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    Notes on the Bembidiina of Taiwan with description of three new species (Coleoptera: Carabidae)

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    Toledano, Luca (2009): Notes on the Bembidiina of Taiwan with description of three new species (Coleoptera: Carabidae). Acta Entomologica Musei Nationalis Pragae 49 (2): 577-598, DOI: 10.5281/zenodo.532056

    Bembidion (Plataphodes) kmecoi sp. nov. from China, Sichuan (Coleoptera: Carabidae: Bembidiina)

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    Toledano, Luca, Nakládal, Oto (2011): Bembidion (Plataphodes) kmecoi sp. nov. from China, Sichuan (Coleoptera: Carabidae: Bembidiina). Zootaxa 2882: 64-68, DOI: 10.5281/zenodo.27758

    Figs. 10–12 in Notes on the Bembidiina of Taiwan with description of three new species (Coleoptera: Carabidae)

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    Figs. 10–12. Bembidion spp., head and pronotum. 10 – B. (Trichoplataphus) taiwanum Netolitzky, 1939; 11 – B. (Trichoplataphus) jelineki sp. nov., holotype; 12 – B. (Trichoplataphus) seijii sp. nov., holotype.Published as part of Toledano, Luca, 2009, Notes on the Bembidiina of Taiwan with description of three new species (Coleoptera: Carabidae), pp. 577-598 in Acta Entomologica Musei Nationalis Pragae 49 (2) on page 587, DOI: 10.5281/zenodo.532056

    FIGURES 3–4 in Bembidion (Plataphodes) kmecoi sp. nov. from China, Sichuan (Coleoptera: Carabidae: Bembidiina)

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    FIGURES 3–4. Median lobe of the aedeagus in left lateral view of: 3) B. farkaci (holotype) and 4) B. kmecoi sp. nov. (holotype), scale = 0.5 mm.Published as part of Toledano, Luca & Nakládal, Oto, 2011, Bembidion (Plataphodes) kmecoi sp. nov. from China, Sichuan (Coleoptera: Carabidae: Bembidiina), pp. 64-68 in Zootaxa 2882 on page 67, DOI: 10.5281/zenodo.27758

    Plataphodes

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    Key to the Chinese species of Plataphodes 1 Eyes very large, SW China, Sichuan...................................................................... 2 - Eyes normal................................................................ B. fellmanni Mannerheim, 1823 2 Two discal elytral pores..................................................... B. farkaci Toledano & Sciaky, 1998 - Three discal elytral pores................................................................. B. kmecoi sp. nov.Published as part of Toledano, Luca & Nakládal, Oto, 2011, Bembidion (Plataphodes) kmecoi sp. nov. from China, Sichuan (Coleoptera: Carabidae: Bembidiina), pp. 64-68 in Zootaxa 2882 on page 65, DOI: 10.5281/zenodo.27758

    Bembidion (Plataphodes) farkaci Toledano & Sciaky 1998

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    Bembidion (Plataphodes) farkaci Toledano & Sciaky, 1998 (Figs. 1, 3) Examined material. Holotype, ♂, " China, Sichuan, Gongga Shan, Hailuogou, btw. Camp 2 & 3, 2500–2900 m, 29 ° 35 N 102 °00E, 5.VII. 1996 " (CFKP). 1 paratype, ♀, " China, Sichuan, Gongga Shan, Hailuogou, above Camp 3, 3050 m, 29 ° 35 N 102 °00E, 8.VII. 1996 " (NHMB).Published as part of Toledano, Luca & Nakládal, Oto, 2011, Bembidion (Plataphodes) kmecoi sp. nov. from China, Sichuan (Coleoptera: Carabidae: Bembidiina), pp. 64-68 in Zootaxa 2882 on page 65, DOI: 10.5281/zenodo.27758

    FIGURE 5. Male aedeagus and right paramere. Scale bar 0.1 in Phylogenetic relationships of the South American ground beetle subgenus Chilioperyphus Jeannel (Coleoptera: Carabidae: Trechinae: Bembidiini: Bembidion Latreille)

