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    Amputoearinus fernandezi Sharkey

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    Amputoearinus fernandezi Sharkey (Fig. 2 d) Diagnosis: This species can be distinguished from all other species of Amputoearinus by the following characters; forewing infuscate with yellow patch in basal quarter and yellow band in apical 3 rd quarter; propodeum with distinct carinae. Description: Holotype Ψ Body length: 6.0 (5.8–6.1) mm. Color: body yellow except melanic as follows: flagellomeres, pedicel, scape laterally, mid and hind tibia distally, midtarsus distally, hind tarsus, ovipositor sheaths; posterior three terga with black patches laterally; forewing infuscate with yellow patch in basal quarter and yellow band in apical 3 rd quarter (cf. Fig. 4 e). Head: antenna with 37 (37–40) flagellomeres Mesosoma: scutellar sulcus lacking longitudinal carina (cf. Fig. 6 d); projection of propleuron blunt, not dorsoventrally flattened, and without ridges (cf. Fig. 6 f); median areola of mesosoma well defined with sharp carinae laterally and posteriorly (cf. Fig. 7 c); propodeum mostly smooth with pair of short carinae fused anteriorly to appear as an inverted v-shape (cf. Fig. 5 g); margin between metepimeron and metepisternum smooth, lacking transverse carinae (cf. Fig. 7 b); mesopleural suture with transverse carinae; carina bordering subpronope posteriorly, extending to anterior margin of pronotum (cf. Fig. 6 a); length of midtibial spur 0.4mm; length of midbasitarsomere 0.5mm; midtibia with 3–4 pegs at midlength, lacking pegs apically; hind tibia with 5 pegs apically; forewing length 6.0mm Metasoma: median syntergite 2 + 3 length 1.4mm, width 1.2mm. Variation: Most specimens have a smooth propodeum with sculpture reduced to a small protuberance anteromedially. Specimen Examined: HOLOTYPE. Ψ, GUYANA, Dubulay Ranch, 5 ° 40.95 ”N 57 ° 51.52 ”W, Malaise, Feb–May 1999, Sharkey & Brown (HIC). PARATYPES. BRAZIL, Ψ, Para, Baker, [1 ° 26 ’ 60 ”S 48 ° 28 ’ 60 ”W], (USNM). GUYANA, Ψ, Dubulay Ranch, 5 ° 40 ’ 9.45 ”N 57 ° 51 ’ 52.4 ”W, Malaise Trap, 16–23.iv. 1999, M. Sharkey & B. Brown, (HIC). SURINAM: Ψ, Paramaribo, Plantation Ma Retraite, [5 ° 49 ’ 60 ”N 55 ° 10 ’W], Jan. 10, 1964, sweep forest, D.C. Geijskes, (head detached and placed on triangle), (RMNH). Ψ, Paramaribo, [5 ° 49 ’ 60 ”N 55 ° 10 ’W], June 10 –13, 1963, Malaise trap, J.v. d Vecht., (RMNH). TRINIDAD: ♂, BWI [British West Indies], Maracas, [10 ° 40 ’ 60 ”N 61 ° 23 ’ 60 ”W], July- 13-1953, Collector F.J. Simmonds, (head missing), (AEI). Ψ, Asa Wright N.C. [Nature Center], [10 ° 37 ’ 60 ”N 61 ° 16 ’ 60 ”W], Jan. 15, 1981, G.E. Bohart, (EMUS).Published as part of Lindsay, Cheryl & Sharkey, Michael, 2006, Revision of the genus Amputoearinus (Hymenoptera: Braconidae: Agathidinae) with fourteen new species, pp. 1-27 in Zootaxa 1329 on pages 9-10, DOI: 10.5281/zenodo.17417

    Oroceguera andersoni Seltmann and Sharkey, sp. n.

