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    Entrevista a Francisco Serrano

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    Entrevista a Francisco Serrano, treballador d'ENASA (Pegaso) des del 1966 i afiliat a CCOO. Relata el moviment obrer dels anys 70 a Sagrera i a Zona Franca. Explica com va viure les reivindicacions polítiques i econòmiques dels treballadors i la seva participació en la lluita obrera

    A Tejon Serrano

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    Serrano man, head-and-shoulders portrait, facing front.Edward S. Curtis Collection (Library of Congress).Curtis no. 4152.Published in: The North American Indian / Edward S. Curtis. [Seattle, Wash.] : Edward S. Curtis, 1907-30, Suppl. v. 15, p. 513

    SERRANO, Juan

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    Telegram from Mr. Juan Serrano to Gen. Alvaro Obregón congratulating him on his return to Sonora. Thank-you reply. File S-010 / Telegrama del Sr. Juan Serrano al Gral. Alvaro Obregón, felicitándolo por su regreso a Sonora. Respuesta agradeciendo. Exp. S-01

    Geocharis antheroi Serrano & Aguiar, new species

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    Geocharis antheroi Serrano & Aguiar, new species (Figs 4 a– 4 h, 5 c– 5 d, 6 b) Type material: Holotype, 3, Portugal, Carção (Vimioso) (U.T.M. coordinate: 29 TQG0010), 22.IV. 2010, (A. Serrano leg.). Paratypes, 8 3 and 12 Ƥ (23, 1Ƥ gold coated), same locality of holotype, 22.IV. 2010, (A. Serrano leg.), 1 Ƥ, Santulhão (Vimioso) (U.T.M. coordinate: 29 TPG 9806), 22.IV. 2010, (A. Serrano leg.). Diagnosis. Anophtalmous; body slightly depressed, brown with integument microreticulated. Sparse pubescence mainly on pronotum and elytra. Elytron with vestigial striae, humeral region strongly punctured, disk with one anterior and one posterior seta. Male forelegs with the first tarsomere dilated. Males and females without a median tooth on the internal margin of the metafemora. Mesotibiae with dense pubescence in both margins. Hind tibiae more or less straight. Aedeagus as in Figs 5 c– 5 d. Description. Length of holotype: 1.9 mm. Length of paratypes: 1.6–1.9 mm (males and females). Head (Fig. 4 a) more or less as long as wide [length: 0.31–0.37 mm (males), 0.27–0.37 mm (females); width: 0.32–0.38 (males), 0.33–0.38 mm (females)], microsculpture distinct. Cephalic chaetotaxy (fixed setae of right side): L 3 +C 1 +F 1 +SA 1 +SP 1 + V 1 +O 1 +P0+G 2. Frontal and clypeal setae inserted in two large and two small sulci, respectively. Antennae light brown, antennomeres 1–2 longer than the others, the latter subpyriform, the third and the fourth are the shortest ones and subpyriforms, antennomeres 5–10 gradually longer and oval-shaped, the last one acuminate. Mouth-parts (Fig. 4 b) show the general pattern of the genus. Pronotum cordiform (Fig. 4 c), transverse, about 1.2–1.3 times wider than long [length: 0.35–0.43 mm (males), 0.34–0.41 mm (females); width: 0.43–0.51 mm (males), 0.42–0.50 mm (females)]; anterior angles not produced, widely rounded off, lateral channel not recurved inside of anterior angles; disk slightly convex, depressed between the two basal pits, with a superficial central sulcus which does not reach the anterior margin; anterior margin slightly straight and posterior margin slightly arcuate outwards; lateral margins with two or three slight denticles just before the posterior angles, which are right and dentate. Vestiture (pubescence): surface covered with scattered erect pubescence; one seta on the lateral margin in the broadest part of the pronotum, another one near the posterior angle; two additional setae inserted near the anterior angles. Elytra (Figs. 4 d– 4 e) 1.7–1.8 times longer than wide [length: 0.96–1.12 mm (males), 0.94–1.08 mm (females), width: 0.54–0.63 mm (males and females)], slightly convex, parallel and oval posteriorly, with vestigial traces of striae. Tegument microsculptured, disk more punctured in the shoulders, punctures sparser and disappearing to apex (Figs. 4 d– 4 f); lateral margin narrow, finely serrate from the humeral angles, which are rounded, to the seventh seta of the umbilicate series. Vestiture: part of the pubescence of the disk is arranged in six irregular lines, these setae are erect and slightly directed anteriad (Fig. 4 e); umbilicate series follows the pattern of the genus. The longest setae of this series are the 2 nd, the 6 th and the 9 th with the 3 rd, 5 th, 7 th and 8 th, more slightly inserted within the elytral margin; besides these setae there are one parascutellar basad, two discal (one anterior and one posterior) and one apical seta (Figs 4 d– 4 f). Last abdominal ventrite with one pair of medium sub-marginal setae in males, two pairs of medium sub-marginal setae in females (Figs 4 g– 4 h). Male legs with the protarsomere 1 dilated; tarsomere 1 in all legs more pigmented (light brown) than the others; mesotibiae with a strong pubescence on both margins; hind femora (males and