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    Metandania Stephensen 1925

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    <i>Metandania</i> Stephensen, 1925 <p> <i>Metandania</i> Stephensen, 1925: 136.</p> <p> <i>Metandania</i>: Schellenberg, 1953: 187.</p> <p> <i>Andaniotes</i>: Barnard, 1969: 441 (new synonymy).</p> <p> <i>Metandania</i>: Watling and Holman, 1981: 219.</p> <p>Monotypi c genus.</p> <p> <i>Remarks</i></p> <p> The present genus is very distinct from <i>Andaniotes</i>, with which Barnard (1969) synonymized it (for details, see <i>Remarks</i> under <i>M. islandica</i>). However, its phylogenetical relationships with the other genera in the Stegocephalidae will not be further discussed herein, as a cladistic analysis of the entire family will be published separately (Berge and Vader, in press).</p>Published as part of <i>Berge, JØrgen, 2001, Revision of the Amphipod (Crustacea: Stegocephalidae) genera Andaniotes Stebbing, 1897 and Metandania Stephensen, 1925, pp. 787-832 in Journal of Natural History 35 (6)</i> on page 825, DOI: 10.1080/00222930152123620, <a href="http://zenodo.org/record/5277015">http://zenodo.org/record/5277015</a&gt

    Grandidierella bonnieroides Stephensen 1948

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    Grandidierella bonnieroides Stephensen, 1948 Grandidierella bonnieroides Stephensen, 1948: 12, fig. 3.— Myers, 1970: 141.— Myers, 1981 b: 218.— Asari & Myers, 1982: 252, figs 9–10.— Lowry & Stoddart, 2003: 70 (catalogue). Grandidierella megnae.— Chilton, 1921: 548, fig. 10.— Stephensen, 1933: 434.— Shoemaker, 1935: 70. Unciolella lunata.— Schellenberg, 1928: 669, fig. 207. Grandidierella bonnieri.— K.H. Barnard, 1935: 299.— K.H. Barnard, 1951: 708.— K.H. Barnard, 1952: 279, fig. 1.—Pannikar & Aiyar, 1937: 294.—Schellenberg, a: 215.— Shoemaker, 1948: 11, fig. 3.— Ruffo, 1958: 58, figs 8, 9.— Nayar, 1959: 38, pl. 14, figs 1–5.— Nayar, 1966: 161, fig. 17 f. — Myers, 2009: 271, figs 39–40, pl. 2 C. Material. 5 males, 4 females (NHMUK 2015. 3099–3107), llBS, grab on bare sand benthos in phase A 1, M. Nithyanandan, 25 May 2014; 1 male, 7 females (NHMUK 2015. 3108–3115), tunicate samples taken by diving in Marina in phase A 2, D.K. Raja, 1 April 2014. Remarks. This apparently worldwide species may be a species complex.Published as part of Myers, Alan A. & Nithyanandan, Manickam, 2016, The Amphipoda of Sea City, Kuwait. — The Senticaudata (Crustacea), pp. 401-429 in Zootaxa 4072 (4) on page 408, DOI: 10.11646/zootaxa.4072.4.1, http://zenodo.org/record/26675

