3,479 research outputs found
A review of Ancorina, Stryphnus, and Ecionemia (Demospongiae, Astrophorida, Ancorinidae), with descriptions of new species from New Zealand waters
No full textKelly, Michelle, Sim-Smith, Carina (2012): A review of Ancorina, Stryphnus, and Ecionemia (Demospongiae, Astrophorida, Ancorinidae), with descriptions of new species from New Zealand waters. Zootaxa 3480: 1-47, DOI: 10.5281/zenodo.28235
Review of the sponge genus Penares (Demospongiae, Tetractinellida, Astrophorina) in the New Zealand EEZ, with descriptions of new species
No full textSim-Smith, Carina, Kelly, Michelle (2019): Review of the sponge genus Penares (Demospongiae, Tetractinellida, Astrophorina) in the New Zealand EEZ, with descriptions of new species. Zootaxa 4638 (1): 1-56, DOI: 10.11646/zootaxa.4638.1.
Diplopodospongia Sim-Smith & Kelly, 2011, gen. nov.
No full textGenus Diplopodospongia gen. nov. Type species: Diplopodospongia rara gen. nov. sp. nov., here designated. Diagnosis. Thinly encrusting deep-water Podospongiidae with a thin, granular, leathery, wrinkled surface that is not easily removed from the underlying crumbly choanosome. Oscules are not visible in life. Megascleres are anisoxeas, frequently centrotylote or polytylote, occasionally slightly sinuous, forming thick loose swathes that emerge from the base of the sponge and meander towards the surface, bifurcating at least once, and forming rough brushes at the surface. Microscleres are symmetrical diplospinorhabds in which the basal spine(s) and basal whorl, and apical whorl and apical spine(s), are similar in ornamentation and orientation, but one whorl might be slightly wider than the other. Protospinorhabds are linear, and shaped like a bow-tie. Microscleres form a thick crust in the ectosome and are abundant in the choanosome. Etymology. Named for the characteristic dumbbell- or diplaster-shaped microscleres, the ends of which are not always identical, but very similar (diplo =Latin for double). Remarks. Diplopodospongia gen. nov. has diplospinorhabd microscleres with linear or bow-tie-shaped protorhabds, with barely discernible tract development compared to that of other podospongiids; Diacarnus, Sigmosceptrella, Negombata, and Neopodospongia gen. nov. have a range of plumose fibre development with Diacarnus having the most complex and Neopodospongia gen. nov. having the least complex arrangement. In general terms, the diplospinorhabd microscleres are reminiscent of the dumbbell forms of spinorhabds in S. fibrosa, but the microscleres of this species are quite regular with only rare, if any, bifurcation of the spines in the apex, and are more like those of Diacarnus spp. The protospinorhabds of Diplopodospongia gen. nov. are linear as in Diacarnus, with expansions at each end, not sigmoidal as in Sigmosceptrella, Negombata, Podospongia, and Neopodospongia gen. nov., indicating quite a different development path for these microscleres. In the initial study of specimens, acanthostyles were occasionally present in almost all spicule preparations and sections of Diplopodospongia gen. nov. sp. nov., suggesting affinity with Acanthancora Topsent, 1927 (Poecilosclerida: Hymedesmiidae); two species, A. aenigma (Lundbeck 1910) and A. cyanocrypta (de Laubenfels 1930), have diplaster-like microscleres amongst the highly ornate, highly modified arcuate chelae. However, further examinations confirmed that they were foreign inclusions, and there is no evidence whatsoever of the microscleres of Diplopodospongia being derived from chelae. The diplospinorhabds are also highly reminiscent of microscleres in the hadromerid genera Diplastrella Topsent, 1918, and the ‘sclerosponge’ genera Acanthochaetetes Fisher, 1970, and Willardia Willenz & Pomponi, 1996, but all have spiraster-like microscleres in addition to diplasters, and the principal megascleres are tylostyles. The latter genera also have a basal calcareous skeleton.Published as part of Sim-Smith, Carina & Kelly, Michelle, 2011, Two new genera in the family Podospongiidae (Demospongiae: Poecilosclerida) with eight new Western Pacific species, pp. 32-54 in Zootaxa 2976 on page 46, DOI: 10.5281/zenodo.20073
Acanthancora equiformis Sim-Smith & Hickman & Kelly 2021, sp. nov.