females) without any median tooth on the internal margin (Figs. 4 g– 4 h). Male genitalia (Figs 5 c– 5 d) with median lobe arcuate (lateral view), apex strongly sharp and bent down (lateral view), arcuate inwards in the right side and broadly rounded in the left side (dorsal view); basal lobe with apophysis prominent, basal edge arcuate; internal sac as in figures 5 c– 5 d; left and right parameres with 2 apical setae, left paramere with dorso-basal edge expanded (Fig. 5 c). The female genitalia follows the general pattern described for the other species of the genus (e.g., Zaballos & Jeanne 1987, Zaballos 1998, Zaballos, 2005). Female genitalia (Fig. 6 b) shows the gonocoxite IX sickle-shaped, and has a long ensiform seta in the middle region of the external margin, one ensiform seta in the middle dorsal region and a double nematiform seta in the internal margin near the beginning of the apical third. Gonosubcoxite IX without special features; laterotergite IX with a variable number of setae (8–10). Internal genitalia (not shown) with spermathecal duct long, parallel and enlarged very slightly near the bursa copulatrix (length: 0.16 mm), a spherical spermatheca (width: 0.019 mm), duct of spermathecal gland short and thin, gland fusiform (lenght: 0.096 mm), middle region membranous, apical portion sclerotized. Etymology. This new species is dedicated to the Portuguese naturalist Anthero Frederico Ferreira de Seabra, who greatly contributed to the knowledge of the Coleoptera of Portugal in the first half of the XX century. Morphological affinities. The two new species share with most species of Geocharis two setae on the elytral disk, one anterior and one posterior (Table 1). In Table 1 some other morphological characters like the presence or absence of elytral striae, a tooth on the internal margin of the hind femora and the number of setae of the left paramere are compared in all species of Geocharis. Taking into account the presence on the elytral disk of two setae, the absence of a tooth on the internal margin of the hind femora of males and females, as well as the fact that the adults present traces of elytral striae, the new species are close to G. b i v a r i Serrano & Aguiar, G. julianae Zaballos, G. k o r b i (Ganglbauer), G. massinissa (Dieck), G. montecristoi Zaballos and G. testatretafoveata Zaballos (all the adults of these species exhibit the above characters). Nevertheless, in G. b i v a r i and G. julianae the elytral striae are stronger than in the new species (see Zaballos 1989, Serrano & Aguiar 2004). Geocharis bivari, G. julianae and the remaining other four species are also easily segregated from the new ones by the shape of the median lobe of aedeagus plus the pattern of internal sac sclerites (Zaballos 1989, 2005, Serrano & Aguiar 2004 a). The new species are easily separated from one another by the aedeagus conformation (cf. Figs 5 a– 5 b and 5 c– 5 d) though they much resemble each other on external morphology. The growing knowledge of the genus Geocharis in terms of number of species and morphological peculiarities, corroborates former conclusions (e.g. Zaballos 2005) about the difficulty of identifying its species based only in the external morphology, with the exception of some taxa (see Table 1). Characters that are not mentioned in Table 1 like ovulate elytra and the presence of a vestigial sutural stria (G. coiffaiti and G. femoralis), left paramere features, the pronotum shape or even hind tibiae shape are useful for identifying particular Geocharis species. This happens with adults of G. leoni Zaballos which present the left paramere with lamellar and membranous scales in the apex instead of setae, the adults of G. portalegrensis which exhibit a pronotal disk strongly flattened, the adults of G. iborensis Zaballos which present the hind tibiae arcuate inwards instead of the general straight pattern, and the adults of G. julianae which present a fold in the internal margin of this structure (Zaballos 1989, 1990, 1998, Serrano & Aguiar 2000). Indeed, the best characters to identify and segregate the Geocharis species, besides the ones mentioned above, are found in the aedeagus, as are the general conformation of median lobe (lateral view), the apex of the same structure (dorsal view) and the armature pattern of the internal sac. Faunistic data on other Geocharis taxa and Hypotyphlus lusitanicus. Geocharis olisipensis Schatzmayr, 1937. Material examined: Bucelas (U.T.M. coordinates: 29 SMD 9305), 3.III.2011, 53, 4 Ƥ, 22.III.2011, 13, 2 Ƥ, 30.III.2011, 23, 1 Ƥ. Within the genus Geocharis this species was the first one described for Portugal based on two specimens collected near Lisbon (Schatzmayr 1937). Serrano & Aguiar (2004 a) after some efforts to locate this species found it in the outskirts of Lisbon (Valejas and Fanhões) and more recently in Serra de Montejunto (almost 50 km north of Lisbon) (Serrano & Aguiar 2008). The new locality near Bucelas is close to the former ones (Fig. 7). Geocharis