    Metopa abyssalis Stephensen 1931

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    Metopa abyssalis Stephensen, 1931 Fig. 17–18. Metopa abyssalis Stephensen, 1931 Material examined. Morphological examination: ZMUC-CRU 5019. Male. Ingolf Station 101 (7?). 66 23 ’ N, 12 05’ W. 1011m. Temp. -0.7 C. Ingolf Station 138. 63 26 ’ N, 7 56 ’ W. 887m. -0.6 C. (for comparison only, not dissected.) Morphological redescription of type material. This description is based on the same slides that Stephensen (1931) presumably used when he created the species, they are at least made by Stephensen, have his handwriting, and are of the same specimens that Stephensen based his description on. They do, however, miss several of the pieces normally used for a description. There are no other specimens of this species present in the Copenhagen collections than the two samples from Stephensen. Due to the very small sample size of the collections, no further specimens were dissected. Stephensen presumed the slides were of a male; we have found nothing to contradict this. Stephensens slides from station 138 were not found in the collections, these have been of an adult female. Head (Fig. 17): epistome reduced and rounded, lateral cephalic lobes little prominent, rounded; eye small, indistinct, only consisting of a few scattered ommatidia. Antenna 1 (Fig. 17): slightly shorter than antenna 2; basis of peduncle ad broad as half head, peduncle as long as half flagellum; flagellum 10 -articulate, few short simple setae on the distal articles. Antenna 2 (Fig. 17): peduncle articles 1–3 short; articles 4 and 5 each longer than articles 1–3 together; flagellum 8 -articulate, shorter than peduncle, 2 short simple setae distally on the last 6 articles. Mandible (Fig. 17): palp 3 -articulate, cross-section circular, article 2 3 x as long as article 1, with one simple seta, article 3 as short as article 1, one simple seta at apex; incisor coarsely serrate, lacinia mobilis more finely serrate, no raker setae on the mandible examined, no molar. Maxilla 1 (Fig. 17): inner plate with one small conical seta; outer plate with one largish cuspidate seta, two simple setae and one serrate seta; palp 1 -articulate, distal margin serrate with one short simple seta in each serration, two longer simple setae and one type A seta at distal tip. Maxilla 2 (Fig. 17): outer plate in normal position to inner plate; inner plate with one short and two long simple setae; outer plate with one short and strong seta and four long and simple setae. Maxilliped (Fig. 17): inner plate fused with a ridge along the midline, a row of short strong setae at distal margin; outer plate very small; palp 4 -articulate, simple setae along inner margin of palp and outer plate; article 3 of palp with a cushion of flat serrate setae and two type A setae at distal end; dactylus (article 4) with a line of short flat serrate setae on inner margin, slightly curved. Pereon: Smooth. Gnathopod 1 (Fig. 18): simple to vaguely subchelate; merus with a patch/cushion of several short, flat slightly serrate setae at posterior margin, four type A setae at posterodistal corner; carpus as long as propodus, suboval with long type A setae at posterior margin and anterodistal corner, propodus oval with two short simple setae where dactylus folds in, the palm recognizable from the setation, as long as posterior margin; dactylus curved with a small tooth-like projection at tip, short simple setae along inner margin, and one long simple seta at outer margin. Gnathopod 2 - pereopod 4: no data from slide, hidden under coxa 2–4. Pereopod 5: basis slender. Pereopod 6 and 7: basis posteriorly expanded. Urosome: smooth. Uropod 1 (Fig. 18): longer than uropod 2; peduncle slightly longer than rami, 3 short setae at inner margin; biramous; rami of equal length, one short and simple seta at inner margin of outer ramus. Uropod 2 (Fig. 18): slightly longer than uropod 3; two short and simple setae at inner margin; biramous, inner ramus as long as peduncle, outer ramus shorter than inner ramus. Uropod 3 (Fig. 18): peduncle approximately as long as ramus; ramus 2 -articulate, outer article slightly longer than inner, one short simple seta on inner article. Telson (Fig. 18): entire, sharp rounded tip; two short cuspidate setae at proximal half. Ecology: Very little is known about the ecology of this species. We only know it from the two samples in the collections in Copenhagen (see map D on Fig. 22), from 887 and 1011m depth, and subzero temperatures of the water. Geographic distribution of C: Metopa groenlandica and D: M. abyssalis. Other material. Stephensen (1931) used dissected material from both stations when he described M. abyssalis. The slide from station 138 was not found in the collections now, and is presumed lost. By agreement with the museum we did not dissect the last remaining specimen in the tube from station 138. We have not been able to find any other specimens of M. abyssalis in other museums collections. Remarks. Stephensen (1931) dissected out a gnathopod 2 from both a male and a female, they do not show any conspicuous sexual differences. Gnathopod 2 is subchelate, with a subovate coxa; slender basis and subquadrate ischium; merus short, with a few setae at posterior margin, carpus subtriangular, carpal lobe not free, and reaching 1 / 4 of posterior margin of propodus; propodus suboval, little more than double length of carpus, with a clear palm, oblique, but very clearly marked by a strong tooth at palmar corner, palm crenulate; dacylus smooth, as long as palm.Published as part of Tandberg, Anne Helene S. & Vader, Wim, 2009, A redescription of Metopa species (Amphipoda, Stenothoidae) based on the type material. 1. Zoological Museum, Copenhagen (ZMUC), pp. 1-36 in Zootaxa 2093 on pages 25-33, DOI: 10.5281/zenodo.18753

