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Ditaxiporinae Stach 1935
Subfamily Ditaxiporinae Stach, 1935 Diagnosis. Colony erect, jointed, comprising sterile internodes that are mostly multizooidal and biserial, rarely unizooidal in part or all of the colony; fertile internodes always multizooidal and biserial. Frontal shield varied; multi-costate, or mostly with vestigial suboral costae so that there is an extensive gymnocyst, typically foraminate; or the gymnocyst reduced owing to expansion of pore-chambers frontally so that most of the shield is effectively cryptocystal. Ovicells likewise variable, ectooecium well-developed, smooth, with a suture or foramina, or ectooecium reduced, exposing extensive granular endooecium. Dorsal surface of zooids smooth or with pore-chambers variably developed. Avicularia typically lateral-oral and mostly with complete cross-bar. Remarks. Stach (1933, 1934, 1935) subdivided the Catenicellidae into several subfamilies, basing them primarily on the position of the ovicell within the segment. He named five subfamilies — Vittaticellinae Stach, 1933, Scuticellinae Stach, 1934, Catenicellinae Stach, 1935, Cornuticellinae Stach, 1935, and Ditaxiporinae Stach, 1935. Additionally, Cheetham (1963) introduced a new family-rank taxon, Ditaxiporinidae. Gordon & Braga (1994) commented on these arrangements, synonymising some. They regarded biseriality (i.e., multizooidal) as having more fundamental significance within the family than the position of the ovicell (or the morphology of the frontal shield), which they interpret as having differentiated within different lineages later. For these reasons they subsumed the Ditaxiporinidae into the Ditaxiporinae.Published as part of Vieira, Leandro M., Gordon, Dennis P. & Correia, Monica D., 2007, First record of a living ditaxiporine catenicellid in the Atlantic, with a description of Vasignyella ovicellata n. sp. (Bryozoa), pp. 49-58 in Zootaxa 1582 (1) on page 51, DOI: 10.11646/zootaxa.1582.1.5, http://zenodo.org/record/509762
Acerentomon dispar Stach 1954
<i>Acerentomon dispar</i> Stach, 1954 <p> <i>Annales Zoologici</i>, 16 (3): 23</p> <p> <b>Lectotype.</b> Poland. Tatra Mountains: 1♀ 8/VIII/1923 Stach leg., on slide (NHMD 212202).</p> <p> <b>Current status:</b> Valid name.</p> <p> <b>Additional notes:</b> Lectotype designated by Tuxen (1961a).</p>Published as part of <i>Eusébio, Rita, Sendra, Alberto, Enghoff, Henrik & Reboleira, Ana Sofia P. S., 2018, Catalogue of the type material in the entomological collection of the Natural History Museum of Denmark: basal hexapods, pp. 201-236 in Zootaxa 4457 (2)</i> on page 203, DOI: 10.11646/zootaxa.4457.2.1, <a href="http://zenodo.org/record/1341991">http://zenodo.org/record/1341991</a>
Willemia scandinavica Stach 1949
<i>Willemia scandinavica</i> Stach, 1949 <p> – <i>Willemia scandinavica</i> Stach, 1949a: 249.</p> <p> <b>KS, LO:</b> Kahrarian 2014.</p> <p> <b>Ecology and habitat in Iran:</b> soil and leaf litter under oak trees.</p> <p> <b>General distribution:</b> Holarctic (Fjellberg 1998; Babenko <i>et al.</i> 2019).</p>Published as part of <i>Mayvan, Mahmood Mehrafrooz, Greenslade, Penelope & Sadeghi-Namaghi, Hussein, 2023, An annotated checklist of the Collembola (Hexapoda) from Iran, pp. 1-101 in Zootaxa 5275 (1)</i> on page 13, DOI: 10.11646/zootaxa.5275.1.1, <a href="http://zenodo.org/record/7898948">http://zenodo.org/record/7898948</a>
Willowsia bartkei Stach 1965