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    FIGURE 5. Male aedeagus and right paramere. Scale bar 0.1 mm. A: B. orregoi (Argentina: Chubut: Rio Azul at Lago Puelo, 200m, 42.0933°S 71.6221°W; Maddison voucher V100667. B: B. mendocinum (Argentina: Mendoza: Villa 25 de Mayo, 875m, 34.5901°S 68.5608°W; Maddison voucher V100668).Published as part of Maddison, David R., Toledano, Luca, Sallenave, Soledad & Roig-Juñent, Sergio, 2013, Phylogenetic relationships of the South American ground beetle subgenus Chilioperyphus Jeannel (Coleoptera: Carabidae: Trechinae: Bembidiini: Bembidion Latreille), pp. 547-560 in Zootaxa 3636 (4) on page 557, DOI: 10.11646/zootaxa.3636.4.3, http://zenodo.org/record/526246

    Bembidion (Trichoplataphus) miwai Jedlicka 1946

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    Bembidion (Trichoplataphus) miwai Jedlička, 1946 (Figs. 6, 21) Bembidion (Trichoplataphus) miwai Jedlička, 1946: 2. Bembidion (Trichoplataphus) miwai: JEDLIČKA (1965): 116; MARGGI et al. (2003): 271; LORENZ (1998): 211; LORENZ (2005): 227. Bembidion miwai: TERADA et al. (2005):170; TERADA (2006): 12. Type material examined. HOLOTYPE: J, ‘ TAIWAN / Raisya / 24-XI-1934 / S. Issiki // TYPUS (red) // Mus. Nat. Pragae / 23964 Inv. (orange) // miwai / sp.n. [handwritten] / DET. ING. JEDLIČKA [pink]’ (NMPC). Additional material examined. TAIWAN: Near Mt. Ari, Takesari, 24.xi.1937, Y. Yano lgt. (‘ Yano 7.20, miwai Jedl. Typo collatum det. Netolitzky [handwritten]’), 1 J (NHMW); Kungliao, Taipei Hsien, 18.ii.2002, Terada lgt., 1 J (CTVR); Kao Hsiung, Co., 10–11 km NE Chiahsen, ca. 300 m, 3.–8.vii.1980, forest, D. R. Davis lgt., 2 ♀♀ (USNM, CTVR). Intraspecific variability. The type specimen and the specimen from NHMW (Fig. 6) have lighter colours (entire body light reddish) in comparison to the other specimens listed here. The specimen from Kungliao is darker and has less convex and smaller eyes, but it seems conspecific in all the other characters; Fig. 21 shows the male genitalia of this specimen because the type specimen is immature and the other examined specimen (NHMW) has the endophallus everted. Distribution. Taiwan (Raisya, Takesari, Taipei Hsien, Kao Hsiung).Published as part of Toledano, Luca, 2009, Notes on the Bembidiina of Taiwan with description of three new species (Coleoptera: Carabidae), pp. 577-598 in Acta Entomologica Musei Nationalis Pragae 49 (2) on page 585, DOI: 10.5281/zenodo.532056

    Bembidion

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    Bembidion subg. Plataphodes Ganglbauer, 1891 Systematic notes. The Holarctic subg. Plataphodes Ganglbauer, 1891 belongs to the “ Plataphus complex” (Toledano 2008). It includes 37 species, of which 18 Palaearctic, 3 Holarctic and 16 Nearctic. A revision of the subgenus is still pending in order to make clear the systematic relationships of the species. The synapomorphy shared by all the species is the basal elytral margin reaching the apical end of stria 5 and then extended with a transverse, straight prolongation reaching the apical end of stria 4. In the species of the subg. Plataphus Motschulsky, 1864, probably the subgenus more strictly related to Plataphodes, the basal elytral margin ends at the beginning of stria 5, without this transverse prolongation. Otherwise, for remaining external characters both subgenera are very similar to one another. We were able to examine the type of B. elatum Andrewes, 1930 (BMNH), formerly attributed to Plataphodes: actually the basal end of the lateral margin of B. elatum exactly reaches the beginning of stria 5 without prolongation to base of stria 4. Furthermore this species belongs to the group of subgenera with supernumerary abdominal setae (Toledano & Schmidt 2010), and more precisely to Blepharoplataphus Netolitzky, 1920 because it shows supernumerary, short setae in a single row on each of the last abdominal sternites, including the last one (which in Trichoplataphus Netolitzky, 1914 is almost entirely covered by setae).Published as part of Toledano, Luca & Nakládal, Oto, 2011, Bembidion (Plataphodes) kmecoi sp. nov. from China, Sichuan (Coleoptera: Carabidae: Bembidiina), pp. 64-68 in Zootaxa 2882 on page 65, DOI: 10.5281/zenodo.27758