No full textAcanthancora equiformis sp. nov. (Fig. 20) Material examined. Holotype — MCCDRS9456, Nameless Island, 0.67° S, 90.586° W, 8 m, 9 Aug 2003. Type locality. Nameless Island. Habitat and distribution. Only known from the type locality. Found growing in a small recess on a vertical rock wall; 8 m. Description. Very thinly encrusting sponge, less than 1 mm thick, densely covered in small oscules and ostia that are flush with the surface. Faint asterose drainage canals are visible on the surface. Colour in life is pale grey with tinges of white, colour in ethanol is light beige (Fig. 20A). Texture is soft. Skeleton. Unknown, specimen is too thin to section, but assumed to be consistent with the skeletal architecture in species included in this genus, as diagnosed above. Spicules. Megascleres— smooth strongyles with slightly inequiended tips, one tip being more bulbous than the other, sometimes faintly polytylote; 160 (125–182) × 4 (3–6) µm (n = 20) (Fig. 20C–D). Acanthostyles, strongly tapered with a prominent spined head; 84 (71–91) µm long (n = 20) (Fig. B). Microscleres— heavily spined isochelae; 20 (17–22) µm long (n = 20) (Fig. 20E). Etymology. Named for the form of the microscleres which resemble prancing horses (Latin adjective agreeing with Acanthancora = ‘horse-shaped’). Remarks. Only one species of Acanthancora has been described from the eastern Pacific: A. cyanocrypta (Laubenfels, 1930) from California. Acanthancora equiformis sp. nov. differs from A. cyanocrypta in that the latter species is blue in life, possesses two size classes of acanthostyles, and has smaller chelae (10–15 µm; measurements from Van Soest 2002a) with blunt spines. Genus Hemimycale Burton, 1934 Diagnosis. Thickly encrusting to massive, with striking and characteristic areolated porefields; skeleton of plumose columns of slightly inequiended strongyles, which are probably derived from tornotes; styles and oxeas may also be present. The choanosomal columns fan out near the surface and strengthen the raised porefields; no microscleres other than raphides (emended Van Soest, 2002 a and Huguenin et al. 2018).Published as part of Sim-Smith, Carina, Hickman, Cleveland & Kelly, Michelle, 2021, New shallow-water sponges (Porifera) from the Galápagos Islands, pp. 1-71 in Zootaxa 5012 (1) on pages 44-45, DOI: 10.11646/zootaxa.5012.1.1, http://zenodo.org/record/515806
Penares angeli Sim-Smith & Hickman & Kelly 2021, sp. nov.
No full textPenares angeli sp. nov. (Fig. 16) Material examined. Holotype — MCCDRS9410, Maria Elena Islet, Pinzón Island, 0.592° S, 90.684° W, 13 m, 20 Jan 2003. Type locality. Maria Elena Islet, Pinzón Island. Habitat and distribution. Only known from type locality, growing in a rock crevice; 13 m. Description. Massive, lobose sponge, with a single, large oscule, 5–8 mm in diameter, on the apex of each lobe. The surface is smooth, almost fleshy. Colour in life dark grey-brown on top fading to white at the base (the colour of the underlying choanosome), colour in ethanol medium brown fading to tan at the base (Fig. 16A–B). Texture firm, compressible. Skeleton. The cortex is approximately 500 µm thick and comprises a dense crust of microrhabds and microxeas that are lying in no particular orientation. A thin, dense layer of microrhabds exists on the surface of the sponge. Regular pores pierce the cortex. Orthotriaenes are arranged perpendicular to the surface with their cladome at the base of the cortex (Fig. 16C). Oxyasters, microxeas, and microrhabds are abundantly scattered throughout the choanosome (Fig. 16D). Spicules. Megascleres— oxeas, moderately long and stout, often curved; 1808 (1240–2672) × 31 (19–44) µm (n = 20). Orthotrianes are moderately long with a curved cladome; rhabdome length 1270 (926–1668) µm, protoclad length, 283 (226–367) µm, cladome width, 530 (389–640) µm (n = 20) (Fig. 16E). Microscleres— oxyasters, small with several long, slender acanthose rays; 14 (6–26) µm in diameter (n = 20) (Fig. 16H). Microxeas are curved and centrotylote, 135 (101–191) × 7 (4–10) µm (n = 20) (Fig. 16F). Microrhabds are centrotylote and lightly acanthose; 48 (29–88) × 4 (2–5) µm (n = 20) (Fig. 16G). Etymology. Named after Angel Chiriboga for his invaluable assistance with the collection and photography of the sponges. Remarks. Four species of Penares have been previously described from the Galápagos Islands: P. saccharis (De Laubenfels, 1930), P. scabiosus Desqueyroux-Faúndez & Van Soest 1997, P. foliaformis Wilson, 1904, and P. apicospinatus Desqueyroux-Faúndez & Van Soest 1997. The first three species possess dichotriaenes whereas P. angeli sp. nov. possesses orthotriaenes. Penares apicospinatus has two size classes of oxyasters and oxeas that are much longer (2496 (1426–3130) µm; measurements from Desqueyroux-Faúndez & Van Soest 1997) than those of P. angeli sp. nov.Published as part of Sim-Smith, Carina, Hickman, Cleveland & Kelly, Michelle, 2021, New shallow-water sponges (Porifera) from the Galápagos Islands, pp. 1-71 in Zootaxa 5012 (1) on pages 36-37, DOI: 10.11646/zootaxa.5012.1.1, http://zenodo.org/record/515806
Stryphnus poculum Kelly & Sim-Smith, 2012, sp. nov.