    Cystisoma latipes Stephensen

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    Cystisoma latipes (Stephensen) (Figs 10­13) Thaumatops latipes Stephensen, 1918: 62 ­63, fig. 21. – Schellenberg 1927: 621, fig. 29. Cystisoma latipes – Pirlot 1938: 377, 379, 380, 384, 386. Pirlot 1939: 34. Brusca 1973: 9 (Table), 14. Brusca 1981 a: 19 (key), 39, fig. 5 d. Brusca 1981 b: 358, fig. 12 c. Vinogradov et al. 1982: 254 ­255, fig. 125. ? Cystisoma parkinsonii Stebbing, 1888: 1332 ­1333. ? Thaumatops parkinsonii – Stephensen 1918: 66 ­68, figs 28, 29. Schellenberg 1927: 623, fig. 32. ? Thaumatops longipes [misidentification] – Bovallius 1887 b: 558. Bovallius 1889: 47 ­52 (spec. B, non A). Type material The unique type of C. latipes, a male measuring 44 mm, is in the ZMUC (CRU 2828). The type locality is the North Atlantic Ocean, south of Ireland, 49 ° 22 ’N, 12 ° 52 ’W (Thor Stn. 81), 1350 m. Type material of synonyms The unique type of C. parkinsonii, a male measuring about 50 mm, is in the BMNH (1889.5.15.198). Stebbing (1888) originally described this species, referring to it as “ Cystisoma. Specimen F”, and proposed the name C. parkinsonii “if, in view of the great length of the upper antennae, it be necessary to separate this specimen from the others”. The specimen is very similar to C. latipes, differing in having slightly shorter first antennae, mandibles with only a medial tooth, and the basis of pereopod 7 slightly narrower but still relatively inflated proximally. Despite these differences, it appears to be the same as C. latipes, although the length of the first antennae could not be determined accurately, because the specimen is in pieces. Amongst the material examined there are other specimens of C. latipes with mandibles without lateral teeth and the basis of pereopod 7 relatively inflated proximally, compared to other species. Material examined (56 specimens) Types. Holotype male of C. latipes, approx. 44 mm: in spirit. Holotype male of C. parkinsonii, approx. 50 mm, specimen “F”, Challenger Stn. 196, north of Amboina, 0° 48 ’ 30 ”S, 126 ° 58 ’ 30 ”E, trawled 825 fathoms, 13 th October, 1874: 3 microscope slides of head, mouthparts, Us, A 1, G 1 & 2 and P 3­7 from right; remainder in spirit. Other material examined. North Atlantic: 8 lots (ZMUC), 1 lot (USNM), 9 specimens. South Atlantic: 1 lot (SAM), 2 lots (ZMUC), 3 specimens. North Indian: 1 lot (ZMUC), 1 specimen. South Indian: 1 lot (SAM), 10 lots (ZMUC), 12 specimens. Central Indo ­Pacific: 2 lots (ZMUC), 2 specimens. North Pacific: 2 lots (CAS), 1 lot (LACM), 2 lots (USNM), 5 specimens. South Pacific: 2 lots (ZMUC), 2 specimens. Diagnosis (males only) Body length up to 50 mm. Head as long as depth posteriorly; as long as first 4.5 pereonites combined; wedge­shaped when viewed laterally, with slightly concave anterior margin when viewed dorsally. Marginal spines 14­15; anterior ventral spine sometimes smaller than glandular spine; oral spines 2­4. Eyes obliquely­oval, barely touching anteriorly, and widely separated posteriorly. First antennae as long as head and pereon combined, with one small terminal article, medial surface with dense brush of aesthestascs, slightly inflated proximally. Mandibles with one medial tooth and one lateral tooth (absent in some specimens). Pereopod 7 with pear­shaped basis, inflated proximally, maximum width about half­length, or 4 x width, near distal margin. Urosome (with uropods) slightly shorter than pleon. Uropod 1; exopod subequal in length to endopod, about 0.4 x length of peduncle. Uropod 3 similar to U 1, exopod length slightly more than half­length of peduncle. Female unknown. Remarks Unusual characters of this species are the relatively long first antennae with aesthestascs, and the wedge­shaped head. In all other species of Cystisoma the head is more oval and, except for C. gershwinae sp. nov., the first antennae are without a brush of aesthestascs medially. The eye shape is also unusual, and the eyes occupy less of the lateral part of the head than other species, suggesting that this species may inhabit deeper water than its congeners. According to water­colour drawings and colour notes made from living material by Stephensen, the specimens are colourless, except for articles 4 and 5 of pereopod four, articles 2­6 of pereopods 5­7 and the first uropods, all of which are pink/red at the margins. Some specimens examined have all the characters of C. latipes except that the mandibles lack lateral teeth (e.g. USNM 39002). In this respect they are similar to the specimen described as C. parkinsonii by Stebbing (1888) (Fig. 12). Thus, it would seem that the presence or absence of a lateral mandibular tooth is variable in C. latipes. As these specimens are slightly larger than the type this character is not size dependent. Amongst the material from the ZMUC, which was seen by Stephensen (1918), are three specimens with relatively long first antennae that are tentatively assigned to C. latipes. The first of these (ZMUC CRU 2838), described by Stephensen as Thaumatops bovalli (= C. longipes), is a male measuring 46 mm from Thor Stn. 73. This specimen differs from C. latipes in having slightly shorter first antennae without aesthestascs, and pereopod 7 with a narrower basis. The medial surface of the first antennae has a broad white band, which may represent developing aesthestascs. It is possible that this is a recently moulted specimen in which the aesthestascs have not yet developed (as found in other species of hyperiideans), and the basis of pereopod 7 has not differentiated. A second specimen (ZMUC CRU 2833), described by Stephensen (1918) as Thaumatops parkinsonii, is a juvenile male measuring 38 mm from Thor Stn. 72. It is very similar to the previous specimen but the mandibles lack a lateral tooth. The third specimen (ZMUC CRU 2834) is Bovallius’ (1887 b) “ type ” specimen B of C. longipes (Fig. 13), which Stephensen (1918) also refers to T. parkinsonii. This specimen, measuring 46 mm, seems to be a juvenile female with developing brood plates. Both Stephensen (1918) and Bovallius (1889) refer to it as a male, probably because pereonite 7 appears to have a pair of gonopores. However, the presence or absence of testes could not be determined as the specimen is in poor condition. It differs from C. latipes in that the mandibles lack lateral teeth and pereopod 7 has a narrow basis. In many respects it is similar to the type of C. parkinsonii, particularly in the generally more slender uropods and in the relatively longer exopods, which are clearly longer than the endopods. This may be a female character of this species. If this specimen is C. latipes then it is the only confirmed female specimen of the species. The type of C. parkinsonii was re­examined but the sex could not be confirmed from the remains. Distribution Previously known only from the North Atlantic. Now also known from scattered records from the warm waters of the Indian and Pacific Oceans. Cystisoma gershwinae sp . nov. (Figs 14 & 15) Material examined (14 specimens) Types. Holotype. Male, 43 mm (USNM cat. 266786). Off Queen Charlotte Islands, British Columbia, Canada, 51 ° 57 ’N, 131 °05’W, 0­360 m, Halibut Commission, University of Washington, M.H. Pettibone: in spirit. Paratypes. 1. Male, approx. 53 mm (urosome missing) (USNM cat. 266788). Baja California, Mexico, 39 °00’N, 121 °00’W to 29 ° 39 ’N, 121 °00’W, mid­water trawl 1140 fathoms, R. Wisner & Parby, 27 April 1951: in spirit. 