<i>Willowsia bartkei</i> Stach, 1965 <p> – <i>Willowsia bartkei</i> Stach, 1965: 361.</p> <p> <b>NK:</b> Mehrafrooz Mayvan <i>et al.</i> 2022a.</p> <p> <b>Ecology and habitat in Iran:</b> soil under oak trees.</p> <p> <b>General distribution:</b> Vietnam (Zhang 2015) and Iran.</p>Published as part of <i>Mayvan, Mahmood Mehrafrooz, Greenslade, Penelope & Sadeghi-Namaghi, Hussein, 2023, An annotated checklist of the Collembola (Hexapoda) from Iran, pp. 1-101 in Zootaxa 5275 (1)</i> on page 38, DOI: 10.11646/zootaxa.5275.1.1, <a href="http://zenodo.org/record/7898948">http://zenodo.org/record/7898948</a>
Ceratophysella adexilis Stach 1964
<i>C. adexilis</i> Stach, 1964 <p> Distribution: China, Beijing (Stach 1964), Jiangsu province, Nanjing (Stach 1964). Note. A member of the <i>C. communis</i> sensu Yosii (1962) complex (see note on <i>C. communis</i> auctt.).</p>Published as part of <i>Jia, Junli, Skarżyński, Dariusz & Li, Youlian, 2010, Ceratophysella taiguensis sp. nov. (Collembola, Hypogastruridae) from China, with an annotated checklist of Chinese Ceratophysella Börner, 1932, pp. 57-63 in Zootaxa 2644</i> on page 61, DOI: <a href="http://zenodo.org/record/198659">10.5281/zenodo.198659</a>
Anurida ellipsoides Stach 1949
<i>Anurida ellipsoides</i> Stach, 1949 <p> – <i>Anurida ellipsoides</i> Stach, 1949b: 34.</p> <p> <b>EA, WA, GI:</b> Cox 1982.</p> <p> <b>Ecology and habitat in Iran:</b> soil.</p> <p> <b>General distribution:</b> Palaearctic (Fjellberg 1998).</p>Published as part of <i>Mayvan, Mahmood Mehrafrooz, Greenslade, Penelope & Sadeghi-Namaghi, Hussein, 2023, An annotated checklist of the Collembola (Hexapoda) from Iran, pp. 1-101 in Zootaxa 5275 (1)</i> on page 25, DOI: 10.11646/zootaxa.5275.1.1, <a href="http://zenodo.org/record/7898948">http://zenodo.org/record/7898948</a>
Folsomia ksenemani Stach 1947
<i>Folsomia ksenemani</i> Stach, 1947 <p> – <i>Folsomia ksenemani</i> Stach, 1947: 134.</p> <p> <b>MAZ:</b> Yoosefi Lafooraki 2014; Yoosefi Lafooraki & Shayanmehr 2014a, b; <b>TE:</b> Qazi & Shayanmehr 2016b.</p> <p> <b>Ecology and habitat in Iran:</b> soil and leaf litter under plum, acacia and oak trees.</p> <p> <b>General distribution:</b> Europe, West Caucasus, Lebanon (Potapov 2001) and Iran.</p>Published as part of <i>Mayvan, Mahmood Mehrafrooz, Greenslade, Penelope & Sadeghi-Namaghi, Hussein, 2023, An annotated checklist of the Collembola (Hexapoda) from Iran, pp. 1-101 in Zootaxa 5275 (1)</i> on page 42, DOI: 10.11646/zootaxa.5275.1.1, <a href="http://zenodo.org/record/7898948">http://zenodo.org/record/7898948</a>
Hypogastrura crassaegranulata Stach 1949
Hypogastrura crassaegranulata (Stach, 1949) sensu nov. Figs 1–8 Neogastrura crassaegranulata crassaegranulata: Stach 1949: 84 Neogastrura crassaegranulata dobsinensis Stach, 1949 syn. nov. Type material. Lectotype male on slide (formerly in alcohol) by present designation, Tatra Mountains (Polish Carpathians), Ðwinica mountain (2301 m a. s. l.), in short dense moss growing in the crakes of granite blocks, 3. VIII. 1909, leg. and det. J. Stach (ISEAC); 3 paralectotypes in alcohol, same data as above. Stach (1949) described H. c. crassaegranulata on the basis of six specimens collected in two localities in the Tatra Mts, but he did not designate the holotype. Consequently five specimens from the first mentioned locality (see Stach 1949) are syntypes. Since the syntype accurately examined by Stach is lost (Weiner in lit.) one of remaining four collected and determined by Stach and generally consistent with the original description was designated as the lectotype. Diagnosis. H. crassaegranulata