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    Arrenurus (Dividuracarus) neogereckei Smit, 2010, n. sp.

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    Arrenurus (Dividuracarus) neogereckei n. sp. (Figs. 3 A–F) Material examined. Holotype male, Small lake Hawkwood Road, 13 km S of Mundaburra, Queensland, Australia, 25 ° 39.910 S 151 ° 13.941 E, 31 October 2005 (QM). Paratypes: 3 females (QM), 2 females (ZMAN), same data as holotype. Diagnosis. Characterized by its complex petiole; central idiosoma part with a rectangular hump. Description. Male: Idiosoma 964 long (without petiole) and 729 wide, divided in three parts, idiosoma colour green. Idiosoma anteriorly truncated, slightly concave. D 1 on a small hump, D 2 slightly anteromedial of D 1. Dorsal idiosoma posteriorly with a rectangular hump (Fig. 3 A). Anterior coxae not extending to anterior idiosoma margin. Medial margins of third and fourth coxal plates of equal length (or medial length of third coxal plate larger). Gonopore rounded, 50 long. Genital plates indistinct, slightly bowed, laterally widened, extending to lateral idiosoma margin (Fig. 3 B). Petiole complex, protrusible, laterally with hyaline lamellae, two triangular structures each with two stout setae. In lateral view posteriorly open with a hook-like extension (Fig. 3 C, D). Lengths of PI-PV: 36, 96, 62, 100, 50; PII medially with three setae, PIV anteroventrally rounded (Fig. 3 E). Lengths of I-leg- 4-6: 136, 144, 150. Lengths of IV-leg- 4-6: 259, 184, 166; IV-leg- 4 without spur. Second, third and fourth legs with numerous swimming setae. Female: Idiosoma 1069 (940–1158) long and 875 (850–988) wide, anteriorly rounded. Idiosoma strongly truncated anteriorly, and less truncated posteriorly. Dorsal shield complete, 705 (640–745) long and 607 (595– 668) wide. First coxae not extending to anterior idiosoma margin. Medial margin of third coxae larger than medial margin of fourth coxae, medial corner of fourth coxae rounded. Gonopore extending laterally, 162 long and 283 wide. Gonopore medially with chitinized patches, which extends along anterior and posterior margin (Fig. 3 F). Genital field 535 wide. Genital plates short and broad. Lengths of PI-PV: 36, 88, 50, 104, 47; palp as in male. Lengths of I-leg- 4-6: 130, 132, 108. Lengths of IV-leg- 4-6: 235, 211, 166. Second, third and fourth legs with numerous swimming setae, first leg with a few swimming setae. Etymology. Named for its resemblance with A. gereckei Smit. Remarks. This is the third species of the subgenus Dividuracarus Smit, a subgenus thus far confined to Australia. The new species is close to A. gereckei Smit, 1997 but the hump on the middle idiosoma part of the male is rectangular (rounded in gereckei) and the petiole is of different shape (with four stout setae instead of two, laterally with hyaline lamellae which are absent in gereckei). Moreover, the idiosoma shape is more stocky, while the hump of the dorsal idiosoma part is less pointed. The female is also close to A. gereckei, but the latter species does not have the chitinized patches on the gonopore.Published as part of Smit, Harry, 2010, Australian Arrenurus (Acari, Hydrachnidia) with the description of eleven new species, pp. 1-26 in Zootaxa 2541 on pages 5-7, DOI: 10.5281/zenodo.19669

    Erratum to: Effect of moderate red wine intake on cardiac prognosis after recent acute myocardial infarction of subjects with Type 2 diabetes mellitus (Diabetic Medicine, (2006), 23, 9, (974-981), 10.1111/j.1464-5491.2006.01886.x)