2. Male, approx. 53 mm (SAMA C 5906). San Clemente Basin, California, 500­1000m, IKMT, A. Collins, December, 1997: in spirit. 3. Male, 40 mm (LACM CR 15000057). Locality data uncertain, given as “ 8346 ” but is possibly off southern California. 4. Male, 40 mm (ZMUC CRU 3728). Equatorial North Atlantic between Africa and South America, 12 ° 11 ’N, 32 ° 49 ’E (Dana Stn. 1165), 1000 MW, 9 September 1921: in spirit. 5. Male, 45 mm (ZMUC CRU 3729). Equatorial North Atlantic between Africa and South America, 10 ° 16 ’N, 40 ° 41 ’W (Dana Stn. 1166), 1000 MW, 11 September 1921: in spirit. Other material. Six males, 35­45 mm, in poor condition, collected by the Dana (ZMUC CRU 3730­3735) from the equatorial North Atlantic (Stn. 1163), Banda Sea (Stn. 3677), western North Pacific (Stn. 3751) and the eastern North Indian Ocean (Stns. 3902 & 3903). Also, two males in very poor condition, collected by the Galathea (ZMUC), from the Celebes Sea (Stn. 448) and the western tropical Indian Ocean (Stn. 241). Diagnosis (males only) Body length up to 53 mm. Head as long as depth; as long as first 5 pereonites combined; oval when viewed laterally, with straight anterior margin when viewed dorsally. Marginal spines 11; anterior ventral spine shorter than, or subequal to, glandular spine; oral spines absent. Eyes oval, barely separated medially. First antennae slightly longer than head and pereon combined, with one tiny terminal article, medial surface with dense brush of aesthestascs. Mandibles with one medial tooth, lateral teeth absent. Pereopod 7 with basis slightly inflated proximally. Urosome (with uropods) about as long as pleon. Uropod 1; exopod slightly shorter than endopod, slightly less than half­length of peduncle. Uropod 3 similar to U 1, exopod length almost half­length of peduncle. Female unknown. Description of holotype male Body length 43 mm. Head as long as deep; as long as first 5 pereonites combined; oval laterally, with relatively straight anterior margin when viewed dorsally. Marginal spines 11, the anterior one the largest. Anterior ventral spine slightly shorter than glandular spine. Oral spines absent. Eyes oval, barely separated medially. First antennae as long as head, pereon and first 1.5 pleonites combined, with tiny terminal article, medial surface with dense brush of aesthestascs. Mandibles with one medial tooth, lateral teeth absent. Gnathopod 1, relatively small, length slightly more than half G 2, about 0.2 x P 3; distinctly chelate; basis about as long as remaining articles combined. Gnathopod 2, similar to G 1, also relatively small, length about 0.4 x P 3; basis slightly longer than remaining articles combined. Pereopod 3, about half as long as P 5; basis to propodus with denticulate posterior margin, denticles on propodus very small; merus, carpus and proximal half of propodus with transverse rows of setae, mainly medially; basis about as long as merus and carpus combined; carpus slightly longer than merus; propodus slightly shorter than carpus but slightly longer than merus. Pereopod 4, as long as 1.5 x P 3 and 0.7 x P 5; ornamentation similar to P 3 but merus with less, and propodus with more, rows of setae; basis slightly shorter than merus and carpus combined; carpus length about 1.3 x merus, slightly shorter than propodus. Pereopod 5 is the longest pereopod; basis to propodus with denticulate anterior margin, denticles on propodus very small; basis to carpus also with row of denticles posteriorly; basis only slightly longer than carpus; merus length nearly 0.7 x basis; propodus slightly longer than basis. Pereopod 6, similar to P 5, length about 0.8 x P 5; basis slightly longer than carpus; merus length nearly 0.6 x basis; propodus slightly longer than carpus but shorter than basis. Pereopod 7; length slightly less than half P 5; basis to propodus with denticulate anterior margin, denticles on propodus smaller; basis and merus also with row of denticles posteriorly; basis pear­shaped, maximum width proximally almost 3 x width near distal margin, 0.4 x length; basis as long as merus and carpus combined; merus marginally longer than carpus; carpus with fringe of long setae along distal margin; propodus slightly longer than carpus, with transverse rows of setae medially and along distal margin; distal margin concave with anterior tooth; dactylus curved, closing against distal margin of propodus, slightly longer than width of propodus. Urosome (with uropods) subequal in length to pleon. Uropod 1; peduncle length about 3 x maximum width; exopod slightly shorter than endopod, slightly less than halflength of peduncle. Uropod 3; peduncle length about 3 x maximum width, about 0.8 x length of peduncle of U 1; exopod subequal in length to endopod, almost half­length of peduncle. Telson very small, U­shaped, about 0.3 x as long as third pleonite. Etymology Named for Lisa­ann Gershwin, Department of Integrative Biology, University of California, Berkeley, USA, for her continuous support and encouragement in my research of hyperiideans and for being responsible for locating paratype 2 of this unusual species. Remarks The paratypes are like the holotype, except for paratype 1, which has the urosome missing, and slightly shorter first antennae (as long as head and pereon combined), and anterior ventral spines slightly larger than the glandular spines. The first antennae of Paratype 4 are similarly relatively shorter than for the holotype. The non­type material exhibits the characteristic features of the holotype (viz. the oral and mandibular spines, long first antennae and pereopod 7 with an inflated basis) but have not been designated paratypes because of their poor condition. The distal margin of the propodus of pereopod 7 is similar in shape to that found in juvenile females of other species. However, the holotype, paratypes and the other specimens are clearly males, as they have distinct testes, and, or lack the developing brood plates usually found in females of a similar size. The combination of characters of long first antennae with aesthestascs, the absence of oral spines, mandibles lacking lateral spines, and pereopod 7 with a pear­shaped basis, and subchelate propodus and dactylus, readily distinguish this species from all its congeners. Cystisoma gershwinae resembles C. latipes in the length of the first antennae, which have aesthestascs, and the shape of the basis of pereopod 7. It differs from C. latipes mainly in lacking an arched row of oral spines, a character that is only shared with C. fabricii. The shape of the head and eyes is also more like other congeners than C. latipes. Distribution This species has been collected from off Queen Charlotte Islands, British Columbia, Canada, the San Clemente Basin, California, Baja California, Mexico and from the equatorial North Atlantic (type material). Other specimens assigned to this species (non­types) have, in addition, been collected from the western North Pacific, the central Indo­Pacific region and the tropical eastern and western Indian Ocean. This species probably occurs in other regions of the world’s oceans, having been mistaken for C. latipes.Published as part of Zeidler, Wolfgang, 2003, A review of the hyperiidean amphipod family Cystisomatidae Willemöes­Suhm, 1875 (Crustacea: Amphipoda: Hyperiidea), pp. 1-43 in Zootaxa 141 on pages 27-37, DOI: 10.5281/zenodo.15637