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    Oroceguera andersoni Seltmann and Sharkey, sp. n. Holotype: Female, COSTA RICA. Guanacaste Providence, Guanacaste Conservation Area, Pitilla Field Station, 600m., Berlese leaf litter, low montane rain forest, 2.v. 1995, R.S. Anderson. Deposited in the Hymenoptera Institute Collection, University of Kentucky. Etymology: The species name is in honor of the collector, Robert S. Anderson. Holotype Description: Body: length 1.27mm; generally ant-like in appearance. Color: head orange to brown; antenna orange-brown basally becoming light yellow apically; mesosoma dark orange to brown, carinae pronounced and darker brown than rest of mesosoma; legs light yellow; metasoma orange to brown basally becoming yellow-white apically; ovipositor pale yellow. Head: entirely smooth with sparse hairs on the face; eye small, malar space subequal to eye height; ocelli absent; antenna broken after the 10 th flagellomere (7 th on the other). Mesosoma: wingless; tegula absent; scutellum raised; propodeum subequal in length to mesonotum + metanotum, coriaceous-rugose. Legs: foretibia with single row of seven stout spines along anterior edge; dorsal surface of hind coxa rugose. Metasoma: first tergum with distinct irregular median and lateral longitudinal carinae and a basal transverse ridge that defines a concave area anteriorly, concave area fits into a bulge on propodeum; remainder of terga smooth, with few sparse setae; ovipositor about 0.4 times length of body. Male: Unknown.Published as part of Seltmann, Katja & Sharkey, Michael, 2007, A new genus and species of apterous Doryctinae (Hymenoptera: Braconidae) from Costa Rica, pp. 17-24 in Zootaxa 1415 on page 19, DOI: 10.5281/zenodo.27369

    FIGURE 3. A in Description of three new species of Helconichia Sharkey & Wharton (Hymenoptera: Braconidae: Ichneutinae) with a revised key to all species

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    FIGURE 3. A. Lateral habitus of female holotype of Helconichia trichiops Sharkey & Wharton. B. Dorsal habitus of female holotype of H. trichopteryx Sharkey & Wharton. Inset is lateral view of head, pronotum, and mesopleuron.Published as part of Sharanowski, Barbara J. & Sharkey, Michael J., 2007, Description of three new species of Helconichia Sharkey & Wharton (Hymenoptera: Braconidae: Ichneutinae) with a revised key to all species, pp. 45-53 in Zootaxa 1502 on page 50, DOI: 10.5281/zenodo.27380

    Gelastagathis frosti Sharkey, NEW SPECIES

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    Gelastagathis frosti Sharkey NEW SPECIES (Fig. 4) Description ♂. Length. 5.3 mm. Color: Testaceous except for terminal metasomal segments black, antenna melanic, wings infuscate, propodeum melanic. Head: in frontal view 1.7 x wider than high (measured from top to apex of clypeus). Antennal flagellum with 27 flagellomeres. Mesosoma: Notaulus complete, smooth anteriorly, weakly pitted posteriorly. Hind tibia with 1 apical peg. Propodeum entirely rugose with two irregular longitudinal carinae defining a medial region. Metasoma: First median tergite with weak longitudinal striae. Second median tergite with a curved transverse groove. Material examined. Known only from the holotype ♂: USA, Massachusetts, Natick, Sept. 1935, C.A. Frost. Approximately 42 ° 17 ’N, 71 ° 20 ’W and 50 m. elevation. (USNM). Etymology. Named after the collector of the type specimen.Published as part of Sharkey, Michael J. & Chapman, Eric G., 2015, The Nearctic genera of Agathidinae (Hymenoptera: Braconidae) with a phylogenetic analysis, illustrated generic key, and the description of three new genera, pp. 49-72 in Zootaxa 4000 (1) on pages 61-62, DOI: 10.11646/zootaxa.4000.1.2, http://zenodo.org/record/28924

    Austroearinus Sharkey 2006, n. gen.