No full textStryphnus poculum sp. nov. (Fig. 2 A, 3, 6A–D, 8) Material examined. Holotype ― NIWA 62046: 3 nautical miles east of North Cape, north-eastern tip of Northland, 34.412 ° S, 173.149 ° E, RV Kaharoa, 133–210 m, 19 Apr 1999, additional vouchers of the holotype are in the CRRF reference collection (0 CDN 6695 -M) and at the USNM (USNM 1182994). Type locality. North Cape. Distribution. Known only from the type locality. Description. Tall vase-shaped sponge, 200 mm high x 120 mm wide, with a 40 mm thick body wall (Fig. 2 A). Ectosome is 1.5–3 mm thick and clearly differentiated from the underlying choanosome. Margin of bowl is thinner than the body wall and slightly undulating. Surface is smooth, granular to the touch. Oscules and ostia not visible with unaided eye, texture is firm, barely compressible, interior highly siliceous, can be torn relatively easily from margin to base. Colour in life is light grey, interior tan. Colour in ethanol medium brown, interior tan. Skeleton. Ectosome is 1500–3000 µm deep, cavernous, diaphanous, well differentiated from the underlying choanosome by a wispy 50–100 μm thick layer of granular cells (Fig. 3). Outer ectosome has a thin, dispersed crust of amphisanidasters, below which these are only lightly scattered. Dichotriaenes form a rough radiating layer in the ectosome, with cladomes protruding slightly beyond the surface. Choanosome is dense and heavily pigmented, dichotriaenes also form an irregular, roughly radial to oblique layer at the choanosome/ectosome boundary (Fig. 3), interspersed with oxeas. Microscleres are sparsely scattered throughout the choanosome. Spicules. Megascleres (Fig. 6 A–B) are oxeas (Fig. 6 A), very large, stout, slightly curved, 2004 (1825–2552) x 34 (21–50) μm; dichotriaenes (Fig. 6 B) with stout, conical rhabdome, 723 (453–897) μm long, cladome width 347 (183–461) μm, protoclads 82 (63–109) μm long, protruding forward at an angle of approximately 45 °, deuteroclads slightly longer than protoclads, 100 (44–154) μm. Microscleres (Fig. 6 C–D) are oxyasters (Fig. 6 C), choanosomal, with 4–6 very stout acanthose rays, 35 (28– 43) μm diameter; amphisanidasters (Fig. 6 D), with a short thick axis and numerous short conical acanthose spines protruding in a cluster from each end, or from along the shaft. Spines are frequently of different lengths, 11 (10–13) μm. Substrate, depth range, and ecology. Attached to deep rocky reef interspersed with sandy substrate, between 133– 210 m. Etymology. Named for the vase-shaped morphology of the species (poculum = Latin for cup or bowl). Remarks. Stryphnus poculum sp. nov. has the same general skeletal, cellular arrangements, and spicule categories as the type species S. niger from Port Jackson, South Australia; it has a well demarcated diaphanous ectosome, short-shafted dichotriaenes, large oxeas, oxyasters and amphisanidasters. However, the spicule dimensions differ considerably between the two species; S. poculum sp. nov. has smaller oxeas and amphisanidasters, but larger dichotriaenes and oxyasters than S. niger (Table 3). Furthermore, the dichotriaenes of S. poculum sp. nov. are present in both the ectosome and choanosome, unlike in S. niger where they are confined to the ectosome. The two species are easily separated on field characters; S. poculum sp. nov. is a tall, light greyish brown cup-shaped sponge whereas S. niger and all other known species, with the exception of S. ponderosus (Bowerbank 1866) are massive, lobate, or thickly encrusting. Stryphnus ponderosus from the Arctic and Atlantic coasts of Europe sometimes forms a cup or goblet but its amphisanidasters have longer, thinner spines than those of S. poculum sp. nov.Published as part of Kelly, Michelle & Sim-Smith, Carina, 2012, A review of Ancorina, Stryphnus, and Ecionemia (Demospongiae, Astrophorida, Ancorinidae), with descriptions of new species from New Zealand waters, pp. 1-47 in Zootaxa 3480 on page 21, DOI: 10.5281/zenodo.28235
Neopodospongia bergquistae Sim-Smith & Kelly, 2011, gen. nov.