    Epimeria (Urepimeria) monodon Stephensen 1947

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    Epimeria (Urepimeria) monodon Stephensen, 1947 Epimeria monodon Stephensen, 1947: 53, fig. 19. Epimeria monodon – J.L. Barnard 1961: 103 (key). — McCain 1971: 161. — Thurston 1972: 34. — De Broyer & Klages 1991: 164 (key). — Coleman 1994: 560, 565–569, figs 9–12; 1998b: 224; 2007: 44, fig. 22a–b, colour plate 2c, map 12 (rhomb). — Watling 1981: 206. — Wakabara & Serejo 1999: 644 (key). Material examined SOUTHERN OCEAN: 1 exuvia, King George Island, Admiralty Bay, kept in aquarium, 14 Apr. 1992, coll. C. De Broyer (RBINS, INV. 132831); 1 exuvia, expedition ARCTOWSKI 1992, King George Island, Admiralty Bay, kept in aquarium, 1 Apr. 1992, coll. C. De Broyer (RBINS, INV. 132832); 1 exuvia, expedition ARCTOWSKI 1992, King George Island, Admiralty Bay, kept in aquarium, 16 Apr. 1992, coll. C. De Broyer (RBINS, INV. 132833); 1 exuvia, expedition ARCTOWSKI 1992, King George Island, Admiralty Bay, kept in aquarium, 30 Mar. 1992, coll. C. De Broyer (RBINS, INV. 132834); 2 specs, expedition ARCTOWSKI 1992, King George Island, Admiralty Bay, kept in aquarium, year 1992, coll. C. De Broyer (RBINS, INV. 132835). Body length Up to 29 mm. Distribution South Orkney Islands; South Shetland Islands; Palmer Archipelago; 0–15 m (De Broyer et al. 2007). Epimeria monodon was recorded from rocky intertidal communities (Aghmich et al. 2016) and was found amidst macro-algae (Richardson 1977). Thurston (1972) stated: “apparently associated with algae growing on rocks, boulder or gravel bottoms, 3–15 m ”. Remarks The record of E. monodon from the eastern shelf of the Weddell Sea at 254 m listed in the data compilation of De Broyer et al. (2007) falls outside of the usual geographical and bathymetric range of the species. It is considered as very suspect. It might be based on E. annabellae, which is common at such depth in the eastern Weddell Sea. It seems possible that E. monodon, which is a very shallow-water species, uses algal rafting as a method of dispersal between islands and archipelagoes. Thurston (1972) recorded a slight sexual dimorphism in the species: “the male has larger eyes and a slightly more prominent projection on pleon segment 3”.Published as part of d'Acoz, Cédric d'Udekem & Verheye, Marie L., 2017, Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea), pp. 1-553 in European Journal of Taxonomy 359 on pages 158-159, DOI: 10.5852/ejt.2017.359, http://zenodo.org/record/385569

