197,185 research outputs found

    Campodea (Campodea) portacoeliensis Sendra & Jimenez 1986

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    Campodea (Campodea) portacoeliensis Sendra & Jiménez, 1986 Material studied. Central System Mountains, Miraflores de la Sierra, near to Morcuera Mountain Pass; 1 ♂, scree slope MSS Reajo Cazado Stream, 13 VI 2016. GIBSES leg. Habitat and distribution. This epiedaphic species founds in the uppermost soil layer between the leaves. It is distributed in humid spots in the Iberic Mountains, Eastern of the Iberian Peninsula (Sendra & Jiménez 1986; Sendra 1989b; Sendra & Moreno 2004).Published as part of Sendra, Alberto, Jiménez-Valverde, Alberto, Gilgado, José D., Ledesma, Enrique, Baquero, Enrique, Pérez-Suárez, Gonzalo, Cuesta, Eva, Herrero-Borgoñón, Juan J., Jordana, Rafael, Tinaut, Alberto, Barranco, Pablo & Ortuño, Vicente M., 2017, Diplurans of subsurface terrestrial habitats in the Iberian Peninsula, with a new species description (Diplura: Campodeidae), pp. 61-80 in Zootaxa 4291 (1) on page 70, DOI: 10.11646/zootaxa.4291.1.4, http://zenodo.org/record/82936

    Mueggejapyx Sendra & Komericki 2021, gen. nov.

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    Mueggejapyx Sendra & Komerički gen. nov. urn:lsid:zoobank.org:act: 924B9ABA-1630-4832-86EE-19CE73C3E7CB Type species Mueggejapyx brehieri Sendra & Komerički gen. et sp. nov. Diagnosis Large and elongated body; antennae with 36 antennomeres; trichobothria on antennomeres 4–6 in a 3/5/5 pattern; a trichobothria in distal position; 1–2 placoid sensilla on medial and distal antennomere; apical antennomere with 16–18 placoid sensilla; dorsal and ventral side of head with numerous uniformly distributed sM and sm, and several M; five laminae pectinate on maxilla; elongated thoracic segments; and cuticle with tiny micro-holes, pronotum and mesonotum each with 5+5 M, metanotum with 2+2 M; strong internal Y cuticular furcisternite structures in pro-presternites and pro-, meso- and metasternites; urotergites with at most 1+1 M (ma), 1+1 M 1, 1+1 M 2–5; urite X longer than wide, with marked carinae with subparallel margins, with 2+2 intracarinal macrosetae; acropygium rounded; lateral angles of urotergites VI and VII with weakly to moderate lobiform projection; first urosternite with multiperforated surface and rounded protrusions, with 3+3 M on prescutum and 11+11 M on scutum; median glandular organ with seta-shaped sensilla; lateral subcoxal organs with one to three rows of short glandular setae (GS), one row of sensory setae (SS), and a row of setae with large socket; sternites with abundant sm and strong M; cerci strong, elongated, rectilinear and becoming curved subapically, heavily sclerotized with external dorsal and ventral carinae. Both cerci without medial tooth, but with one short row of small round denticles on the right cercus and two rows of small round denticles on the left cercus. Etymology This genus is dedicated to the memory of the American entomologist Mark Alan Muegge (pronounced Meggy), who died young due to an unfortunate car accident in 2015. He left behind a promising career and was devoted, among other groups, to diplurans; alongside other taxa, he provided a description of the highly cave-adapted japygid from Texas, Mixojapyx reddelli Muegge, 1992.Published as part of Sendra, Alberto, Komerički, Ana, Lips, Josiane, Luan, Yunxia, Selfa, Jesús & Jiménez-Valverde, Alberto, 2021, Asian cave-adapted diplurans, with the description of two new genera and four new species (Arthropoda, Hexapoda, Entognatha), pp. 1-46 in European Journal of Taxonomy 731 on pages 28-29, DOI: 10.5852/ejt.2021.731.1199, http://zenodo.org/record/442255