    Bembidion (Blepharoplataphus) teradai Toledano 2009, sp. nov.

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    Bembidion (Blepharoplataphus) teradai sp. nov. (Figs. 5, 20) Type locality. Taiwan, Nansichi, Wulai, Taipei Hsien. Type material. HOLOTYPE: J, ‘ TAIWAN, 23-xii-2001 / Nansichi, Wulai / Taipei Hsien / Terada – 77’ // HOLOTYPUS / Bembidion teradai sp.n. / Det. L. Toledano, 2009 [red]’ (MSNV). PARATYPES: 6 JJ 9 ♀♀, same date and locality as the holotype (NTUC, CTHJ, CTVR); 2 JJ 6 ♀♀, ‘ Wulai / Taipei Hsien / Taiwan // 4-XII-2001 / K. Terada leg.’ (CMTJ); 1 J 1 ♀, ‘ Wulai Taiwan / 23-XII-2001 / K. Terada leg.’ (CMTJ, CTVR). All paratypes labelled as ‘ PARATYPUS / Bembidion teradai sp.n. / Det. L. Toledano, 2009 [red]’. Diagnosis. A Blepharoplataphus species with very wide elytra and humeri, very similar to Bembidion nirasawai Morita, 2008 from southwestern Japan, but distinguishable from this last for the apex of the aedeagus, rounded in B. nirasawai (MORITA 2008) and ventrally hooked in B. teradai. Description. Body length 5.38–5.84 mm. Black with faint metallic greenish reflections. Legs and antennae piceous black (Fig. 5). Head with deep, wide and parallel frontal furrows, not extending on clypeus. Eyes convex, antennae rather short (bl/al = 1.98–2.09). Pronotum transverse (pw/pl = 1.42–1.48), rather narrow in relation to elytra (ew/pw = 1.75–1.76). Sides rounded, briefly sinuate before the short, slightly obtuse hind angles. Lateral channel very wide, anterior transverse impression short, in some specimens with some punctures, median line superficial, basal foveae somewhat square, very rugose, laterally delimited by a carina. Posterior margin straight in middle and oblique at sides. Elytra (el/ew = 1.40–1.43) oval, very wide and convex, with humeri wide and rather rounded. Elytral striae 1–7 complete, visibly impressed, punctate-sulcate, scutellar stria long, apical stria long and connected with apical end of stria 5. Two discal elytral pores in interval 3 near stria 3. Microsculpture distinct, consisting of small isodiametric, convex cells on head and pronotum and of transverse, rather irregular flat cells on elytra. Male genitalia (Fig. 20). Apex of median lobe ventrally hooked. For the other characters, typical shape and endophallus of the subgenus Blepharoplataphus. In particular, the endophallus shares the structure of the sclerites with B. nirasawai. Etymology. Dedicated to Katsuyuki Terada, an accomplished specialist in Japanese and Taiwanese Carabidae, who gave me the opportunity to study the interesting fauna of Taiwan by kindly sending me the specimens dealt with in this paper. Distribution. So far known only from the type locality. Remarks. The morphological diversity within Blepharoplataphus is very low: the male genitalia and habitus of all species are very similar; B. teradai sp. nov. seems closely related to B. nirasawai, with which it shares almost identical male genitalia, except for the apical shape.Published as part of Toledano, Luca, 2009, Notes on the Bembidiina of Taiwan with description of three new species (Coleoptera: Carabidae), pp. 577-598 in Acta Entomologica Musei Nationalis Pragae 49 (2) on pages 582-585, DOI: 10.5281/zenodo.532056
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