    Grandidierella bonnieroides Stephensen 1947

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    Grandidierella bonnieroides Stephensen Grandidierella bonnieroides Stephensen, 1948: 12, fig. 3.— Myers, 1970: 141, fig. 1–2.—1972: 790.— Asari & Myers, 1982: 252, figs. 9–10.— Ledoyer, 1982: 245, fig. 89. Grandidierella bonnieri.—Appadoo & Steele, 1998: 640. Material examined. 733, 12♀♀, AM P60856, from Acanthophora spicifera, Anse la Raie (19°59.5'S 57°37.5'E), 15 May 1998. 733, 8♀♀, AM P60857, from Ulva lactuca and Ulva reticulata, Le Bouchon (20°28'S 57°40.5'E), 27 October 1998. Diagnosis. Sternal process present on pereon segment 1. Coxal plates discontinuous, broader than deep. Antenna 2 robust, articles 4 and 5 subequal, flagellum 5-articulate. Gnathopod 1 carpus slender, subquadrate, 1.7× as long as broad, with parallel anterior and posterior margins, posterior margin with few setae, inner face of carpus with a small process, distal margin with a medial finger-like process followed by a strong process on the posterodistal margin; propodus subquadrate, 2.5× as long as broad, anterior and posterior margins appear parallel; dactylus robust, inner margin crenulate with robust setae. Gnathopod 2 carpus, subovate, 2× as long as broad; propodus subrectangular, 0.6× length of carpus. Uropod 3 uniramous, ramus slightly over twice as long as peduncle, slender, 5× as long as broad with setae on margins and on distal end. Telson with two medial and one distal setae on each side. Female, gnathopod 1 carpus 2.3× as long as broad, propodus medially expanded, slightly shorter than carpus, palm oblique. Variation. In younger males (e.g., 2.8 mm) the sternal process is not developed. The gnathopod 1 carpus anterior margin is more convex and appears globular, distal fingerlike process poorly developed; the propodus is 2× as long as broad, inner margin is more convex with strong groups of setae; dactylus is slender and inner margin is toothed with short setae. Type locality. Salinja Paloe Lechi, Bonaire. Habitat. Grandidierella bonnieroides was collected from depths less than 1 m on the north, east and south coasts of the island. It lives mostly amongst the green alga, Ulva sp. and red algae. It was very abundant at one site on the southeast coast, Le Bouchon, a site characterized by claylike suspended matter in seawater and extensive green algae. Distribution. Madagascar, Mauritius, India, Bonaire, Caribbean, Gulf of Mexico. Remarks. The present material agrees with the description given by Asari & Myers (1982: 252, fig. 9–10). One small difference being the uropod 3 peduncle, which is broader in the material described by Asari & Myers (1982). The present material also agrees with the description given by Ledoyer (1982: 245, fig. 89) where the uropod 3 peduncle is slender. Two sternal processes are illustrated by Ledoyer (1982: 246, fig. 89), but only one is observed in the current material. Myers (1970: 138–139, fig. 2) points out that there may be considerable variation in the relative development of sternal processes on pereon segments 1 and 2 when he reviewed material of G. bonnieroides from the Caribbean and Gulf of Mexico.Published as part of Appadoo, C & Myers, A. A., 2004, Corophiidea (Crustacea: Amphipoda) from Mauritius, pp. 331-362 in Records of the Australian Museum 56 (3) on pages 353-35

    Holophryxus acanthephyrae Stephensen 1912

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    <i>HOLOPHRYXUS ACANTHEPHYRAE</i> STEPHENSEN, 1912 <p> <i>Synonym:</i> <i>Isophryxus concavus</i> Schultz, 1977.</p> <p> <i>Original description:</i> Stephensen (1912 —as <i>H. Acanthephyrae</i>): 112–117, figs 13 (top), 15–21 (♀, ♂ and epicaridium).</p> <p> <i>Additional descriptions:</i> Jones & Smaldon (1986): 306–312, figs 1–6 (♀ and ♂). Schultz (1977 —as <i>I. concavus</i>): 93–96, figs 17, 18A-H, O, P (♀ and ♂). Wasmer (1988): 21–23 (♀ and ♂). Alves-Júnior <i>et al.</i></p> <p>(2019): 14 (♀). Boyko & Williams (2021b): 114–116, fig. 2A-C (♀ and ♂).</p> <i>Hosts: Acanthephyra acanthitelsonis</i>, <i>A. pelagica</i> (type host) and <i>A. purpurea</i> (Pleocyemata, Acanthephyridae). <p> <i>Distribution:</i> North Atlantic Ocean, south-western Atlantic Ocean, South Pacific Ocean, Southern Ocean (Table 2).</p> <p> <i>Differential diagnosis: Holophryxus</i>. Body length 22 mm (♀; <i>N</i> = 1), 0.2 mm (epicaridium) (Stephensen, 1912); 12.18–13.94 mm (♀♀; <i>N</i> = 2), 18.28–22.68 mm (ovigerous ♀♀; <i>N</i> = 2), 3.33 mm (♂; <i>N</i> = 1) (Jones & Smaldon, 1986); 8.5–21.6 mm (♀♀; <i>N</i> = 7), 2.6–3.4 mm (♂♂; <i>N</i> = 3) (Schultz, 1977); 22.7 mm (ovigerous ♀; <i>N</i> = 1), 2.3 mm (♂; <i>N</i> = 1) (Wasmer, 1988); 20.33 mm (♀; <i>N</i> = 1) (Alves-Júnior <i>et al.</i>, 2019); 13.7 mm (♀; <i>N</i> = 1), 2.0 mm (♂; <i>N</i> = 1) (Boyko & Williams 2021b). Colour in alcohol reddish brown spotted with yellow (Stephensen, 1912) or light yellow (Wasmer, 1988) in ♀; yellowish white in ♂ (Wasmer, 1988).