    Gollumjapyx Sendra & Ortuno

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    Gollumjapyx Sendra & Ortuño gen. nov. Etymology: The name of this genus comes from joining the prefix Gollum to the suffix Japyx. Gollum is the mythological character created by J.R.R. Tolkien, the dweller of the caves located below the Misty Mountains of Middle-earth. Description: Body elongate and restricted dorso-ventrally, unpigmented cuticle; legs, thoracic segments and urite X extremely elongate (Figs 1 and 2 a). Head. 53 to 55 antennomeres per antennae, manifesting 13 typical trichobothria (a clearly distal); 14 to 16 placoid sensilla over apical antonnomere, scattered in 2 groups and with at least one or two supernumerary placoid sensilla over the penultimate antennomere (Figs 3 a, b). Buccal pieces typical from this family with all five laminae pectinate of the maxillae. Thorax. Exceptionally elongate. Pronotum 6 + 6 M (5 + 5 M typical plus 1 + 1 M posterolateral) (Fig. 4 c). Mesonotum 5 + 5 M typical (Fig. 4 d). Metanotum 4 + 4 M (M 2 absents) (Fig. 4 e). Legs very elongate. Abdomen. Tergite I: 0+0 M. Tergite II: 1 + 1 M. Tergites III–VII: 5 + 5 M and 1 + 1 m 3 (Fig. 5 a). Tergite X with carinae distinct of subparallel margins with 4 + 4 M intracarinae (D) and 5 + 5 M carenae (L) (Fig. 3 c and 5 b). Urosternite I: 22 + 22 M (Fig. 6 a). Urosternite II-VII: 28 + 28 M. Simple lateral subcoxal organs showing one or two rows of very short subequal glandular setae and a row of sensory setae (Fig. 6 b). Median glandular organ with no apparent operculum and no pseudospori (Fig. 6 a). Styli and exsertile vesicles typical of this family. Cerci. (Fig 3 c, d and 5 b). Very protracted and robust with strongly sclerotized angles; slightly longer (in young specimens) or shorter (in adults) than the usually uncovered part of tergite X. Right cercus with a row of tubercles and simple denticles, with a strong sharp tooth clearly proximal; left cercus with double row of tubercles and postmedial tooth. Affinities: Gollumjapyx gen. n. shares certain affinities with the controversial genus Burmjapyx Silvestri, 1930. Since its description, the taxonomy of the Burmajapyx genus has undergone several difficulties. Paclt (1957) groups as Burmjapyx nine genera sharing the same model of cercal armature, a few of them have well defined characteristics and geographical distribution (Silvestri, 1948), such as: Protjapyx Silvestri, 1948 and some species of genus Megajapyx (Verhoeff, 1904), scattered around the Mediterranean region; Hapljapyx Silvestri, 1949 and Merojapyx Silvestri, 1948, found in the Neotropical region; Austrajapyx Silvestri, 1949, whose species range from the Neotropical region to the Ethiopean; Xenjapyx Silvestri, 1949, exclusive to the Ethiopean and, Henicjapyx Silvestri, 1949 from the Eastern region. Paclt (1957) grouped all the above with the true Burmjapyx and also included in this group some twenty more species, many of which belong to the heterogeneous genus Japyx Haliday, 1864. Pagés (1961, 1977, 1994, 2000) has repeatedly noted the artificiality of Burmjapyx sensu Paclt (1957), providing enough taxonomic evidence to return Burmjapyx to its initial taxonomic position. Gollumjapyx shares several features with the Burmjapyx s. str., particularly in the cercal armature and, to a lesser extent, also in the antennal supernumerary placoid sensilla, which are similar in distribution and number (in some specimens) to those described by Pagés (1977) for Burmjapyx inferus (Carpenter, 1932) found at Batu Caves (Malaysia). However, Gollumjapyx does not show a median glandular organ with setae-shaped sensilla (Silvestri’s “pseudospori”). In the new taxon, these sensilla appear on both sides of the central area. One more difference should be noted: a lack of setae with large setal socket preceding the glandular setae of the subcoxal organs, which are clearly recognisable in Burmjapyx. Among the species of Japyx included by Paclt (1957) in Burmjapyx only Japyx goliath (Parona, 1888) from Guatemala, described again later by Silvestri (1928), manifests affinities with the new taxon. Although the type material is unknown -almost certainly disappeared-, from Silvestri’s (1928) text and illustrations a few similarities with Gollumjapyx can be observed, for instance in the number of antennomeres, the robustness of the cercal armature and the morphology of the glandular organs in the first urosternite. In the future, with new specimens, the true meaning of these affinities could be resolved. The general lengthening of the body, thorax and legs in particular, is so extraordinary as to distinguish Gollumjapyx from other known genera of the Japygidae family. This troglobite appearance with such elongate body can only be compared to the description of two hypogean species, Troglojapyx hauseri and Mixojapyx reddelli (Muegge, 1992; Pagés, 1980); nevertheless Gollumjapyx smeagol shows a greater thinning of the body, more noticeable in the thoracic prescutum and legs. These traits suggest an adaptation of this new taxon to hypogeous life, as can be ascertained by the fact that they have been collected exclusively in caves. Type-species: Gollumjapyx smeagol sp. n. Sendra & OrtuñoPublished as part of Sendra, Alberto, Ortuño, Vicente M., Moreno, Agustín, Montagud, Sergio & Teruel, Santiago, 2006, Gollumjapyx smeagol gen. n., sp. n., an enigmatic hypogean japygid (Diplura: Japygidae) from the eastern Iberian Peninsula, pp. 35-52 in Zootaxa 1372 on pages 37-42, DOI: 10.5281/zenodo.17494

    Mueggejapyx Sendra & Komericki 2021, gen. nov.