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    In an article by Marfella et al, the author name C. Saron is incorrect and should be listed as C. Sardu. Therefore the correct author list is: R. Marfella, F. Cacciapuoti, M. Siniscalchi, F. C. Sasso, F. Marchese, F. Cinone, E. Musacchio, M. A. Marfella, L. Ruggiero, G. Chiorazzo, D. Liberti, G. Chiorazzo, G. F. Nicoletti, C. Sardu, F. D'Andrea, C. Ammendola, M. Verza and L. Coppola.In an article by Marfella et al, the author name C. Saron is incorrect and should be listed as C. Sardu. Therefore the correct author list is: R. Marfella, F. Cacciapuoti, M. Siniscalchi, F. C. Sasso, F. Marchese, F. Cinone, E. Musacchio, M. A. Marfella, L. Ruggiero, G. Chiorazzo, D. Liberti, G. Chiorazzo, G. F. Nicoletti, C. Sardu, F. D'Andrea, C. Ammendola, M. Verza and L. Coppola

    Monatractides polyhymnia Pešić & Smit, 2014, n. sp.

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    Monatractides polyhymnia n. sp. (Figs. 30 A–F, 31 A–C, 37 E–F, 38 E–F) Type series. Holotype male, dissected and slide mounted, Ghana: Kue River, Kyabobo NP, 8 º 31.087 N, 0º 36.049 E, alt. 208 m asl., 25.ii. 2013, Smit. Paratypes: 1 / 1 /0, same data as holotype, 0/ 1 /0 dissected and slide mounted. Diagnosis. P- 3 ventral margin concave, sexual dimorphism of P- 3, P- 4 bowed, ventral margin with setaebearing protrusion in distal half of segment. Male: genital field oval (L/W ratio 1.26); P- 3 with three stout, medially orientated setae at the ventrodistal margin. Female: genital field elongated trapezoid (L/W ratio 1.2); P- 3 distally with ventrolateral seta. Description. General features —Idiosoma roundish; frontal platelets shorter than shoulder platelets (shoulder/ frontal platelets L ratio 1.1–1.2); frontal margin with flat protrusions medial to the eyes (Fig. 31 A); the tips of Cx-I orientated to the lateral side. Cxgl- 4 located far anteriorly, near insertion of I-L; gnathosomal bay deep U-shaped, proximally wide box-shaped; posterior suture lines of Cx-IV extending posteriorly beyond posterior margin of genital field and directed posterolaterally; excretory pore and Vgl– 2 away from the line of primary sclerotization, Vgl– 2 posterior to excretory pore; gnathosoma nearly triangular in lateral view, with short, cone-shaped rostrum (Fig. 30 F); palp compact, ventral margin of P- 2 proximally with sharp bend, distally straight, bearing ventral seta, P- 3 ventral margin concave, sexual dimorphism of P- 3, P- 4 bowed, ventral margin with setae-bearing protrusion in distal half of segment. Male: medial margin of Cx-II/III relatively short; genital field small and oval in shape; P- 3 ventrodistally bearing three stout (short and thick) setae, orientated medially and one ventrolateral seta (Figs. 30 C–D). Female: suture lines Cx-I/II complete, not meeting in the medial line, but separately ending at anterior genital field margin (Fig. 31 B); genital field pentagonal; P- 3 distally with ventrolateral seta (Fig. 31 C). Measurements. Male (holotype)—Idiosoma (ventral view: Figs. 30 A, 38 E) L 665, W 499; dorsal shield (Figs. 30 B, 37 E) L 559, W 418, L/W ratio 1.34; dorsal plate L 516; shoulder platelets L 142–147, W 69–70, L/W ratio 2.07–2.09; frontal platelets L 120–127, W 58–59, L/W ratio 2.03–2.2; shoulder/frontal platelets L ratio 1.12–1.22. Gnathosomal bay L 131, Cx-I total L 281, Cx-I mL 150, Cx-II+III mL 22; ratio Cx-I L/Cx-II+III mL 12.8; Cx-I mL/Cx-II+III mL 6.8. Genital field L/W 106 / 84, ratio 1.26; distance genital field-excretory pore 175, genital fieldcaudal idiosoma margin 241. Gnathosoma vL 184; palp total L 232–233, dL/H, dL/H ratio: P-1, 21– 22 / 28, 0.78; P- 2, 68/ 47, 1.43; P-3, 49/ 43, 1.15; P-4, 68/ 28, 2.4; P-5, 26/ 13, 2.0; P- 2 /P- 4 ratio 1.0; dL of I-L- 5–6 (Fig.): 122, 128; I-L- 6 H 58–59, dL/H I-L- 6 ratio 2.2. Female (paratype from Kue River, n = 1)—Idiosoma (ventral view: Figs. 31 B, 38 F) L 731, W 584; dorsal shield (Fig. 37 F) L 641, W 494, L/W ratio 1.3; dorsal plate L 594; shoulder platelets L 156–159, W 78–81, L/W ratio 2.0; frontal platelets L 131–134, W 67, L/W ratio 1.95 –2.0; shoulder/frontal platelets L ratio 1.19. Gnathosomal bay L 147, Cx-I total L 267. Genital field L/W 173 / 145, ratio 1.19; distance genital field-excretory pore 209, genital field-caudal idiosoma margin 284. Gnathosoma vL 199; chelicera total L 240; palp total L 229, dL/H, dL/H ratio: P-1, 23/ 28–29, 0.81; P-2, 69/ 48, 1.43; P-3, 48/ 38–39, 1.24; P-4, 64/ 29, 2.16; P-5, 25/ 12, 2.06; P- 2 /P- 4 ratio 1.09; dL of I-L- 2–6: 69, 92, 111, 122, 131; I-L- 6 H 55, dL/H I-L- 6 ratio 2.4. Etymology. The species is named after Polyhymnia (Ancient Greek: Πολυύμνια), one of nine Muses from Greek mythology, who was a patron of sacred poetry and song. The species name is a noun in apposition (in the nominative case). Discussion. The new species shows a remarkable sexual dimorphism, as in the male P- 3 has a group of three stout, medially orientated setae at the ventrodistal margin, while these setae are lacking in the female. Due to the shape of P- 3 with stout, medially oriented setae at the ventrodistal margin, the new species from Ghana seems to be closely related to M. spinipalpis (K.O. Viets, 1981) a species described by K.O. Viets (1981) based on a single male from South Africa. Later on, Goldschmidt & Smit (2009) reported a single male of M. spinipalpis from Kwazulu- Natal Province and provided an additional description of this species. In the original description, K.O. Viets (1981) described this species as close to M. mitchelli (K.O. Viets, 1965), but differing in a stout, medially oriented setae at the ventrodistal margin of P- 3 lacking in the latter species. Obviously, he was not aware that the difference in the shape of P- 3 might be results of the sexual dimorphism, as the latter species was known only from two females from Transvaal (K.O. Viets 1965). It is possible that M. mitchelli represents a senior synonym of M. spinipalpis but this should be checked by finding additional material, including the male, from the locus typicus of M. mitchelli. Monatractides spinipalpis differs from the new species from Ghana in larger idiosoma dimensions, a more elongated genital field (L/W ratio 1.41–1.55, data taken from K.O. Viets 1981 and Goldschmidt & Smit 2009), P- 2 shorter than P- 4 and by having a well developed ventrodistal protrusion on P- 3. The female of M. mitchelli can be separated by the more elongated genital field, strongly tapering to the caudal end (see K.O.Viets 1958, Fig. 9). Remarks. No ejaculatory complex of Monatractides spinipalpis has been described yet (see: K.O. Viets 1981, Goldschmidt & Smit 2009) Also in males in the present study this organ was not found, but sclerotized framework immediately above the genital field can be hardly visible. Habitat. A sandy/bouldary streams, shaded by riparian vegetation (see Fig. 45 C in Pešić et al. 2013). Distribution. Ghana; known only from the type locality.Published as part of Pešić, Vladimir & Smit, Harry, 2014, Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Ghana, pp. 1-80 in Zootaxa 3820 (1) on pages 46-49, DOI: 10.11646/zootaxa.3820.1.1, http://zenodo.org/record/28630

    FIGURES 1A–F. Aspidiobates imperfectus n in The water mite genus Aspidiobates Lundblad, 1941 from Australia (Acari: Hydrachnidia: Hygrobatidae) with the description of two new species