sensu nov. together with H. franconiana stat. nov., H. hohi Babenko, 1994 (Caucasus), H. microspina Babenko, 1994 (Siberia) and H. ghirkani Babenko, 1994 (Azerbaijan) fully conform to the crassegranulata group sensu Babenko et al. (1994). They are characterised by coarse skin granulation, thick and short ant. IV sensilla, labrum with papillae, broad basal empodial lamella, 1, 1, 1 clavate tibiotarsal hairs, more than 4 + 4 VT setae, quadridentate retinaculum and dens with 7 setae, fine granules and without ventroapical swelling. Morphological differences between species mentioned above are presented in Tab. 2. Redescription. Body length 1 –1.6 mm. Antennae, dorsal part of the body and legs from grey to black, ventral part of the body usually slightly paler. Granulation coarse, 3–6 granules between setae p 1 on abd. V (Figs 2 & 4). Dorsal chaetotaxy of th. II–III and abd. III–VI as in Figs 1 & 2. Chaetotaxy of head typical of the genus. Setae short and smooth. Body sensilla (s) fine and smooth 2–3 times longer than ordinary setae. Th. I with 2 + 2 setae. Th. II with 3 + 3 ordinary m setae (m 2, m 4, m 6 ’, m 6 absent), th. III with 2 + 2 (m 2, m 3, m 4, m 6 ’, m 6 absent). Setae p 3 and p 7 on abd. IV and p 2 on abd. V absent. Chaetotaxy rather variable. Subcoxae I, II, III with 1, 3–4, 3 – 4 setae respectively. Microsensillum on th. II present. Ant. IV with simple apical vesicle, subapical organite, microsensillum, 6–9 short, thick sensilla of variable size and shape (Fig. 5). AOIII with two long (lateral) and two short (internal) curved sensilla (Fig. 5). Microsensillum on ant. III present. Ant. I with 8 setae. Ocelli 8 + 8. Postantennal organ typical of the genus, clearly smaller than neighbour ocelli (Fig. 3). Accessory boss invisible. Labrum with 5, 5, 4 setae and 4 prelabrals. Head of maxilla of the tullbergi type. Outer lobe with 2 sublobal hairs. Labium of the tullbergi type. Tibiotarsi I, II, III with 19, 19, 18 setae respectively. 1, 1, 1 A 1 clavate setae present. Claws with inner tooth. Empodial appendage with broad basal lamella and apical filament reaching middle of inner unguis (Fig. 6). VT with 5–7 + 5–7 setae (Fig. 7). Retinaculum with 4 + 4 teeth. Furca well developed. Dens with fine granules and 7 setae. Mucro narrow with low lamellae (inner usually lower than outer) (Fig. 8). Ratio: dens/mucro=2.5–3.5. Anal spines short, situated on low basal papillae (Figs 2 & 4). Distribution. Tatra Mountains (Polish Carpathians) and Slovenski Raj (Slovak Carpathians). Ecological preferences. This species inhabits mosses growing on the rocks (calcareous mainly) in cold and humid places: shaded steep slopes in the subalpine and alpine zones, glacial cirques and entrances to caves and deep ravines in lower altitudes. Discussion. According to Stach (1949) H. c. crassaegranulata differs from H. c. dobsinensis in number of thick ant. IV sensilla, shape of mucro, claws and empodium (Tab. 1). Analysis of available material showed that real morphological differentiation is low. H. c. crassaegranulata has 6–9 thick ant. IV sensilla, 5–7 VT setae and 3–6 granules between setae p 1 on abd. V, while H. c. dobsinensis has respectively 7–8 thick ant. IV sensilla, 5 VT setae and 4–6 granules between setae p 1 on abd. V. Consequently there are no morphological characters distinguishing these species and H. c. dobsinensis is hereby placed in synonymy with H. crassaegranulata sensu nov. Hypogastrura franconiana (Stach, 1949) stat. nov. Figs 9–16 Neogastrura crassaegranulata franconiana: