia (Zettel et al. 2017). First records for Vietnam. Remarks See Remarks under M. cultellata sp. nov.Published as part of Ha, Tuyet Ngan & Tran, Anh Duc, 2021, Taxonomy of Micronectidae (Heteroptera: Nepomorpha) from Vietnam, with descriptions of 11 new species, pp. 1-82 in European Journal of Taxonomy 756 on pages 28-29, DOI: 10.5852/ejt.2021.756.1407, http://zenodo.org/record/503301
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