No full textNeopodospongia bergquistae gen. nov. sp. nov. (Fig. 1 E, 4 I –M) Material examined. Holotype ― NIWA 62016, Poor Knights Islands, 35.467 ° S, 174.733 ° E, 18 m, collected by Dr Christopher N. Battershill, on SCUBA, 25 Oct 1984. Type location. Poor Knights Islands. Distribution. Known only from type location. Description. Thinly encrusting sponge, about 3–5 mm thick, spreading in small patches on rock walls. Surface is smooth and glossy, with an open, lacy appearance, and is easily detached in life. Oscules with elevated rims about 1–2 mm high and 2–3 mm in diameter are scattered over the surface of the sponge (Fig. 1 E). Texture is leathery. Colour of the ectosome in life is translucent white, the underlying choanosome is dark beige. Skeleton. Ectosome is 550–650 mm thick, translucent, densely collagenous, and clearly differentiated from the underlying choanosome. Aciculospinorhabds are lightly packed in the ectosome and are arranged more or less vertically with the apical spire pointing towards the sponge surface. They do not extend beyond the surface. Choanosome is thick, fleshy, composed of very thick tracts of megascleres, around 300 µm thick, that meander vertically from the collagenous basal layer, radiating towards the surface and dividing into numerous fine secondary fibres that expand again into delicate fans that terminate below the surface. Aciculospinorhabds are only lightly dispersed throughout the choanosome. Megascleres (Fig. 4 I, Table 2)― Strongyloxeas, faintly polytylote: 405 (355–457) x 6 (5–8) µm. Microscleres (Fig. 4 J–M, Table 2)― Aciculospinorhabds, asymmetrical, with clearly differentiated ends. The basal whorl is composed of numerous spines that extend obliquely away from the shaft towards the base of the spicule. The median whorl consists of a number of smooth bifurcating spines arranged in pairs that encircle the shaft horizontally (Fig. 4 M). The apical whorl is a ring of bifurcate spines that bifurcate in the vertical plane. The apex is a thick spire with a distal ring of small spines. The protoaciculospinorhabd is sigmoidal in shape (Fig. 4 L). The spicules are strictly aciculospinorhabds in one size category; 35 (28–40) µm long x 17 (15–22) µm wide. Substrate, depth range, and ecology: Encrusting a flat rock surface at 18 m. Etymology. Named after Dame Professor Patricia Rose Bergquist of New Zealand (deceased), for her enormous and enduring contribution to our knowledge of sponges in general, and to our knowledge of New Zealand sponges in particular. Remarks. Apart from their disparate type localities, N. bergquistae sp. nov. is difficult to differentiate from N. pagei sp. nov. in life; Neopodospongia bergquistae sp. nov. is slightly thinner with a differential colouration of the choanosome and ectosome. However, the two species can be reliably differentiated by their skeletal architecture and spicule complement. The ectosome of N. bergquistae sp. nov. is thicker and more collagenous than that of N. pagei sp. nov., which is packed and granular with projecting microscleres. The fibre architecture of N. bergquistae sp. nov. is more intricate than that of N. pagei sp. nov., forming fine sprays of secondary fibres quite deeply within the choanosome. The megascleres of N. bergquistae sp. nov. are shorter and thinner than those of N. pagei sp. nov., and the aciculospinorhabds, while being similar in morphology to those of N. pagei sp. nov. are markedly shorter and narrower. There is only one discernable category of aciculospinorhabds in N. bergquistae sp. nov. and they are considerably more spinose than those of N. pagei sp. nov.Published as part of Sim-Smith, Carina & Kelly, Michelle, 2011, Two new genera in the family Podospongiidae (Demospongiae: Poecilosclerida) with eight new Western Pacific species, pp. 32-54 in Zootaxa 2976 on pages 43-45, DOI: 10.5281/zenodo.20073
Diplopodospongia macquariensis Sim-Smith & Kelly, 2011, gen. nov.