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    Mueggejapyx Sendra & Komerički gen. nov. urn:lsid:zoobank.org:act: 924B9ABA-1630-4832-86EE-19CE73C3E7CB Type species Mueggejapyx brehieri Sendra & Komerički gen. et sp. nov. Diagnosis Large and elongated body; antennae with 36 antennomeres; trichobothria on antennomeres 4–6 in a 3/5/5 pattern; a trichobothria in distal position; 1–2 placoid sensilla on medial and distal antennomere; apical antennomere with 16–18 placoid sensilla; dorsal and ventral side of head with numerous uniformly distributed sM and sm, and several M; five laminae pectinate on maxilla; elongated thoracic segments; and cuticle with tiny micro-holes, pronotum and mesonotum each with 5+5 M, metanotum with 2+2 M; strong internal Y cuticular furcisternite structures in pro-presternites and pro-, meso- and metasternites; urotergites with at most 1+1 M (ma), 1+1 M 1, 1+1 M 2–5; urite X longer than wide, with marked carinae with subparallel margins, with 2+2 intracarinal macrosetae; acropygium rounded; lateral angles of urotergites VI and VII with weakly to moderate lobiform projection; first urosternite with multiperforated surface and rounded protrusions, with 3+3 M on prescutum and 11+11 M on scutum; median glandular organ with seta-shaped sensilla; lateral subcoxal organs with one to three rows of short glandular setae (GS), one row of sensory setae (SS), and a row of setae with large socket; sternites with abundant sm and strong M; cerci strong, elongated, rectilinear and becoming curved subapically, heavily sclerotized with external dorsal and ventral carinae. Both cerci without medial tooth, but with one short row of small round denticles on the right cercus and two rows of small round denticles on the left cercus. Etymology This genus is dedicated to the memory of the American entomologist Mark Alan Muegge (pronounced Meggy), who died young due to an unfortunate car accident in 2015. He left behind a promising career and was devoted, among other groups, to diplurans; alongside other taxa, he provided a description of the highly cave-adapted japygid from Texas, Mixojapyx reddelli Muegge, 1992.Published as part of Sendra, Alberto, Komerički, Ana, Lips, Josiane, Luan, Yunxia, Selfa, Jesús & Jiménez-Valverde, Alberto, 2021, Asian cave-adapted diplurans, with the description of two new genera and four new species (Arthropoda, Hexapoda, Entognatha), pp. 1-46 in European Journal of Taxonomy 731 on pages 28-29, DOI: 10.5852/ejt.2021.731.1199, http://zenodo.org/record/442255