quartaui Serrano & Aguiar, 2004. Material examined: Serra de Sicó (Pombal) (U.T.M. coordinates: 29 SNE 3319), 31.III.2008, 1Ƥ, 15.IV.2008, 13, 3 Ƥ; Serra do Sicó (Pombal) (U.T.M. coordinates: 29 SNE 3419), 15.IV.2008, 23 3, 10 Ƥ. This species was described on the basis of several specimens collected near Alcobaça (Carvalhal), a locality close to Serra de Aire e Candeeiros and recorded later in Serra de Montejunto (Serrano & Aguiar 2008). This new localization in Serra do Sicó (almost 40 km north of Serra de Montejunto), indicates for this species a wider distribution than the previously thought (Fig. 7). Interesting also is the fact that the Serra de Sicó male specimens exhibit a stronger median tooth on the internal margin of the hind femora than that observed in specimens of the other localities. Hypotyphlus lusitanicus Serrano & Aguiar, 2004. Material examined: Aldeia do Catarredor (Serra da Lousã) (U.T.M. coordinates: 29 TNE 6636), 2.IV.2008, 2 3; Fundeira (Pampilhosa da Serra) (U.T.M. coordinates: 29 TNE 8731), 3.IV.2008, 4 3, 3 Ƥ; Silvares (Fundão) (U.T.M. coordinates: 29 TPE 1141), 4.IV.2008, 1 Ƥ; Folgosa (Peso da Régua) (U.T.M. coordinates: 29 TPF 1357), 29.IV.2009, 1 Ƥ. One male and 1 female from Fundeira are deposited in the collection of Vicente Ortuño; all the remaining specimens are deposited in the collection of the first author, Departamento de Biologia Animal (Faculdade de Ciências da Universidade de Lisboa). This remarkable species was found in the centre of Portugal (Aldeia do Mato near Tomar) (Serrano & Aguiar, 2004 b). Lately it was found in a new locality (Barragem do Cabril, Pedrógão Grande) almost 60 km farther north (Serrano & Aguiar 2008). The new localities notably increase east- and northwards the distribution of this species (Fig. 7). Ecological and geographical considerations. The two new species of Geocharis inhabit the endogean environment in the same way than other congeneric taxa. This means that they live in the soil, usually at different depths of the B-horizon. They are found under sunken stones laying at different depths, from superficial (epigean) to well-buried (edaphic or endogean) environments. Individuals of endogean carabids are more easily found close to the superficial horizon layers after heavy rains because then the soil reaches higher percentage of humidity (saturation or close to saturation), pushing the beetles upwards. References to this behaviour are not abundant (e.g. Zaballos 2005; Zaballos & Pérez-González 2011), though we sampled several specimens of G. sacarraoi Serrano & Aguiar and Typhlocharis passosi Serrano & Aguiar, 2005 after rain in these conditions. However, the soil horizons do not always present the same depth and many times the bed rock is closer to the epigean environment. In this case endogean individuals may occur for instance between plates of sunken schistose rocks (e.g. Fig. 2). Though only one species of Geocharis occurs at a given locality, there are some recorded exceptions (e.g. Zaballos 2005; Serrano & Aguiar 2001, 2006, 2008). Sometimes Geocharis species can also be syntopic with species of the genus Typhlocharis (e.g. Dieck 1869, Serrano & Aguiar 2008). To date, all the Geocharis species located north of Tejo River in Portugal were recorded from localities ranging between the Lisbon region and Serra de Aire e Candeeiros. Interestingly, despite several field surveys to the north of Tejo River, only those conducted in the western region north of Lisbon led to the discovery of Geocharis species until now. Geocharis quartaui (also collected in this study) was found in Serra de Sicó, a mountain also located in the western region. The finding of the two new species here reported from surveys near the Douro River and Vimioso has increased the distribution of this genus 240 km farther north of Serra de Sicó. Yet, more remarkable was the discovery of Hypotyphlus lusitanicus in Folgosa in syntopy with G. barcorabelo n.sp. The presence of the former species north and south of Serra da Estrela (Fig. 7) corroborates the hypothesis that H. lusitanicus is a zoogeographical relict of a lineage that once must have extended westward from the Lionigurian massif to the Lusitanian massif (Serrano & Aguiar 2004 b). Finally we would like point out that G. barcorabelo n. sp. and G. antheroi n.sp. were not found within the limits of any Natural Park or protected area and as such are not currently under any conservation protection. From the 20 Geocharis species known in Portugal (including the new ones) (Table 1), only 7 (G. b i v a r i, G. boieiroi, G. monfortensis, G. m o s c a t e l u s, G. portalegrensis, G. quartaui and G. rodriguesi) were recorded inside the geographical limits of a protected area.Published as part of Serrano, Artur R. M. & Aguiar, Carlos A. S., 2011, Two new species of the genus Geocharis Ehlers, 1883 and new data on Anillina species from Portugal (Coleoptera: Carabidae), pp. 33-46 in Zootaxa 3116 on pages 38-45, DOI: 10.5281/zenodo.20716