No full textDiplopodospongia macquariensis gen. nov. sp. nov. (Fig. 1 H, 5 L–M) Material examined. Holotype ―QM G 331915: NIWA station TAN0803/ 89, Seamount 8, south of Macquarie Island, Macquarie Ridge (Australian EEZ), 55.381 ° S, 158.427 ° E, 504–637 m, 15 Apr 2008. Paratypes ―QM G 331916: NIWA Stn TAN0803/ 102, Seamount 9 Hjort, south of Macquarie Island, Macquarie Ridge (Australian EEZ), 56.242 ° S, 158.462 ° E, 790–1025 m, 16 Apr 2008; QM G 331917: NIWA Stn TAN0803/ 98, Seamount 9 Hjort, south of Macquarie Island, Macquarie Ridge (Australian EEZ), 56.246 ° S, 158.506 ° E. 676–750 m, 16 Apr 2008. Type location. Seamount 8, south of Macquarie Island, Macquarie Ridge (Australian EEZ). Distribution. Seamounts 8 and 9 in the Macquarie Ridge. Description. Very thinly encrusting sponge, <1 mm thick, growing on dead coral or rock (Fig. 1 H). Surface smooth, soft and friable to the touch, no visible oscules; surface wrinkled. Colour in ethanol beige to light mustard brown. Skeleton. Ectosome a dense layer of diplospinorhabds, which are generally vertically disposed. The choanosome consists of loose bundles of anisoxeas that emerge from the base to form rough brushes in the ectosome. Microscleres are scattered throughout the choanosome. Megascleres (Fig. 5 L, Table 3)― Anisoxeas, straight, centrally thickened, centrally tylote; 442 (394–477) x 9 (7–11) µm. Microscleres (Fig. 5 M, Table 3)― Diplospinorhabds, symmetrical, double-ended, with generally symmetrical ends that are separated by a short shaft. The base and apex are single spines, and the basal and apical whorls consist of four long smooth, bifurcate spines, separated by a smooth thick shaft. Protodiplospinorhabds were rare. Those that were found under light microscopy were linear, shaped like a bow-tie; 27 (20–31) µm long x 23 (20–27) µm wide. Substrate, depth range, and ecology: Growing on the surface of oculinid corals between 500 and 1000 m. Etymology. Named for the type location, the Macquarie Ridge. Remarks. Diplopodospongia macquariensis gen. nov. sp. nov. is superficially quite similar to D. rara gen. nov. sp. nov. in the shape of the diplospinorhabds. However, there are several key differences that we believe are not ecophenotypic variations. The megascleres of D. macquariensis gen. nov. sp. nov. are considerably longer and different in shape; they are centrally thickened with polytylote or centrotylote swellings, and they are very straight, whereas those of D. rara gen. nov. sp. nov. are noticeably curved, almost sinuous. While the diplospinorhabds of D. macquariensis gen. nov. sp. nov. are similar in general style of ornamentation, the bifurcate spines are much longer, and the shaft shorter, making the spicule less compact. The apex and base are frequently only composed of a single robust spine, compared to three spines in D. rara gen. nov. sp. nov. The gross morphology and colouration of the two species also differs; D. rara gen. nov. sp. nov. is more thickly encrusting and white in ethanol, whereas D. macquariensis gen. nov. sp. nov. is more thinly encrusting and beige in ethanol.Published as part of Sim-Smith, Carina & Kelly, Michelle, 2011, Two new genera in the family Podospongiidae (Demospongiae: Poecilosclerida) with eight new Western Pacific species, pp. 32-54 in Zootaxa 2976 on page 48, DOI: 10.5281/zenodo.20073
Dragmacidon hendersoni Sim-Smith & Hickman & Kelly 2021, sp. nov.