</p> <p> <i>FEMALE:</i> Body oblong-ovate, about 1.9–2.2 times as long as maximum width. Cephalon only slightly demarcated from pereon in dorsal aspect by weak constriction bilaterally and transverse surface fold dorsally; dorsal prominence with emarginate (bilobed) anterior margin; part of ventral cephalic ridge clearly discernible in dorsal aspect (Fig. 15H, I), slightly arched, about 0.3 times as long as maximum width, anterior margin slightly sinuous, with pair of distinctive, slit-like pit organs; posterolateral corners of cephalic ridge with rounded processes, representing coxal plates of incorporated first pereonite. Maxilliped present, bilobate. Median sternal plate with paired posteriorly directed appendices. Pereon with five pairs of incubatory plates; oostegite 1 bilobate, with medial projection on proximal lobe; oostegite 5 unilobate, covering 57–63% of ventral side of body, posterior margin with 12–14 spinular extensions. Coxal plates of pereopods 2–5 expressed, represented by rounded processes on lateroventral margins of pereon, separated by transverse folds; those of pereopod 5 weakly developed. Posterior part of pereon without ventrolateral papillae. Pleon clearly demarcated from pereon; about 12–14% of total body length; subtriangular, tapering abruptly in posterior half to rounded extremity; dorsal surface deeply excavated either side of midline near border with pereon.</p> <p> <i>MALE:</i> Body elongate but more compact than in most other species, about 2.5 times as long as maximum width, dorsoventrally flattened. Cephalon completely fused to first pereonite, forming the cephalothorax; original segmentation marked by dorsal groove and weak lobate extensions at posterolateral corners; distinctly wider than long, clearly narrower than pereon; anterior margin rounded. Antennule bilobate, outer lobe with non-articulated flagellum. Antenna 2-segmented; with broad basal part and elongate, posteriorly directed, slender part; not extending to posterior margin of first pereonite. Pereonites 2–6 completely separated, pereonite 7 almost completely fused dorsally with pleon; lateral margins of pereonites 2–6 rounded, those of seventh conical or rounded. Pleon conical, markedly narrower than pereonite 7, representing 31–40% of body length; posterior margin rounded.</p> <p> <i>Variability:</i> Recorded in the number (12–14) of spinular extensions around the posterior margin of the fifth incubatory plate in females (Stephensen, 1912; Schultz, 1977; Jones & Smaldon, 1986; Wasmer, 1988; Alves-Júnior <i>et al.</i>, 2019), and in the shape of the lateral margins of the seventh pereonite in males (Jones & Smaldon, 1986; Wasmer, 1988).</p> <p> <i>Remarks:</i> By far the most detailed account of the adults is that by Wasmer (1988) who pointed out some pertinent errors in previous descriptions. However, his statement (not accompanied by any figures) that the male antennule is 3-segmented and the antenna possibly also displays the same segmentation is almost certainly incorrect (see above). Schultz’s (1977 —as <i>I. concavus</i>) description of the cephalic appendages is grossly inadequate and difficult to interpret. The author states that the antennules and antennae are absent in the female and ‘… some mouth parts [are] distinct, but most not immediately apparent as mouth parts’. It is indeed unclear which appendages Schultz’s figures (his fig. 18C-G) refer to. His description of the male antennule and antennae is similarly ambiguous, the former being described as consisting of three conical segments with a single aesthetasc on the distal one, and the latter as 4-segmented, composed of a broad irregularly-shaped peduncular segment, supporting two subequal, narrow segments, and a small apical segment tipped with several setae.</p> <p> <i>Holophryxus acanthephyrae</i> is the only representative of the genus that is distributed in both northern and southern hemispheres. Within the cephalic type 3 group it is most closely related to <i>Ho. alaskensis</i> but differs from it in a number of significant characters such as the presence of pit organs on the ventral cephalic ridge, and the shape and dorsal sculpturing of the pleon in the female (see above). The epicaridium stage was described by Stephensen (1912: figs 19–21) based on larvae (0.2 mm) recovered from the marsupial cavity of the female; however, the morphology of the pereopods indicate that it was only partially developed (Coyle & Mueller, 1981).</p>Published as part of <i>Huys, Rony, Savchenko, Alexandra S. & Kosobokova, Ksenia N., 2023, Discovery of a new species and host record of Holophryxus Richardson, 1905 (Isopoda: Dajidae) from the central Arctic: a model of enhanced descriptive standards for epicaridean isopods, pp. 592-649 in Zoological Journal of the Linnean Society 198 (2)</i> on pages 636-637, DOI: 10.1093/zoolinnean/zlac105, <a href="http://zenodo.org/record/8011009">http://zenodo.org/record/8011009</a&gt