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    Austroearinus Sharkey n. gen. (Fig. 1b) Diagnosis Austroearinus can be distinguished from all other agathidine genera with the following combination of characters: tarsal claws with a basal tooth; mesoscutum unsculptured and lacking notauli; hind coxal cavities open; propleuron without projection; ovipositor as long as or longer than metasoma. Description Head: Carinae on frons lacking; interantennal space lacking two prominences separated by shallow groove; gena not extended ventroposteriorly into sharp prominence; labial palp with four segments, third segment not reduced; apical antennomere acute; antennal insertion bordered anteriorly, medially, and laterally by moderately elevated ridge. Mesosoma: Mesoscutum smooth; notauli lacking; posteroscutellar depression absent; median areola of metanotum well defined with sharp carinae laterally and posteriorly; propodeum from completely smooth to carinate with anterior transverse carina and with pair of longitudinal carinae enclosing large medial cell; propleuron mildly convex to flat lacking ventromedial prominence; propodeal pseudosternite poorly developed and not separating hind coxal cavities from metasomal foramen. Legs: Foretibia lacking pegs; foretarsal claws with quadrate basal lobe; basal lobe with tuft of pectination basally (c.f. Fig. 2c); midtibia with apical and medial pegs; hind tibia with apical pegs. Wings: RS + M vein incomplete and not tubular throughout, though often more complete than most genera of Agathidinae (Fig.1b); second submarginal cell triangular to weakly quadrate; 3RSb straight; hind wing r and r­m crossveins absent; CUb sometimes present as a nebulous or spectral vein. Metasoma: Median tergites smooth lacking sculpture; median tergite 1 with pair of longitudinal carinae diverging distally; median syntergite 2 + 3 with 1 or 2 transverse depressions, distinct grooves absent; ovipositor mildly decurved, longer than metasoma (slightly shorter than the metasoma in one species) usually as long as body. Type species Bassus rufofemoratus (Muesebeck) (Fig. 1b). Distribution New World, primarily Neotropical. One species, Austroearinus rufofemoratus n. comb., is widespread in central and eastern USA and through Central America, but the genus is far more diverse in warm tropical areas. Species diversity At least two species formerly placed in other agathidine genera belong here: Austroearinus rufofemoratus, originally placed in Bassus (Muesebeck 1927), and subsequently transferred to Earinus (Sharkey 2004); Austroearinus unicolor (Schrottkey) n. comb., originally placed in Orgilus (Schrottkey 1902), but more recently considered as a member of the genus Bassus. The genus is extremely species rich. I estimate there to be more than 100 species and perhaps several times this number. Many species also appear to be common, for example, there has been intensive Malaise trap sampling in La Selva, Costa Rica for more than a decade (http://viceroy.eeb.uconn.edu/ALAS/ALAS.html) and, of all species of Agathidinae, A. chrysokeras has been the most frequently collected. Biology (Dan Janzen, pers. comm.) A specimen of A. rufofemoratus from Guanacaste Province, Costa Rica, was reared from a microlepidopteran, only identified as Tortricidae, feeding on the mature leaves of Amyris pinnata (Rutaceae) and living in a light silk web. The wasp larva emerged from the last instar caterpillar on 6 April 2002 and spun a strong white cocoon tightly glued to the leaf surface. This is unlike other known agathidines which spin inside the caterpillar cocoon. The wasp eclosed 9 days later, which means that all of this happened in the full dry season. The site is 280 m elevation, Lat 11.04562 Long ­85.45742, and locally known as Puente Mena in the Del Oro property adjacent to the north boundary of the Area de Conservacion Guanacaste. The site lies on the interface between ACG dry forests and ACG rain forest. The entire record may be found at http://janzen.sas.upenn.edu, refer to specimen number 02­SRNP­14223. Etymology From the Latin australis which means southern, referring to the southern distribution of this genus and its morphological similarity to Earinus.Published as part of Sharkey, Michael, 2006, Two new genera of Agathidinae (Hymenoptera: Braconidae) with a key to the genera of the New World, pp. 37-51 in Zootaxa 1185 on pages 45-46, DOI: 10.5281/zenodo.264601

    Amputoearinus boringi Lindsay and Sharkey, sp.n.