    A Serrano woman of Tejon

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    Serrano woman, head-and-shoulders portrait, facing front.Edward S. Curtis Collection (Library of Congress)Curtis no. 4141.Published in: The North American Indian / Edward S. Curtis. [Seattle, Wash.] : Edward S. Curtis, 1907-30, Suppl. v. 15, pl. 512

    SERRANO, Francisco Roque (Gral.)

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    Correspondence of Gen. Francisco Roque Serrano, Undersecretary of War and Navy, Mr. Fernando Torreblanca and Gen. Alvaro Obregón, in which Gen. Serrano informs Gen. Obregón of Gen. Francisco Villa’s surrender. Gen. Serrano informs Gen. Obregón that he has made the arrangements to hook up a car to a train he would like to board for his exclusive use. Gen. Obregón informs Gen. Serrano of the terrible travel conditions that some Yaqui families have to endure under the responsibility of Gen. Fructuoso Méndez. Reply stating that he has already taken care of that issue. Gen. Serrano informs Gen. Obregón of the circumstances under which Gen. Francisco Villa surrendered. Gen. Serrano informs Gen. Obregón that Mr. Juan Platt is heading to Los Angeles, CA. Gen. Serrano informs Gen. Obregón that Gen. Benjamín Hill concurs with his view of the Gen. Villa’s matter. Gen. Obregón requests Gen. Serrano to inform Gen. Pedro Zamora of the date he will meet him in his office. Gen. Serrano requests Mr. Fernando Torreblanca to clarify the amount of money spent on the wedding present that Gen. Obregón will gift Gen. Jesús M. Garza. Gen. Obregón requests Gen. Serrano to provide Mr. George M. Bizler with the necessary tools to fix the Packard car. Gen. Serrano informs Gen. Obregón of the arrival of the “Melchor Ocampo” ship at the Mexico Port. Matter concerning Mr. Luis Hostetter’s returning travel to Mexico. Matter concerning the hire of another ship, since there is a case of yellow fever in the “Melchor Ocampo” ship. Gen. Obregón requests Gen. Serrano to see Gen. Juan C. Zertuche, who is bringing a message to him. Gen. Serrano transcribed for Gen. Obregón a message from Gen. Pablo González, who is based in New York. Gen. Serrano informs Gen. Obregón that the bill against Garza proposed by the Chamber of Congressmen was vetoed. Files S-23 and S-09. / Correspondencia entre el Gral. Francisco Roque Serrano, Subsecretario de Guerra y Marina, el Sr. Fernando Torreblanca y el Gral. Alvaro Obregón, en la que el primero comenta al Gral. Obregón sobre la rendición del Gral. Francisco Villa. El Gral. Serrano comunica al Gral. Obregón que ha ordenado se anexe un carro para él en el tren que indique. El Gral. Obregón informa al Gral. Serrano sobre las pésimas condiciones en que viajan familias yaquis al cuidado del Gral. Fructuoso Méndez. Respuesta indicando ya tomó medidas para resolver esta situación. El Gral. Serrano informa al Gral. Obregón de las condiciones en que se rindió el Gral. Francisco Villa. El Gral. Serrano comunica al Gral. Obregón que el Sr. Juan Platt va rumbo a Los Angeles, Cal. El Gral. Serrano comunica al Gral. Obregón que el Gral. Benjamín Hill está de acuerdo con él en el asunto del Gral. Villa. El Gral. Obregón solicita al Gral. Serrano le informe al Gral. Pedro Zamora la fecha en que lo recibirá en su oficina. El Gral. Serrano solicita al Sr. Fernando Torreblanca aclare a cuánto asciende el monto del regalo de bodas que le hará el Gral. Obregón al Gral. Jesús M. Garza. El Gral. Obregón solicita al Gral. Serrano le proporcione al Sr. George M. Bizler los accesorios para que repare el coche Packard. El Gral. Serrano informa al Gral. Obregón de la llegada a Puerto México del buque "Melchor Ocampo". Asunto relativo al viaje de regreso a México del Sr. Luis Hostetter. Asunto relativo a la contratación de otro barco porque en el "Melchor Ocampo" hay a bordo un caso de fiebre amarilla. El Gral. Obregón solicita al Gral. Serrano que reciba al Gral. Juan C. Zertuche, quien le lleva un mensaje. El Gral. Serrano transcribe al Gral. Obregón un mensaje del Gral. Pablo González quien se encuentra en Nueva York. El Gral. Serrano comunica al Gral. Obregón que fue rechazado el dictamen de la Comisión de la Cámara de Diputados en contra de Garza. Exps. S-23 y S-0