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    FIGURES 1A–F. Aspidiobates imperfectus n. sp., A–C = holotype female, D–F = paratype male. A = dorsum; B = venter; C = palp; D = dorsum; E = venter; F = palp (P5 lacking). Scale bars: A–B, D–E = 100 µm; C, F = 50 µm.Published as part of Smit, Harry, 2019, The water mite genus Aspidiobates Lundblad, 1941 from Australia (Acari: Hydrachnidia: Hygrobatidae) with the description of two new species, pp. 115-122 in Zootaxa 4550 (1) on page 119, DOI: 10.11646/zootaxa.4550.1.5, http://zenodo.org/record/262517

    FIGURE 41A–F. Monatractides minuta n in Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo

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    FIGURE 41A–F. Monatractides minuta n. sp., holotype female, Great Lumotok stream: A = frontal margin of idiosoma, dorsal view; B = dorsal shield; C = idiosoma, ventral view; D = palp, medial view; E = I-L-5 and -6; F = gnathosoma. Scale bars = 100 µm.Published as part of Pešić, Vladimir & Smit, Harry, 2014, Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo, pp. 1-72 in Zootaxa 3840 (1) on page 66, DOI: 10.11646/zootaxa.3840.1.1, http://zenodo.org/record/492789

    Monatractides urania Pešić & Smit, 2014, n. sp.