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    Amputoearinus boringi Lindsay and Sharkey sp.n. (Fig. 2 b, 6 c) Diagnosis: This species can be distinguished from all other species of Amputoearinus by the following characters; longitudinal carina of scutellar sulcus present; median areola of mesosoma depressed medially with sharp carinae laterally and posteriorly; projection of propleuron blunt; carina bordering subpronope posteriorly, extending to anterior margin of pronotum. Description: Holotype Ψ Body length: 7.3mm. Color: body orange except for melanic as follows: antenna, hind tarsus, mid and hind tibia with melanic patch distally; forewing with infuscate band apically (cf. Fig. 4 f). Head: antenna with 43 flagellomeres. Mesosoma: scutellar sulcus with weak longitudinal carina (Fig. 6 c); projection of propleuron blunt (cf. Fig 6 f.); median areola of mesosoma slightly depressed medially, with carinae laterally and posteriorly and transverse carina anteriorly, interior of areola rugulose (cf. Fig. 7 c); propodeal sculpture reduced to small protuberance anteromedially (cf. Fig. 5 f); mesopleural suture with weak and reduced carinae along its length; margin between metepimeron and metepisternum smooth, lacking transverse carinae (cf. Fig. 7 b); carina bordering subpronope posteriorly, extending to anterior margin of pronotum (cf. Fig. 6 a); length of midtibial spur 0.6mm; length of midbasitarsomere 0.7mm; midtibia with 3 pegs at midlength; 0–1 peg(s) apically; hind tibia with 3–4 pegs apically; forewing length 7.1mm. Metasoma: median syntergite 2 + 3 length 1.6mm, width 1.5mm. Male: Unknown Etymology: in honor of Andy Boring. Specimen examined: HOLOTYPE. Ψ, COLOMBIA, [Valle del] Cauca, PNN [Parque Nacional Natural] Gorgona, El Saman, 2 º 58 ’N 78 º 11 ’W, 5m, Malaise, 7–20.ii. 2001, H. Torres, Leg. M. 1364, (right hind leg broken off after hind tibia), (IAvH).Published as part of Lindsay, Cheryl & Sharkey, Michael, 2006, Revision of the genus Amputoearinus (Hymenoptera: Braconidae: Agathidinae) with fourteen new species, pp. 1-27 in Zootaxa 1329 on page 8, DOI: 10.5281/zenodo.17417

    Oroceguera Seltmann and Sharkey

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    Oroceguera Seltmann and Sharkey, gen. n. Type species: Oroceguera andersoni sp. n. Etymology: The genus name is a combination of the Spanish words for “gold” (oro) and “blindness” (ceguera) and refers to the golden-yellow coloration, reduced eyes and lack of ocelli. Diagnosis: Only the holotype female is known (Fig. 1 a). They are easily distinguished from all other wingless genera of Doryctinae by the high insertion of the metasoma and the complete absence of wings which are not indicated, even as small scales or wingpads. (Fig. 1 d). Description: Small size, 1.27 mm; female wingless. Head: subcubical flattened and glabrous dorsally; face with median keel and few long setae; eye small, longer than wide; ocelli absent; occipital carina meeting hypostomal carina; postgenal bridge absent; first flagellomere subequal but slightly shorter than second. Mesosoma: highly modified and lacking tegula; pronotal collar equal in length to first flagellomere, pronotal carina distinct and raised; notauli present but short; scutellum conical and with distinct longitudinal carinae apically, coriaceous-rugose with tubercles on either side laterally; scuto-scutellar suture present; epicnemial carina distinct; postpectal carina present but less pronounced; sternaulus distinct; mesopleuron and metaplueron concave; deep sulcus dividing metanotum and propodeum; propodeum concave. Legs: foretibia and midtibia with numerous obvious and stout spines along anterior edge; hind coxa without basal tubercle or tooth. Metasoma: inserted high above hind coxal cavities and separated from them by a wide propodeal bridge; first metasomal tergite petiolate, acrosternite long, 0.5 times as long as first metasomal tergum (Belokobylskij et al., 2004), with distinct dorsope, without basolateral processes, spiracle found at about 0.5 length; longitudinal carinae present on tergum one, terga 2–8 without sculpture; distinct groove between second and third median tergites lacking; laterotergites not present on terga 2–4. Illustrations: Additional images describing Oroceguera are found on MorphBank biological image database (http://morphbank.net/Show/?id= 101864). Distribution: The sole specimen was sifted from leaf litter from a Costa Rican montane rain forest at 600m. Biology: Unknown, but see discussion below.Published as part of Seltmann, Katja & Sharkey, Michael, 2007, A new genus and species of apterous Doryctinae (Hymenoptera: Braconidae) from Costa Rica, pp. 17-24 in Zootaxa 1415 on pages 18-19, DOI: 10.5281/zenodo.27369

    Amputoearinus galbus Lindsay and Sharkey, sp.n.