    "Andrea´s love": love as a necessity

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    La fórmula de Martín Cuenca nos recuerda a la filosofía de André Bazin, a la nostalgia de la nouvelle vague y al hiperrealismo de la posguerra: actores desconocidos, actores- persona para potenciar los elementos de identificación, historias que encontramos en los barrios marginales fotografiados con su propia poética, personajes desheredados nacidos del claroscuro, de la exaltación de los sentimientos, o de los bloqueos emocionales, un drama social que trasciende… Elementos de un cine clásico que ya existió, que ha existido siempre, que en "El amor de Andrea" tienen una expresión peculiar, en una película muy personal de Manuel Martín Cuenca.Depto. de Ciencias de la Comunicación AplicadaFac. de Ciencias de la InformaciónFALSEpu

    SERRANO, Eugenio P.

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    Letter from Mr. Eugenio P. Serrano to Gen. Alvaro Obregón congratulating him on his presidential nomination. File S-22. / Carta del Sr. Eugenio P. Serrano al Gral. Alvaro Obregón felicitándolo por su candidatura a la Presidencia. Exp. S-2

    Typhlocharis mendesi Artur R. M. Serrano & Carlos A. S. Aguiar 2017, sp. n.

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    Typhlocharis mendesi sp. n. Figs 1-17 Holotype: cAS; 1 ♂; Portugal, Bucelas (Estremadura Prov.) (UTM: 29SMD9305), 3.III.2011, Serrano & Aguiar leg. Paratypes: cAS; 3 ♂♂ and 2 ♀♀; same locality and date as holotype. – MHNG; 2 ♂♂ and 2 ♀♀; same locality and date as holotype. – cAS; 4 ♂♂ and 2 ♀♀; Bucelas (Estremadura Prov.) (UTM: 29SMD9305), 22.III.2011. – cAS; 3 ♂♂ and 4 ♀♀; Arruda dos Vinhos (Estremadura Prov.) (UTM: 29SMD9012), 3.III.2011. – cAS; 1 ♂; A. do Mourão (Estremadura Prov.) (UTM: 29SMD9209), 3.III.2011. – cAS; 1 ♀; Bucelas (Estremadura Prov.) (UTM: 29SMD9205), 23.X.2012. – cAS; 5 ♂♂ and 5 ♀♀ (2 ♂♂ and 2 ♀♀ gold coated); Bucelas (Estremadura Prov.) (UTM: 29SMD9305), 7.XII.2012. – MHNG; 1 ♂ and 1 ♀; same locality and date as previous lot. – cAS; 7 ♂♂; 7 ♀♀; Bucelas (Estremadura Prov.) (UTM: 29SMD9305. 1.XI.2013. All Serrano & Aguiar leg. Diagnosis: Anophthalmous. Body parallel, depressed, brown or brownish-yellow with integument microreticulate and scattered pubescence. Antennae moniliforme. Semilunar notch present. Vertex with pars stridens. Pronotum rectangular (1.2 times longer than wide). Apical edge of elytra slightly sinuate, without teeth; discal setae arranged in 4 or 5 rows slightly bent anteriad; umbilicate series with six (4+2) marginal setae; abdominal female sternum II and III with lateral fovea; anterior region of the last abdominal sternum in both sexes showing a smooth or scaly and serrate microsculpture (‘belt’) (Figs 13, 14). Hind trochanters inerm in both sexes (Figs 7-8). Aedeagus (Figs 15, 16) sickle-shaped, basal lamina markedly arcuate, the apex slightly eagle’s beak shaped (lateral view); parameres bisetulose. Female genitalia (Fig. 12) with long tubular gonocoxites, each one with a double apical seta and without a lateral seta. Etymology: This new species is dedicated to our colleague Luís Mendes, eminent Portuguese entomologist, who has been greatly contributing to the taxonomic and faunistic knowledge of the world Zygentoma, Microcoryphia, and Afrotropical Lepidoptera. Description: Length of holotype: 1.14 mm. Length of paratypes: 1.12-1.23 mm (males) and 1.15- 1.27 mm (females). Head (Figs 1, 2, 9, 10): Almost as long as wide [length: 0.21-0.24 mm (males) and 0.22-0.26 mm (females), width: 0.24-0.26 mm (males) and 0.24-0.27 mm females)] with hexagonal microsculpture; vertex with transverse microsculpture which, in the middle area bellow the anterior margin of the pronotum, is arranged in parallel ridges forming a stridulatory organ (pars stridens) (Fig. 9); posterolateral region of the head with a semilunar notch (Fig. 1); labrum sub-rectangular, anterior margin almost straight; anterior margin of clypeus almost straight; moniliform antennae with 11 rounded antennomeres progressively more quadrate (morph 1, subquadrate, as defined in Pérez-González & Zaballos, 2013a), except last one, pyriform (Fig. 10). The last antennomere has a pattern of three anterodorsal and one posterodorsal sensilla coeloconica (type A). Ventral sensilla coeloconica in antennomeres 5° and 6°. Stem of third antennomere short (average proportion of stem length/antennomere body length=0.38). Labium with epilobes and middle tooth slightly sharp. Ligula with convex middle lobe and paraglossae not long. Gula wide with sutures visible through light microscopy. Cephalic chaetotaxy: two pairs of submarginal labral setae (l-s/s-l) and two pairs of marginal labral setae (m-m/m-m), two pairs of clypeal setae (l-s/s-l), one pair close to frontal sulcus; two pairs of supraocular setae (anterior and posterior) and 