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    Monatractides urania n. sp. (Figs. 36 A–H, 37 I, 38 I) Type series. Holotype male, dissected and slide mounted, Ghana, Kintampo Falls, 8 º 05.413 N, 1 º 41.881 W, 3.iii. 2011, Smit. Paratype: 3 /0/0, same data as holotype, 2 /0/0 dissected and slide mounted. Diagnosis (Female unknown). Shoulder platelets shorter than frontal platelets (shoulder/frontal platelets L ratio 0.93–0.96); medial margin with convex protrusions medial to the eyes; medial margin of Cx-II/III very short (21 µm); genital field large (L 140–160 µm) and strongly trapezoid in shape, anteriorly enlarged and pointed; I-L- 6 L/H ratio 2.0– 2.1, distally thickened. Description. Male. General features —Idiosoma elongated-oval; shoulder platelets short, slightly shorter than frontal platelets (shoulder/frontal platelets L ratio 0.93–0.96); frontal margin medially convexly rounded between anterolaterally orientated apodemes (Fig. 36 A); Cxgl– 4 located far anteriorly, near tips of Cx-I; gnathosomal bay, U-shaped, proximally rectangular box-shaped; medial margin of Cx-II/III relatively very short; genital field large and strongly trapezoid in shape, anteriorly enlarged and pointed, laterally convex, tapering towards posterior (Figs. 36 C–D); suture line of Cx-IV extending back beyond posterior margin of genital field; excretory pore slightly away from the line of primary sclerotization, excretory pore anterior to Vgl– 2; gnathosoma nearly triangular in lateral view (Fig. 36 G); P- 2 and P- 3 distal margins without pointed tips, ventral seta on P- 4 short (Figs. 36 E–F); I- L- 6 L/H ratio 2.0– 2.1, basally narrowed, distally thickened (Fig. 36 H). Measurements. Male (holotype, in parentheses measurements of paratypes, n = 1)—Idiosoma (ventral view: Figs. 36 C, 38 I) L 619 (642), W 425 (434); dorsal shield (Figs. 36 B, 37 I) L 525 (550), W 366 (384), L/W ratio 1.43 (1.43); dorsal plate L 491 (509); shoulder platelets L 116–119 (127–131), W 50–52 (55–58), L/W ratio 2.2–2.4 (2.3); frontal platelets L 123–125 (127–131), W 48 (50–52), L/W ratio 2.5–2.6 (2.5–2.6); shoulder/frontal platelets L ratio 0.93–0.96 (0.96). Gnathosomal bay L 150 (153), Cx-I total L 275 (306), Cx-I mL 125 (122), Cx-II+III mL 21 (21); ratio Cx-I L/Cx-II+III mL 12.9 (14.6); Cx-I mL/Cx-II+III mL 5.9 (5.8). Genital field L/W 141 (155)/ 127 (130), ratio 1.11 (1.19); distance genital field-excretory pore 133 (131), genital field-caudal idiosoma margin 181 (189). Gnathosoma vL 157 (155); chelicera total L 189 (198); palp total L 188–190 (188–191), dL/H, dL/H ratio: P-1, 26/ 20, 1.3 (26 / 21, 1.26); P-2, 57/ 38–39, 1.48 (54–55 / 37, 1.48); P-3, 38– 39 / 32, 1.2 (38–39 / 31, 1.25); P-4, 49/ 22, 2.2 (52 / 21–22, 2.4); P-5, 18– 19 / 11–12, 1.6 (18–19 / 12, 1.5); P- 2 /P- 4 ratio 1.16 (1.04); dL of I-L- 2–6: 63 (65), 76 (80), 97 (99), 92 (97), 100 (106); I-L- 6 H 48 (52), dL/H I-L- 6 ratio 2.08 (2.03). Female: unknown. Etymology. The species is named after Urania (Ancient Greek: Οὐρανία), one of nine Muses from Greek mythology, who was a patron of tragedy. The species name is a noun in apposition (in the nominative case). Discussion. The combination of very short medial margin of Cx-II/III, the box-shaped gnathosomal bay, the large and strongly trapezoid genital field, makes the new species close to Monatractides (M.) jucundus (Lundblad, 1927) (syn. M. afer (Lundblad, 1927)), known from East Africa (Lundblad 1952, Walter & Bader 1952). The male of latter species differs from M. urania n. sp., in its larger dimensions of idiosoma and palps, a frontal margin medially straight, a shoulder platelets longer than medial ones and a genital field anteriorly less pointed (see Lundblad 1952, figs. 38 A–B). Remarks. The ejaculatory complex of the male was not found, but