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    Amputoearinus galbus Lindsay and Sharkey sp.n. (Figs. 2 f, 5 e, 7 a, 6 b) Diagnosis: This species can be distinguished from all other species of Amputoearinus by the following character; median areola of propodeum present but lateral edges weak at midlength. Description: Holotype Ψ Body length: 6.8 mm. Color: yellow except melanic as follows: antenna; black patch posterior to eye and anterior of head; hind coxa, hind trochanter, hind femur ventrally, midfemur apically; ovipositor sheaths; apex of forewing infuscate (cf. Fig. 4 f). Head: antenna with 43 flagellomeres. Mesosoma: scutellar sulcus lacking longitudinal carina (cf. Fig. 6 d); projection of propleuron blunt (cf. Fig. 6 f); carina bordering subpronope posteriorly, not extending to anterior margin of pronotum(Fig. 6 b); median areola of mesosoma well defined with sharp carinae laterally and posteriorly (cf. Fig. 7 c); median areola of propodeum present but lateral edges weak at mid-length, transverse carinae located anteriorly (Fig. 5 e); margin between metepimeron and metepisternum with 3 transverse carinae (Fig. 7 a); mesopleural suture with strong transverse carinae; midtibia with 1–2 pegs at midlength, 1 peg apically; forewing length 5.8mm. Metasoma: median syntergite 2 + 3, length 1.2mm, width 1.2mm. Male: Unknown Etymology: from the Latin galbus meaning yellow. Specimen Examined: HOLOTYPE. Ψ, COLOMBIA, Choco, PNN [Parque Nacional Natural] Utria Cop. Manual, Sendero Menglar, 6 º01’01”N, 7 º 20 ’ 52 ”W, 28.iii. 1998, C. Londoño, (front legs broken off at the coxa, left midleg broken off at tarsus, left hind leg broken off at femur, right hind leg broken off at trochanter, right antenna broken), (IavH 3381).Published as part of Lindsay, Cheryl & Sharkey, Michael, 2006, Revision of the genus Amputoearinus (Hymenoptera: Braconidae: Agathidinae) with fourteen new species, pp. 1-27 in Zootaxa 1329 on pages 12-13, DOI: 10.5281/zenodo.17417

    Amputoearinus sharanowskiae Lindsay and Sharkey, sp.n.

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    Amputoearinus sharanowskiae Lindsay and Sharkey sp.n. (Figs. 4 b, 4 f, 7 b) Diagnosis: This species can be distinguished from all other species of Amputoearinus by the following characters; forewing yellow with infuscate band apically; median areola of mesosoma not depressed medially and lacking sharp carinae laterally and posteriorly; projection of propleuron blunt. Description: Holotype Ψ Body length: 6.0mm. Color: body yellow except melanic as follows: head, antenna, propleuron, fore, mid, hind femur, midtibia distally, midtarsus, hind coxa, trochanter, hind tibia basally and distally, hind tarsus, posterior three terga; apex of forewing infuscate (Fig. 4 f). Head: antenna with unknown number of flagellomeres, antennae broken off at scape, one antenna placed on point and broken after 27 th flagellomere. Mesosoma: scutellar sulcus lacking longitudinal carina (cf. Fig. 6 d); projection of propleuron blunt (cf. Fig. 6 f); median areola of mesosoma not depressed medially and lacking sharp carinae laterally and posteriorly (cf. Fig. 7 d); propodeal sculpture reduced to small protuberance anteromedially (cf. Fig. 5 f); mesopleural suture with weak carinae; margin between metepimeron and metepisternum smooth lacking transverse carinae (Fig. 7 b); carina bordering subpronope posteriorly, extending to anterior margin of pronotum (cf. Fig. 6 a); length of midtibial spur 0.5mm, length of midbasitarsomere 0.6mm; midtibia with 3–4 pegs at midlength, lacking pegs apically; hind tibia with 5–7 pegs apically; forewing length 6.1mm. Metasoma: median syntergite 2 + 3, length 1.3mm, width 1.2mm. Male: Unknown Etymology: In honor of Barb Sharanowski. Specimen Examined: HOLOTYPE. Ψ, COLOMBIA, Meta, PNN [Parque Nacional Natural] Macarena Borde Rio, quejar, 3 º 20 ’N 73 º 53 ’W, 460m, Malaise, 24–28.xii. 2001, D. Campos, leg. M. 2609, (head and antennae detached and placed on triangular point), (IAvH).Published as part of Lindsay, Cheryl & Sharkey, Michael, 2006, Revision of the genus Amputoearinus (Hymenoptera: Braconidae: Agathidinae) with fourteen new species, pp. 1-27 in Zootaxa 1329 on pages 21-22, DOI: 10.5281/zenodo.17417