1-2 pairs of setae on the posterior region between vertex and lateral carinae. Pronotum (Fig. 3): Rectangular, slightly longer than wide [length: 0.3-0.33 mm (males) and 0.31-0.34 mm (females); width: 0.26-0.28 mm (males) and 0.27- 0.29 mm (females)], with hexagonal microsculpture, lateral margins slightly arcuate, slightly narrowed posteriorly, with 3-4 small crenulations before the posterior angles, which are marked with one small tooth; disk flattened, anterior margin almost straight and posterior margin slightly arcuate outwards in median region, anterior margin with medial hiatus. Pronotal chaetotaxy: four longitudinal series of erect setae between midline and lateral margins, directed inwards and slightly anteriad; lateral groove of each side with one long seta in the anterior quarter and one long seta on hind angle; a row of five pairs of setae (l-l-l-l-l/l-l-l-l-l) parallel to the anterior margin, two pairs of setae near the posterior margin (l-l/l-l); a row of regularly placed setae in anterior, posterior and lateral margins, more spaced in the later. Elytra (Figs 4, 5, 6): 2.1 times longer than wide [length: 0.58-0.65 mm (males) and 0.59-0.66 mm (females); width: 0.22-0.30 mm (males) and 0.29-0.30 mm (females)], parallel and oval posteriorly, dorso-ventrally flattened on the disk; transverse scutelar organ present; disk with hexagonal and pentagonal micro-sculpture, almost scaly in the lateral regions; scutelar region not punctured; humeral angles rounded; lateral margins serrate, the teeth decreasing in size posteriorly; apical margin without teeth. Elytral chaetotaxy: Part of the pubescence of the disk is arranged since the sutural region in four or five rows of short setae slightly directed anteriad; a very irregular series of minute setae between the outer row and the lateral margins directed outward; umbilicated series aggregated with four seta in the anterior group and two in the posterior group (4+2) (Figs 4, 5). Legs: Femora robust, inner margin of pro- and mesofemora inerm, slightly dentate in metafemora; metatrochanters similar in both sexes and without any special features (Figs 7, 8); meso- and metatibiae without apical teeth (Figs 7, 8); apex of fore tibia with a comb of setae (cleaning organ) similar in both sexes (Fig. 11); male protarsus without dilated segments. Abdomen (Figs 7, 8): Sterna II and III with lateral fovea, deeper in females than in males; last abdominal sternite (both sexes) presents in its anterior region a more or less invaginated area of smooth and scaly zone of microtrichia (abdominal ‘belt’) (Figs 13, 14); last sternite chaetotaxy with sexual dimorphism: male (s-l-s-s-l-(s)s/s(s)-l-s-sl-s), female (s-l-s-s-l-(m)m/m-l-s-s-l-s) (Figs 13, 14). Male genitalia (Figs 15, 16): in lateral aspect with median lobe sickle-shaped, basal lamina markedly arcuate, the apex slightly shaped like an eagle’s beak; in dorsal aspect with apex broadly rounded, slightly bent to left; internal sac with several tangled membranes, in central area with one twisted sclerite in continuous by one fusiform membrane (Fig. 16); parameres bisetulose apically; external tegument of median lobe with several presumably chemo- or chemo-mechano sensilla sunken in pits (Fig. 15). Ring sclerite (IX abdominal sternum) subtriangular-arcuate, the apical margin is slightly projected in a triangular blunt extension, gently tilted. Female genitalia (Fig. 12): Tubular ovipositor gonocoxites weakly sclerotized, each one in ventral aspect without a lateral seta and with a double apical one (Fig. 12); internal genital tract with spermathecal duct short, spermatheca spherical (Fig. 17) and short (length: 0.022 mm); conical spermathecal gland (length: 0.047 mm) sclerotized in distal region (Fig. 17). Variability: There is a certain degree of intraspecific variation affecting some characters, such as the chaetotaxy of the labrum, the number and marking of posterolateral denticles in the pronotum (from faintly marked to almost absent), or the chaetotaxy of the last ventrite, showing some variability of the number and length of marginal seta. It is also common that some individuals show short and long setae irregularly mixed in the anterior row of setae in pronotum. DISCUSSION The following traits and others more have been used to discuss affinities among Typhlocharis species or to justify their close geographical proximity or even syntopy (Ortuño & Gilgado, 2011; see also Andújar et al., 2010): conformation of clypeus, presence or absence of pars stridens, type of gula, discal setae on the elytron, elytral pores, tarsal formula, hind trochanter shape, sexual dimorphism [e.g. presence or not of: 1) a median tubercle or carina in the sternum II (males), 2) a more or less deep foveae in the sterna II, III or even IV (predominantly in females) and, 3) thorn-shaped hind trochanter (males); different