sclerotized framework immediately above the genital field can be visible. For a discussion on the ejaculatory complex in M. jucundus see the following species. Habitat. A sandy/bouldary streams, shaded by riparian vegetation. Distribution. Ghana, known only from the type locality. Monatractides microstoma (Koenike, 1898) — species complex (Figs. 39 A–G, 40 A–G, 41 A–D, 42 A–H, 43 A–I, 44 E–L, 46 A–C, Tabs. 4–5) New records. Ghana: Apkonu stream, downstream of falls, 6 º 53.054 N, 0º 28.024 E, alt. 362 m asl., 21.ii. 2013, Smit, 10 / 14 /0 (2 / 2 /0 Form ''A'' [2 / 1 /0 mounted] + 8 / 13 /0 Form ''B'' [6 / 2 /0 mounted]); Small rainforest stream upstream Sagyimase, Atewa Hills, 6 º 13.964 N, 0º 33.116 W, alt. 654 m asl., 8.iii. 2011, Smit, 0/ 1 / 1 (Form ''B''); Kuluga River, N of Biakpa, 6 º 51.365 N, 0º 25.101 E, alt. 388 m asl., 19.ii. 2013, Smit, 0/ 1 /0 (mounted) (Form ''B''); Kuluga River, upstream, hyporheic, 6 º 51.223 N, 0º 25.141 E, alt. 410 m asl., 20.ii. 2013, Smit, 4 / 3 (one juvenile)/0 (3 /0/0 mounted) (1 /0/0 Form ''A'' [mounted] + 2 / 3 /0 Form ''B'' [2 /0/0 mounted]); Agumatsa River at first bridge, Agumatsa Wildlife Sanctuary, 7 º 06.830 N, 0º 35.760 E, alt. 253 m asl., 22.ii. 2013, Smit, 5 / 9 /0 (Form ''A''); stream Boti Falls, 6 º 11.508 N, 0º 13.010 W, alt. 273 m asl, 9.iii. 2011, Smit, 0/ 1 /0 (Form ''B''); Kintampo Falls, 8 º 05.413 N, 1 º 41.881 W, alt. 235 m asl, 3.iii. 2011, Smit, 8 / 7 (one juvenile)/0 (3 / 1 /0 mounted) (4 / 5 /0 Form ''A'' [1 / 1 /0 mounted] + 4 / 2 /0 Form ''B'' [2 /0/0 mounted]); Afiaso stream, Kakum NP, 5 º 30.087 N, 1 º 26.373 W, alt. 114 m asl., 12.ii. 2013, Smit, 0/ 1 /0 (Form A); Fuller Falls, 8 º 04.975 N, 1 º 47.842 W, alt. 189 m asl, 6.iii. 2011, Smit, 0/ 1 /0 (Form ''A''); Supon stream near Asiakwa, Atewa Hills, 6 º 15.530 N, 0º 30.642 W, alt. 250 m asl, 7.iii. 2011, Smit, 1 /0/0 (Form ''A''); Tagbo River, downstream of falls, 7 º 00.708 N, 0º 34.326 E, alt. 394 m asl., 23.ii. 2013, Smit, 19 / 11 (one juvenile)/0 (6 / 2 /0 mounted) (10 / 5 /0 Form ''A'' [4 / 1 /0 mounted] + 8 / 6 /0 Form ''B'' [2 / 1 /0 mounted]); Akaa Falls, 6 º 10.516 N, 0º 11.723 E, alt. 180 m asl., 9.iii. 2011, Smit, 8 / 12 / 1 (1 /0/0 mounted) (Form ''B''); Nubui River, Agumatsa Wildlife Sanctuary, 7 º 06.986 N, 0º 35.548 E, alt. 254 m asl., 22.ii. 2013, Smit, 10 / 14 (2 juveniles)/0 (Form ''A''); Laboun River, downstream of falls, Kyabobo NP, 8 º 19.836 N, 0º 35.487 E, alt. 342 m asl., 24.ii. 2013, Smit, 2 /0/0 (1 /0/0 Form ''A'' + 1 /0/0 Form ''B''); stream downstream of falls, Wanjakli River, Likpe Todome, 7 º 09.976 N, 0º 36.456 E, alt. 478 m asl., 23.ii. 2013, Smit, 3 / 9 / 2 (1 / 1 /0 Form ''A'' + 0/ 1 /0 Form ''B'' mounted); Plunge pool Agumatsa Falls, Agumatsa Wildlife Sanctuary, 7 º 06.350 N, 0º 36.476 E, alt. 258 m asl., 22.ii. 2013, Smit, 12 / 8 /0 (1 / 2 /0 mounted) (Form ''A''). Morphology. General features —Idiosoma elongated oval; frontal platelets only slightly shorter than shoulder platelets; frontal margin medially straight or slightly convex, between flat, anterolaterally pointed protrusions (Figs. 39 A, 40 A); Cxgl– 4 located far anteriorly, near tips of Cx-I; gnathosomal bay moderately deep, U-shaped, proximally rectangular box-shaped; suture line of Cx-IV strongly extending posteriorly beyond posterior margin of genital field, laterally curved (Figs. 39 C, 40 C); excretory pore and Vgl– 2 away from the line of primary sclerotization, excretory pore anterior to Vgl– 2; ventral seta on P- 4 short (Figs. 39 D–E, 40 D–E). Male: medial margin of Cx-II/III relatively short; genital field elongated, anteriorly enlarged. Female: genital field elongated pentagonal, anteriorly rounded. ......continued on the next page Apkonu Apkonu M. procursa, K.O. Viets & Böttger 1974, Congo stream, ♂, stream, ♀, holotype ♀ paratype ♂, ♂, n = 8 ♀, n = 9 n = 5 n = 2 slide no. 4768 slide no. 4797 ......continued on the next page Remarks. There appear to be two forms of this species (or two sibling species within Monatractides microstoma species - complex, see above for a discussion). At one extreme end (Form ''A''), specimens are generally of smaller dimensions, the dorsal plate with the area of primary sclerotization bears four dorsoglandularia (Figs. 39 B, 41 A, 43 A), the gnathosoma is more elongated (Fig. 39 F) and the male genital field is anteriorly projecting and strongly pointed (Figs. 39 C, 42 A). At the other extreme end (form ''B''), specimens are generally of larger dimensions, the dorsal plate with the area of primary sclerotization bears two dorsoglandularia (Figs. 40 A, 41 C, 43 B), the gnathosoma is more triangular in shape (Fig. 40 F), P- 4 has a small but clearly visible denticle near insertion of ventral setae (Figs. 40 D–E) and the male genital field is anteriorly less projecting and slightly pointed (Figs. 40 C, 42 C–D). The figures 39 A–G and 40 A–G illustrate the two extremes. However it is necessary to stress that in many cases we observed intermediate characters (in shape of male genital field, see Figs. 42 A–D) between these two extremes, even though at most sampling sites where these two forms are living together it is easily possible to identify the two groups (see ''New records''). There is a large group of African species sharing the more or less elongated idiosoma and the box-shaped gnathosomal bay. This group includes several similar but still weakly defined species, i.e., Monatractides microstoma (Koenike, 1898), M. amplipalpis (Lundblad, 1951), M. longicoxalis (Cook, 1966), M. dolichosoma (Cook, 1966), M. congoensis (K.O. Viets, 1974), M. rivalis (K.O. Viets, 1974), M. procursa (K.O. Viets, 1974) and M. nigromontanus Goldschmidt & Smit 2009 (see Fig. 45 for their distribution in Africa). Due to the similar shape of the idiosoma and palp the specimens from our study most closely resemble Monatractides microstoma (Koenike, 1898) and M. procursa (K.O. Viets, 1974). The latter species was described by K.O. Viets (in K.O. Viets & Böttger (1974) from Congo (Zaire). Due to the dorsal plate with the area of primary sclerotization bearing two dorsoglandularia and similar shape of the genital field, M. procursa resembles more the specimens of Form ''B'' in our study. Differences with figures and measurements in the original description are found in the excretory pore shifted to the line of primary sclerotization in the female (see Fig. 75 in K.O. Viets & Böttger 1974, postgenital area in male not illustrated). Monatractides microstoma was decribed by Koenike (1898) from Cameroon based on a single female. Later on a detailed description of the female (from Manoka near Duala, prep. no. 1390) was given by K. Viets (1958). Cook (1966) reported M. microstoma as the most frequently collected and most widely distributed torrenticolid species in Liberia and provided illustrations and descriptions of the male. K.O. Viets (1971) published from Zimbabwe a female slightly larger than those from Liberia also under the name microstoma. Due to the dorsal plate with the area of primary sclerotization bearing four dorsoglandularia, M. microstoma is more similar to the specimens of Form ''A'' of our study. Males from Liberia differ from those in our study in a medial suture of Cx- II+III of moderate length, the genital field anteriorly less projecting and slightly pointed and a less extended postgenital area (see Cook 1966, fig. 262). Among the specimens from Ghana we found a single male from Tagbo stream which fits well Cook’s (1966) description of microstoma (see Figs. 46 A–C, and measurements in Tab. 4). This specimen also agrees in