    Pneumagathis Sharkey

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    Pneumagathis Sharkey NEW GENUS (Fig. 5) Type species. Pneumagathis spiracularis (Muesebeck) NEW COMBINATION. Original combination = Bassus spiracularis Muesebeck (1927). Diagnosis. Propodeal spiracle elongate (Fig. 5); first median tergite mostly smooth with two longitudinal carinae; frons not margined by carinae; tarsal claws simple with a large basal lobe. The only other agathidines with elongate propodeal spiracles belong to the Disophrini and these have cleft claws. Description. Head. Lateral carina on frons (as in members of Alabagrus) absent; interantennal space with a raised triangular prominence; gena not extended ventroposteriorly into sharp prominence; gena lacking sharp angle posterad eye; mandible dorsoventrally flattened; labial palpus with 4 segments, third segment reduced, less than ½ length of apical segment. Mesosoma. Propleuron weakly convex ventrally and lacking a sharp bump; notauli impressed, with or without pits or crenulae; posteroscutellar depression absent but sculpture usually present in this area; propodeum areolate rugose, usually with pair of longitudinal carinae and a transverse carina that define distinct areolae; propodeal spiracle large and elongate, more than twice as long as wide; sclerite between hind coxal cavities and metasomal foramen narrow but complete, its ventral margin situated ventral to dorsal margin of hind coxal cavities. Legs. Fore tibia lacking dull pegs; mid and hind tibia with blunt apical or preapical pegs; all tarsal claws with basal lobe. Wings. Fore wing RS+M vein mostly absent; second submarginal cell triangular and petiolate; fore wing 3 RSb straight and strong throughout; hind wing r and r-m cross veins absent; hind wing vein CUb tubular as a short stub, or lacking tubular stub, with a long pigmented non-tubular extension; wings strongly infuscated. Metasoma. First median tergite longer than wide apically, with or without weak rugose sculpture and always with pair of longitudinal carinae; second and third median tergites smooth; second tergite with a strong curved transverse depression; ovipositor about as long as metasoma, slightly thickened preapically. Diversity and Distribution. There are two described species, Pneumagathis spiracularis (Muesebeck) and P. brooksi (Sharkey ) new combination (original combination = Bassus brooksi Sharkey (in Janzen et al. 1998)) and perhaps a few more undescribed species. The genus is widespread and rather common over most of the contiguous states of the US, south through the dryer regions of Costa Rica. Biology. Parasitoids of skippers (Hesperiidae), P. spiracularis has 4 published hosts Cogia outis, Gesta gesta, Erynnis zarucco, Systasea pulverulenta (Marsh et al. 1979) and Pneumagathis brooksi new combination parasitizes 24 known species of pyrgine hesperids (Janzen et al. 1998). Janzen et al. (1998) discuss other aspects of the life-history of P. brooksi. Etymology. From the Greek agathis, meaning “ball of thread”, and the Greek pneuma, meaning “breath”. The latter word is a reference to the elongate propodeal spiracle (Fig. 5 D). The gender is feminine. Molecular data. H 1172 (COI: ATRMK 483 - 11); H 1173 (COI: ATRMK 488 - 11)Published as part of Sharkey, Michael J. & Chapman, Eric G., 2015, The Nearctic genera of Agathidinae (Hymenoptera: Braconidae) with a phylogenetic analysis, illustrated generic key, and the description of three new genera, pp. 49-72 in Zootaxa 4000 (1) on pages 62-63, DOI: 10.11646/zootaxa.4000.1.2, http://zenodo.org/record/28924
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