chaetotaxy of the last abdominal segment]. Being useful for practical purposes, the morphological species groups proposed by Zaballos & Ruíz-Tapiador (1997), Zaballos & Wrase (1998), and more recently by Pérez-González & Zaballos (2013b), are not always coherent, sometimes even for the geographic distribution of the species within each group (e.g. Zaballos & Pérez- González, 2011). This is evident, for instance, for the diecki- and outereloi- groups, whose diagnoses are based almost exclusively on the presence of a maximum number of two teeth in the apical margin of each elytron and on the pattern of the umbilicate series (4+3 – diecki -group, 4+2 or 4+1 – outereloi -group) (Zaballos & Wrase, 1998; Serrano & Aguiar, 2008). The latter character presents a degree of variability (4+2, 4+3 or 4+1, 4+2, 4+3) within several other Typhlocharis groups also (e.g. Zaballos & Pérez-González, 2011; Pérez-González et al., 2013; see also Pérez-González & Zaballos, 2012: table 1). At a first sight, the absence of a median tooth in clypeus and teeth in the apical margin of elytra allows the inclusion of T. mendesi sp. nov. in the silvanoides- or diecki -group. The former group includes six species (see Serrano & Aguiar, 2014: table 1) and is known for the stability of the 4+4 umbilicate series pattern and the absence of teeth in the apical margin of elytra (see Pérez- González & Zaballos, 2012: table 1). Thus, T. mendesi sp. nov. shares with all species of the silvanoides -group the absence of teeth in the apical margin of the elytra, but it is well differentiated by the 4+2 umbilicate series pattern. The diecki -group includes eleven species (see Serrano & Aguiar, 2014: table 1), all with 4+3 umbilicate series pattern and of which only T. armata Coiffait, 1969 and T. deferreri Zaballos & Pérez-González, 2011 present an apical margin of elytra without teeth also (see Pérez- Gonzalez & Zaballos, 2012: table 1). So, for both the silvanoides -and diecki -group the 4+2 umbilicate series pattern of the new species is apparently incompatible. This could be solved redefining both groups by encompassing species with 4+2 umbilicate series pattern, which would allow the inclusion of T. mendesi sp. nov. in one of them. Taking into account only this pattern of umbilicate series, the new species could be included in some of the remaining groups too. Thus, apart the difficulty into which species group the new species should be included, it is easily differentiated from all species of the silvanoides- or diecki -group by the 4+2 umbilicate series pattern and from part of the species of these two groups by the presence of abdominal ‘belt’ in males and females also (see Serrano & Aguiar, 2014: table 1). On the other hand, it is easily differentiated from all species of the monastica -group by the female genitalia model (tubular gonocoxite vs. unguiform gonocoxite shapes), from the quadridentata - group [all species except T. quadridentata (Coiffait, 1969)] by the tarsal formula (5+5+5 vs. 4+4+4) and the absence of elytral teeth in the apex and from the gomezi-, carpetana-, outereloi- and baetica- group ‒ among other traits ‒ by the absence of elytral teeth in the apex (see Pérez-González & Zaballos, 2012: table 1; 2013b: table 1). However, by the female tubular genitalia model (submodel with long tubular gonocoxites without lateral setae and two apical nematiform setae) (Pérez-González & Zaballos, 2012), the tarsal formula (5+5+5), the absence of a median clypeal tooth and of meso- and metatibial apical teeth, it seems closer to the outereloigroup. This group is characterized by the presence of at least one sutural tooth or yet additional one at the end of the 7th elytral striae. Taking into account the above set of characters, we propose that Typhlocharis mendesi sp. nov. must be incorporated in outereloi -group. This implies also that the characterization of the group needs to be redefined by the inclusion of ‘absence of teeth in the apical margin of elytra’. This redefinition avoids the creation of a new species group, which would greatly complicate the artificial systematics of the genus. The outereloi -group contains a variety of morphologically diverse species (e.g. Serrano & Aguiar, 2008: table 2; Pérez-González & Zaballos 2012: table 1). Within the group, the new species differs from T. elenae Serrano & Aguiar, 2002 and T. gomesalvesi Serrano & Aguiar, 2002 by the umbilicate series pattern (4+2 vs. 4+1), but shares with 5 species (T. outereloi Novoa, 1979, T. bazi Ortuño, 2000, T. laurentii Magrini, 2000, T. singularis Serrano & Aguiar, 2000 and T. gomesalvesi) the presence of a pars stridens as well as with 3 species (T. laurentii, T. singularis and T. gomesalvesi) the presence of a medial hiatus in the pronotum. Another feature shared by the new species and both sexes of 8 species of the group (T. outereloi, T. bazi, T. belenae Zaballos, 1983, T. intermedia Zaballos, 1986, T. navarica Zaballos & Wrase, 1998, T. laurentii, T. singularis and T. gomesalvesi) are the ovoid-elongate hind trochanters versus the thornshaped (at least in males) in the remaining species. Additionally, the presence of a fovea in the female abdominal sternum II or sterna II and III is shared by 6 species (T. bazi, T. navarica, T. atienzai Zaballos & Ruiz- Tapiador, 1997, T. estrellae Zaballos & Ruiz-Tapiador, 1997, T. bullaquensis Zaballos & Ruiz-Tapiador, 1997 and T. elenae). It seems that morphologically the new species is more akin to T. singularis, T. gomesalvesi, and T. laurentii. Interestingly, the two former species occur geographically closer to Portugal (Serrano & Aguiar, 2000, 2002), and the latter farther away (Almeria, Spain) (Magrini, 2000). Morever, T. mendesi sp. nov. is also similar to T. laurentii concerning the number of setae of the female gonocoxite (apical and lateral: 2, 0). However, the new species is easily separated from all species of the outereloi -group, including the closely resembling ones, by the absence of teeth in apical margin of elytra and the presence of a slight fovea in the male abdominal sternum II (Figs 6, 7). There are three Typhlocharis species belonging to other species groups with a known geographical distribution in Portugal which is closer to the new species than the other two mentioned above (T. singularis and T. gomesalvesi). They are Typhlocharis rochapitei Serrano & Aguiar, 2008, T. passosi Serrano & Aguiar, 2005, and T. bivari Serrano & Aguiar, 2006, belonging to the diecki and gomezi species groups, respectively. Typhlocharis rochapitei was found in the same region of the new species (see also the ‘Ecological remarks’) and T. passosi and T. bivari were located more northerly and far away (60 km) in the region of Serra d’Aire e Candeeiros (Serrano et al., 2005; Serrano & Aguiar, 2006). These observations reinforce the idea that the existing species groups, eventually with the exception of the baetica -group (Pérez-González et al., 2013; Pérez- González & Zaballos, 2013c), have no biogeographical or phylogenetic meaning and that obviously many traits that define them are probably convergences (e.g. see Ortuño & Gilgado, 2011). Despite the above comments, the inclusion of T. mendesi sp. nov. based on external morphologic characters in one of the already established species groups nevertheless challenges the relationships of the species so far included in each group. ECOLOGICAL REMARKS The new species of Typhlocharis inhabit the endogean environment in the same way as other congeneric taxa. They live in the soil, usually at different depths of the B-horizon, but are found also under sunken stones lying at different depths, even from superficial (epigean) to wellburied (edaphic or endogean) environments. Individuals of endogean carabids are more easily found close to the superficial horizon layers after heavy rains because then the soil reaches higher percentage of humidity (saturation or close to saturation), pushing the beetles upwards (e.g. Serrano & Aguiar, 2011; Zaballos & Pérez-González, 2011). Typhlocharis mendesi sp. nov. was found in three geographically close localities (see description) with calcareous rocky substrate and clayey brown- or brown-reddish colour soils. The habitats in the Bucelas and Arruda dos Vinhos region are dominated by dense herbaceous vegetation, typical of grassland (e.g. gramineous, liliaceous, leguminous, orchidaceous, asteraceous plants) with holm-oaks (Quercus ilex Linnaeus), pine trees (Pinus pinaster Aiton), daphnes (Daphne gnidium Linnaeus), hawthorns (Crataegus monogyna Jacquin), and other woody plants, while in A. do Mourão the vegetation is only grassland. The new species is syntopic with other endogean carabid species in the localities of Bucelas and Arruda dos Vinhos such as Typhlocharis rochapitei, Geocharis olisipensis (Schatzmayr, 1937), and Geocharis capelai Serrano & Aguiar, 2012 and also with other soil endogean arthropods like pseudoscorpionids, zopherid beetles (Doderonymus lusitanicus Binaghi, 1937), curculionid beetles, and hymenopteran formicids [Ponera coarctata (Latreille, 1802) and Solenopsis sp.]. The great concentration of endogean carabid beetles (at least four species) in Bucelas, among other endogean species, confers to this region a statute of a biodiversity ‘hot-spot’. Unfortunately, the region suffers from strong human influence and threats, like quarry industries, offroad racings, and other activities.Published as part of Artur R. M. Serrano & Carlos A. S. Aguiar, 2017, A new species of the genus Typhlocharis Dieck, 1869 (Coleoptera, Carabidae) from Portugal, pp. 39-46 in Revue suisse de Zoologie 124 (1) on pages 40-45, DOI: 10.5281/zenodo.32266
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