the shape of the dorsal shield, gnathosoma elongated in lateral view and P- 4 without lacking denticle at insertion of ventral setae, but it is notably larger than our specimens from Ghana. As the holotype of M. microstoma is lost (see K. Viets 1958), additional material, including the male, and selection of a neotype from the locus typicus is necessary to clarify the taxonomy of this species. Monatractides rivalis (Congo) and M. nigromontanus (South Africa) are two other species similar in having an elongated genital field, a box-shaped gnathosomal bay and a short medial margin of Cx-II+III. The latter species differs in a more elongated idiosoma in both sexes, a more trapezoid shape of the genital field, anteriorly less enlarged and slightly pointed, and a sligtly longer medial margin of Cx-II+III in the male (see Goldschmidt & Smit 2009). Monatractides rivalis can be distinguished by the larger dimensions in both sexes and a characteristically rounded lateral margin Cx-II (see K.O. Viets & Böttger 1974). For the time being we keep the populations from Ghana not assigned to either M. microstoma or M. procursa, mainly because the unclear taxonomy of M. microstoma. Understanding the taxonomic position of the M. microstoma - complex is not possible without additional material from a wide area and/or without the application of molecular techniques. Ejaculatory complex. In male specimens of form ''A'' the ejaculatory complex was not found but a sclerotized framework immediately above the genital field can be hardly visible (see Figs. 44 E–G). In some examined males of form “B” the ejaculatory complex was reduced to a sclerotized framework (Fig. 44 H) as in the previous form. However, in other specimens of form ''B'' a genital skeleton with more or less developed series of membranous chambers attached to a sclerotized framework can be observed (see Figs. 44 I –L, 46 C). This ejaculatory organ was shortened apically, the proximal chamber is large, the proximal and distal arms are small, the carina anterior is not developed and the apical setae is very long (Figs. 42 E–G). Figures 42 G–H illustrate the ejaculatory complex found in the largest specimens of form ''B'', possibly the last stage in the development of the ejaculatory complex. We considered several possibilities: 1) it is possible that populations (or species?) with developed or reduced ejaculatory complex exist? 2), it is possible that populations (or species?) exist in which the coalescence of membranous chambers to make up a normally developed ejaculatory complex is more or less progressed and 3), the ejaculatory complex may be more or less developed in animals of different ages. At the time being we think that differences in development of ejaculatory complex might be the result of growth. No ejaculatory complex has been described for Monatractides microstoma, nor for M. procursa. A very unusually shaped ejaculatory complex was found in Monatractides jucundus (see Fig. 42 I, figure taken from Lundblad 1952). Figure 42 K (taken from Lundblad 1952) shows the genital flap in one of Lundblad’s fixed specimen, with the apical part of the opened ejaculatory complex ending in two apical setae and a two-parted, apically pointed membranous structure, similar to condition found in ejaculatory complex of our specimens illustrated in Fig. 42 G. Malformation. In one male (form ''A'') from Kintampo Falls the anterolateral platelets of dorsal shield were asymetrically developed (Fig. 43 C). Habitat. Sandy/bouldary streams, shaded by riparian vegetation (Figs. 53 A–B, D).Published as part of Pešić, Vladimir & Smit, Harry, 2014, Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Ghana, pp. 1-80 in Zootaxa 3820 (1) on pages 54-70, DOI: 10.11646/zootaxa.3820.1.1, http://zenodo.org/record/28630