No full textDragmacidon hendersoni sp. nov. (Fig. 13) Material examined. Holotype — MCCRDS9412, Nameless Island, 0.670° S, 90.586° W, 9 m, 9 Aug 2003. Type locality. Nameless Island; 9 m. Habitat and distribution. Only known from the type locality. Found in a recess on a vertical wall growing on rock; 9 m. Description. Thinly encrusting sponge, 3 mm thick, with a minutely ridged and punctate surface. A few small oscules are scattered over the surface of the sponge. Texture is compressible, surface is faintly hispid. Colour in life is bright yellow, colour in ethanol is tan (Fig. 13A). Skeleton. Choanosomal skeleton is loosely plumoreticulate, with delicate, multispicular primary tracts and vague secondary tracts/connections. Bundles of styles protrude through the surface of the sponge, in tufts, points uppermost. There is no special ectosome (Fig. 13B). Spicules. Megascleres— Style I (Fig. 13C), small, slender and slightly curved; 231 (209–248) × 6 (4–7) µm (n = 20). Style II (Fig. 13D), very slender and often strongly curved or slightly sinuous; 197 (127–334) × 2.4 (1.2–3.4) µm (n = 20). Etymology. Named after Scott Henderson, who accompanied co-author Cleveland Hickman on numerous trips to the Galápagos from 1983–2016. He assisted Cleve in a variety of roles from local negotiator and logistics organiser, divemaster during harrowing high-current dives, trusted friend and host on his Santa Cruz coffee farm. Of the 149 students Cleve led on 12 Galápagos student field trips, Scott was the only one to make Galápagos his home and life-long conservation project. Remarks. Dragmacidon hendersoni sp. nov. differs considerably from D. raeae sp. nov. by: live colouration (yellow vs bright red); skeleton (compact plumoreticulate vs loosely plumoreticulate to plumose); texture (firm vs compressible); and the possession of two size categories of styles, with both categories being more slender and sinuous than those of D. raeae sp. nov. It also differs considerably from the previously recorded D. oxeon which has much longer megascleres. It differs from all other tropical/subtropical eastern Pacific species by the lack of oxeas. Table 4 provides useful details on all known species of Dragmacidon, many of which appear to fall loosely into two groups: those that were previously identified as species of Pseudaxinella Schmidt, 1875 [D. raeae sp. nov. and other species that form firm dome-shaped sponges, with compact plumoreticulate skeletons that often lack trichodragmata, such as D. australe (Bergquist, 1970), D. coccineum (Keller, 1891), D. debitusae (Hooper & Lévi, 1993)]; and those that have a looser, more plumose skeleton with longer, more sinuous megascleres and often raphides in trichodragmata, resulting in a softer, less firm sponge, such as D. oxeon, D. mutans (Sarà, 1978), D. tumidum (Dendy, 1897), and D. decipiens (Wiedenmayer, 1989). The addition of two species from the Galápagos Archipelago, each of which falls loosely into one or the other group, may lend additional data for future detailed analysis of Dragmacidon, a genus of convenience into which all species commonly previously recorded as Pseudaxinella (a junior synonym of Axinella), were assigned by Alvarez & Hooper (2002: 734).Published as part of Sim-Smith, Carina, Hickman, Cleveland & Kelly, Michelle, 2021, New shallow-water sponges (Porifera) from the Galápagos Islands, pp. 1-71 in Zootaxa 5012 (1) on pages 30-31, DOI: 10.11646/zootaxa.5012.1.1, http://zenodo.org/record/515806
FIGURE 7 in Two new genera in the family Podospongiidae (Demospongiae: Poecilosclerida) with eight new Western Pacific species
No full textFIGURE 7. New Zealand fossil sponge spicules collected from the late Eocene–early Oligocene Oamaru Diatomite (c. 43–34 Ma). Figures reproduced from Plates 11 and 12 of Hinde and Holmes (1892).Published as part of Sim-Smith, Carina & Kelly, Michelle, 2011, Two new genera in the family Podospongiidae (Demospongiae: Poecilosclerida) with eight new Western Pacific species, pp. 32-54 in Zootaxa 2976 on page 51, DOI: 10.5281/zenodo.20073
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