    Metandania islandica : Stephensen 1925

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    Metandania islandica Stephensen, 1925 (®gures 25±27) Metandania islandica Stephensen, 1925: 136. Metandania islandica: Schellenberg, 1953: 187. Andaniotes islandica: Barnard and Karaman, 1991: 678 (A. islandica synonymized with A. corpulentus). Material examined HOLOTYPE: Zoological Museum, Copenhagen: immature 7 mm, 65 ss 14 ¾ N, 30 ss 39 ¾ W, 1318 m. Additional material: Naturhistorisk a Museet, Gotenborg, Sweden: Female 22 mm (Schellenberg, 1953: 187), 19 ss 49 ¾ N, 65 ss 01 ¾ W, 7900± 7625 m. Distribution Known only from two localities in the north Atlantic, 1318±7900 m. Description The following description and ®gures are all from Schellenberg’s female, 22 mm. Rostrum very small. Antennae elongate. Antenna 1 shorter than antenna 2; ¯agellum four-articulate; accessory ¯agellum article 2 absent. Antenna 2 peduncle (articles 3±5) shorter than ¯agellum; article 3 short, about as long as broad; article 4 shorter than article 5. Epistome curved (convex) and smooth; epistomal plate (medial keel) produced into a large conspicuous medial keel. Mouthparts not elongate or pointed. Mandible incisor transverse; smooth and toothed; left lacinia mobilis present; powerful; laterally straight; not conical. Maxilla 1 palp uni-articulate, oval, setae short, robust, cuspidate, apex reaching above the apex of outer plate; outer plate distally rounded; ST in two parallel rows, ®rst marginal and second submarginal, ST ®rst row with six setae (ST1±5, ST7), ST 1 ordinary (similar to ST 2±4), gap between ST 5 and ST 7, ST A±C present, part of second row; inner plate without a well developed shoulder, setae pappopectinate. Maxilla 2 ordinary; outer plate setae without distal hooks or cleft; inner plate setae row A covering the entire margin, appressed to row B, row A setae pappose, ®rst three or four setae diOEerentiated from the other setae (weakly pappopectinate); row B setae proximally pappose, distally with cusps; row C present; row D absent. Maxilliped palp four-articulate; article 2 distally unproduced; dactylus distally simple (pointed); inner plate not exceeding base of palp article 1, with six nodular setae; medial setae-row present, not reduced, transverse, setae cuspidate; distal setaerow present; inner setae-row present, row not reduced (more than two setae), setae conspicuously large and strong; outer plate outer setae-row present, marginal, setae attached normally; setae short, straight; inner setae-row present but strongly reduced, parallel but not appressed to outer row, setae short simple, slender; distal setaegroup present, setae attached normally; setae long robust. Labrum elongate; lobes asymmetrical; right lobe ordinary; left lobe reduced. Labium distally broad, oval. Coxal plates and basis on the pereopods smooth. Coxae 1±3 contiguous. Pereopod 1 coxal plate not as deep as basis; propodus subovate. Pereopod 2 longer and thinner than pereopod 1; ischium not elongate, ratio length:breadth not exceeding 1.5; ischium distal posterior margin with plumose setae; propodus subrectangular; palm absent. Pereopod 4 coxa posteroventral lobe small, reaching about the base of pereon segment 6; basis anterior and posterior margins with long setae, plumose setae on distal anterior margin, no plumose setae on distal posterior margin; ischium with plumose setae on posterior distal margin. Pereopod 6 basis posteriorly expanded, expansion conspicuous, rounded posteriorly; with a row of long plumose setae. Pereopod 7 basis anterior margin straight, distally rounded; with a medial row of setae present, setae short and robust. Oostegites on pereopod 2±5. Gills on pereopods 2±7. Pleonites 1±3 dorsally smooth. Urosomites 2 and 3 free. Uropod 1 peduncle longer than rami, outer ramus as long as inner. Uropod 2 peduncle longer than rami, outer ramus shorter than inner. Uropod 3 peduncle longer than half the length of rami, outer ramus uni-articulate, outer ramus shorter than inner. Telson as long as broad; shorter than uropod 3 peduncle, cleft, apically rounded, submarginal setae on apex of each lobe. Males. Unknown. Remarks Stephensen (1925) described the present species based on a single immature female captured west of Iceland. Later, Schellenberg (1953) identi®ed a mature female captured at 19 ss 49 ¾ N 65 ss 01 ¾ W. The two specimens show some very important diOEerences (see table 1), but due to lack of material, it is impossible to determine whether the two specimens belong to two diOEerent species or if the diOEerences are due to one of them being immature. However, both specimens also share some very important character-states that clearly group them together and at the same time separate them from Andaniotes. First, the epistome is convex, not laterally produced as in all species belonging to the genera Andaniexis, Andaniotes and Parandaniexis. The epistomal plate is large and conspicuous, and the labrum is not reduced, with the right lobe well developed. Furthermore, the mandibular incisor is partly toothed, resembling the state found in Andaniella, Andaniopsis and Bathystegocephalus, and the setae on the palp of the ®rst maxilla are short robust with cusps (versus long simple and slender in Andaniotes). Examination of the left mandible has also revealed a lacinia mobilis of a very diOEerent morphology than found in any other stegocephalid species (this state was impossible to verify for the holotype, as it was partly damaged, but Stephensen’s original description is similar to the state found on Schellenberg’s female): the lacinia mobilis is almost totally reduced, but the mandibular body itself is produced, thus the lacinia mobilis appear to have gone through ®rst a reduction and then a secondary enlargement. Finally, the accessory ¯agellum on the ®rst antenna is conspicuously large, with lateral setae-groups.Published as part of Berge, JØrgen, 2001, Revision of the Amphipod (Crustacea: Stegocephalidae) genera Andaniotes Stebbing, 1897 and Metandania Stephensen, 1925, pp. 787-832 in Journal of Natural History 35 (6) on pages 825-829, DOI: 10.1080/00222930152123620, http://zenodo.org/record/527701

    "Repositioning of craniofacial tumorous bone after autoclaving"

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    Thirteen bone tumours that were invading the craniofacial skeleton were operated on by intracranial procedures. The resected tumorous bone was autoclaved and put back. Follow up of no less than one year included 122mTc scanning, computed tomography, radiography, bone biopsy and clinical examination. In every case, when rigidly fixed, most of the autoclaved bone was gradually revitalised by invading new and normal bone. We conclude that autoclaved bone will be replaced by normal bone, and that the present technique is justified for reconstruction of complicated structures or large areas of bone after operations for tumours invading the craniofacial skeleton

    Lynn C. Stephensen and Lucinda Ann Wilson grave marker decorated for Halloween in Fountain Green, Utah, 1999

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    Lynn C. Stephensen and Lucinda Ann Wilson grave marker in the Fountain Green Cemetery in Fountain Green, Sanpete County, Utah. Image is of a headstone engraved with a mountain range across the top, evergreen trees, a fence and two horses walking through a stream are on the left side. A Utah State Fireman's association emblem is engraved next to the name Stephensen. A Latter-day Saint temple, evergreen trees and an elk are engraved at the right. On top of the headstone is a garland of yellow and orange silk leaves with a rock to possibly hold it down. A plastic scarecrow with a crow sitting on its arm and holding a jack-o'-lantern is attached to a black metal rod inserted into the ground to the right of the headstone. To the left of the headstone is a metal hook holding up a basket of ivy, twigs, cat tails and a green painted wood plaque that reads 'Lynne, We love and miss you! Mom and Dad.' Black string has been wrapped around the headstone and the decorations on either side
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