    FIGURE 4A–F. Subaturus spathulasetus n in The water mite family Aturidae Thor (Acari: Hydrachnidia) from Ghana, with the description of twelve new species

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    FIGURE 4A–F. Subaturus spathulasetus n. sp., holotype male. A = dorsum; B = venter; C = palp; D = I-leg-4–6; E = II-leg-4– 6 left; F = II-leg-4–6 right. Scale bars = 50 µm.Published as part of Smit, Harry, 2016, The water mite family Aturidae Thor (Acari: Hydrachnidia) from Ghana, with the description of twelve new species, pp. 523-543 in Zootaxa 4158 (4) on page 529, DOI: 10.11646/zootaxa.4158.4.5, http://zenodo.org/record/25747

    FIGURE 17A–F in Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo

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    FIGURE 17A–F. Photographs of ventral shield. A–B Torrenticola minuta (Lundblad, 1941): A = male, Kemantis stream; B = female, Great Lumotok stream. C–D T. neoindica n. sp., Mahua stream: C = male holotype; D = female paratype. E T. tjiwalensis (K. Viets, 1935), male, Mahua stream. F T. schilthuizeni n. sp., female holotype, Kibamabangan River.Published as part of Pešić, Vladimir & Smit, Harry, 2014, Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo, pp. 1-72 in Zootaxa 3840 (1) on page 26, DOI: 10.11646/zootaxa.3840.1.1, http://zenodo.org/record/492789

    FIGURES 2 A–F. Gretacarus occidentalis n in The water mite genus Gretacarus K. O. Viets, 1978 from Australia (Acari: Hydrachnidia, Mideopsidae: Gretacarinae), with the description of three new species

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    FIGURES 2 A–F. Gretacarus occidentalis n. sp., A–E holotype male, F paratype female. A = dorsum; B = venter; C = genital field; D = palp; E = IV-leg-4–6; F = venter. Scale bars: A–B, F = 200 µm, C–E = 50 µm.Published as part of Smit, Harry, 2016, The water mite genus Gretacarus K. O. Viets, 1978 from Australia (Acari: Hydrachnidia, Mideopsidae: Gretacarinae), with the description of three new species, pp. 159-166 in Zootaxa 4121 (2) on page 163, DOI: 10.11646/zootaxa.4121.2.5, http://zenodo.org/record/26640
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