Stach 1949: 84 Hypogastrura crassaegranulata burgundiana Gisin, 1960 syn. nov. Hypogastrura crassaegranulata estaranhensis Cassagnau, 1959 syn. nov. Type material. Lectotype male on slide (formerly in alcohol) by present designation, Bavaria, Germany, cave Holzknechtsloch near Sackdilling, leg. Spoeker, det. J. Stach (ISE AC); paralectotype female on slide (formerly in alcohol), same data as above. Stach (1949) described H. c. franconiana on the basis of five specimens collected in Bavaria, but he did not designate the holotype. Since three syntypes accurately examined by Stach are lost (Weiner in lit.) one of remaining two determined by Stach and generally consistent with the original description was designated as the lectotype. Diagnosis. H. franconiana stat. nov. together with H. crassaegranulata sensu nov., H. hohi Babenko, 1994, H. microspina Babenko, 1994 and H. ghirkani Babenko, 1994 fit the definition of the crassegranulata group sensu Babenko et al. (1994) (see diagnosis H. crassaegranulata sensu nov.). Morphological differences between species mentioned above are presented in Tab. 2. Redescription. Body length 1 –1.6 mm. Antennae, dorsal part of the body and legs grey or black, ventral part of the body slightly paler. Granulation coarse, usually 3–5 granules between setae p 1 on abd. V (Figs 10 & 12). Dorsal chaetotaxy of th. II–III and abd. III–VI as in Figs 9 & 10. Chaetotaxy of head typical of the genus. Setae short and smooth. Body sensilla (s) fine and smooth about twice as long as ordinary setae. Th. I with 3 + 3 setae. Th. II with 6 + 6 ordinary m setae (m 2, m 4, m 6 ’ present), th. III with 5 + 5 (m 2, m 3, m 6 ’ present). Setae p 3 and p 7 on abd. IV present, setae p 2 on abd. V present or absent. Chaetotaxy rather variable. Subcoxae I, II, III with 1, 3–4, 3 – 4 setae respectively. Microsensillum on th. II present. Ant. IV with simple apical vesicle, subapical organite, microsensillum, 6–9 short, thick sensilla of variable size and shape (Fig. 13). AOIII with two long (lateral) and two short (internal) curved sensilla (Fig. 13). Microsensillum on ant. III present. Ant. I with 8 setae. Ocelli 8 + 8. Postantennal organ typical of the genus, slightly smaller than neighbour ocellus. Accessory boss present (Fig. 11). Labrum with 5, 5, 4 setae and 4 prelabrals. Head of maxilla of the tullbergi type. Outer lobe with 2 sublobal hairs. Labium of the tullbergi type. Tibiotarsi I, II, III with 19, 19, 18 setae respectively. 1, 1, 1 A 1 clavate setae present. Claws with inner tooth. Empodial appendage with broad basal lamella and apical filament reaching middle of inner unguis (Fig. 14). VT with 5–6 + 5–6 setae (Fig. 15). Retinaculum with 4 + 4 teeth. Furca well developed. Dens with fine granules and 7 setae. Mucro narrow with low lamellae (inner usually lower than outer) (Fig. 16). Ratio: dens/mucro=2.5–3.5. Anal spines large, slightly curved, situated on high basal papillae (Figs 10 & 12). Distribution. Bavaria (Germany), Côte d’Or, SaoneetLoire and Pyrenees (France). Ecological preferences. As for H. crassaegranulata sensu nov. Discussion. Morphological differentiation of H. c. franconiana, H. c. estaranhensis and H. c. burgundiana was found to be low after examination of available material. H. c. franconiana has 7–9 thick ant. IV sensilla (2–3 according to Stach 1949), 5 VT setae and 3–5 granules between setae p 1 on abd. V, whereas H. c. burgundiana and H. c. estaranhensis have 6–7 thick ant. IV sensilla, 5–6 VT setae and 3–4 granules between setae p 1 on abd. V. Moreover this last subspecies is usually devoid of setae p 2 on abd. V. This low level of morphological differentiation was considered to be due to populational variability. As a consequence H. c. estaranhensis and H. c. burgundiana are synonymized with H. franconiana stat. nov.. A single Caucasian specimen of “ H. crassaegranulata ” from the Stach’s collection (Domagor mountain, 3600 m a. s. l., short moss sparely growing on the peak, 31. VII. 1936, leg. R. Wojtusiak, ISEAC) has only weakly visible coarse skin granulation, thick ant. IV sensilla and fully complete chaetotaxy. To consider it as H. franconiana stat. nov. seems to be risky at this moment because at least two “forms” morphologically similar to H. franconiana stat. nov. occur in the Caucasus (see Babenko et al. 1994). The taxonomic status of the H. crassaegranulata complex sensu Babenko et al. (1994) should be resolved by further studies. Stach (1949) suggested a possible synonymy of H. c. franconiana with Achorutes schaefferi Carl, 1901 (= A. affinis Schäffer, 1900). One of the two saved specimens of this last species from the Schäffer’s collection was examined and determined as H. purpurescens (Lubbock, 1867). It has long and thin ant. IV sensilla, chaetotaxy as in Babenko et al. (1994: 114, Fig. 46, 1), 2, 3, 2 tenent hairs (however only A 1 clavate) in arrangement as in Fjellberg (1998: 35, Figs 15 AC) and tridentate retinaculum. In this context synonymy of H. franconiana stat. nov. with A. schaefferi and a synonymy of A. schäfferi with H. sahlbergi (Reuter, 1895) suggested by Gisin (1949, 1960b) seem to be doubtful. A probable synonymy of A. schaefferi with H. purpurescens is also uncertain because we do not know if juveniles originated from the type series were examined. The identity of A. schaefferi should be resolved by further studies.Published as part of Skarżyñski, Dariusz, 2006, A taxonomic revision of Hypogastrura crassaegranulata (Stach, 1949) (Collembola, Hypogastruridae), pp. 27-37 in Zootaxa 1109 on pages 29-35, DOI: 10.5281/zenodo.17151
Eosentomon armatum Stach 1927
<i>Eosentomon armatum</i> Stach, 1927 <p> <i>Sprawozdanie Komisji Fizjograficznej. Polska Akademia Umiejętności</i>, 61: 210</p> <p> <b>Holotype.</b> Poland. Czarny, Dunajec: 1ex. 28/VIII/1920 (no collector available on the label), on slide (NHMD 212543).</p> <p> <b>Current status:</b> Valid name.</p>Published as part of <i>Eusébio, Rita, Sendra, Alberto, Enghoff, Henrik & Reboleira, Ana Sofia P. S., 2018, Catalogue of the type material in the entomological collection of the Natural History Museum of Denmark: basal hexapods, pp. 201-236 in Zootaxa 4457 (2)</i> on page 214, DOI: 10.11646/zootaxa.4457.2.1, <a href="http://zenodo.org/record/1341991">http://zenodo.org/record/1341991</a>
Entomobrya schoetti Stach 1922
<i>Entomobrya schoetti</i> Stach, 1922 <p> – <i>Entomobrya schoetti</i> Stach, 1922: 41.</p> <p> <b>KS:</b> Kahrarian <i>et al.</i> 2014a; Kahrarian 2015, Kahrarian 2019; <b>MAZ:</b> Yahyapour <i>et al.</i> 2018, 2019.</p> <p> <b>Ecology and habitat in Iran:</b> soil in apple orchad.</p> <p> <b>General distribution:</b> Europe (Jordana 2012) and Iran.</p>Published as part of <i>Mayvan, Mahmood Mehrafrooz, Greenslade, Penelope & Sadeghi-Namaghi, Hussein, 2023, An annotated checklist of the Collembola (Hexapoda) from Iran, pp. 1-101 in Zootaxa 5275 (1)</i> on page 35, DOI: 10.11646/zootaxa.5275.1.1, <a href="http://zenodo.org/record/7898948">http://zenodo.org/record/7898948</a>
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