    Cestocampa iberica Sendra & Conde, new species

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    <i>Cestocampa iberica</i> Sendra & Condé new species <p> <b>Diagnosis.</b> Epicuticle without ornamentation. Antennae with 34 to 41 antennomeres, cupuliform organ with 8‒10 sensilla. Sensillum of third antennomere in sternal position. Frontal process not developed. Notal macrochaetae: pronotum with 1+1 <i>ma</i>, 3+3 <i>la1,2,3</i>, 2+2 <i>lp2,3</i>; mesonotum with 1+1 <i>ma</i>, 3+3 <i>la1,2,3</i>, 2+2 <i>lp2,3</i> and 1+1 <i>mp</i>; metanotum with 1+1 <i>ma</i>, 2+2 <i>la1,2</i>, 2+2 <i>lp2,3</i>, 1+1 <i>mp</i>. Elongated legs, femur III with one tergal macrochaeta and tibiae with two sternal macrochaetae. Abdominal macrochaetae: urotergite I with 1+1 <i>post1</i>, II with 2+2 <i>post1,3</i>, III 1 +1 <i>la3</i>, 2+2 <i>post1,3</i>, IV 1 +1 <i>la3</i>, 4+4 <i>post1,3,4,5</i>, V-VII 3+3 <i>la1,2,3</i>, 5+5 <i>post1 to 5</i>, VIII 7 +7 <i>post</i> and IX 9 +9 <i>post</i>. Urosternite I with up to 30 macrochaetae (hypertrichia); urosternites II to VII with 5+5 and VIII with 2+2 macrochaetae. Urosternite I of male without glandular <i>g1-</i> setae; subcylindrical appendages with glandular <i>a1</i> -setae and <i>a2</i> -setae; female with glandular <i>a1</i> -setae. Cerci longer than the body, but less than twice as long.</p> <p> <b>Description.</b> Lengths of body 3.9 to 5.6 mm (males), 3.6‒7.0 mm (females). Cercal length in adults 4.4 to 7.0 mm (n = 4), cercal and body lengths approximately correlated. Epicuticle without ornamentation; clothing setae sparse, glabrous or with short distal barbs.</p> <p> Antennae with 34 to 41 antennomeres: 34, 35, 36 (1 antenna each); 37 (3 antennae); 38 (7 antennae); 39 (2 antennae); 40, 41 (1 antenna each); regenerated antennae with 28, 31 and 33 antennomeres. Sensillum of third antennomere subcylindrical, short, located ventral between macrochaetae <i>d</i> and <i>e</i>. Central antennomeres 2.5× longer than wide. Apical antennomere about 2.3× longer than wide, its cupuliform organ occupying one-sixth of the length, containing 8 to 10 sensilla with 2 or 3 collarettes, each with a reticulated surface (Fig. 2 B). Sensilla well developed, in cross-section concave-convex (Fig. 2 A); one sensillum on the third antennomere, increasing gradually to about 6 sensilla in the medial antennomeres, to 10 sensilla in the distal antennomeres; on each antennomere the sensilla arranged in distal verticils. Frontal process not developed, with an barbed apical macrochaeta along the distal four-fifths and with several setae with a few barbs. Three macrochaetae along the line of insertion of the antennae and <i>x</i> setae with the following relative lengths: <i>a</i> = 0.53, <i>p</i> = 0.64, <i>i</i> = 0.79, <i>x</i> = 1. Labial palp suboval, with subcylindrical lateral-external sensilla similar to sensilla of maxillary palp and third antennomere, two guard setae, up to 11 setae on the anterior border and 150 neuroglandular setae.</p> <p> Thoracic nota elongated. Distribution of macrochaetae: pronotum with 1+1 <i>ma</i>, 3+3 <i>la1,2,3</i>, 2+2 <i>lp2,3</i> (Fig. 3A); mesonotum with 1+1 <i>ma</i>, 3+3 <i>la1,2,3</i> (3+2 on a female of 7 mm), 2+2 <i>lp2,3</i> and 1+1 <i>mp</i>; metanotum with 1+1 <i>ma</i>, 2+2 <i>la1,2</i>, 2+2 <i>lp2,3</i>, 1+1 <i>mp</i>. All macrochaetae thin and short with thin barbs along distal two-thirds. Pronotum with 1+1 well developed setae near center of the notum, <i>la</i> 1 and <i>la</i> 2 less differentiated than the other macrochaetae. Marginal setae thin, barbed along their distal half to two-thirds. Elongated legs, length of metathoracic leg reaching or exceeding abdominal segment X. Femur III with long dorsal macrochaeta with thin barbs except basally, inserted near middle of femur. Ventral femoral macrochaeta shorter and more distal than the dorsal macrochaetae. Usually two well-barbed ventral macrochaetae on tibiae I to III; in a few specimens tibia III with 3 or 4 ventral macrochaetae. All tarsal setae barbed, dorsal setae (<i>sda, sdp</i>) and the 3 lateral setae (<i>sla</i>) barbed in the middle, distal and proximal portions glabrous (Fig. 2 C, D), subapical ventral setae thinner than the others and completely barbed. Subequal claws with ridged lateral crests, without elongated talon. Telotarsal process laminar with dense sternal barbs (Fig. 2 C, D).</p> <p> Usual distribution of abdominal macrochaetae on tergites as follows: tergite I 1 +1 <i>post1</i>, II 2 +2 <i>post1,3</i>, III 1 +1 <i>la3</i>, 2+2 <i>post1,3</i>, IV 1 +1 <i>la3</i>, 4+4 <i>post1,3,4,5</i>, V-VII 3+3 <i>la1,2,3</i>, 5+5 <i>post1 to 5</i>, VIII 7 +7 <i>post</i>, IX 9 +9 <i>post</i>. In some specimens the number of macrochaetae show some variations; for instance, two of the males of Sima del Campillo and Avenc Vinalopó with 2+1 and 2+2 <i>post</i> on urotergite I; one female from Cueva del Tornero with 0+1 <i>la</i> and 2+1 <i>post</i> on urotergite II; one female from Cueva Hermosa with 4+4 <i>post</i> on urotergites V and VI; one male from Avenc Vinalopó and one female of Cova de Galindo with 2+2 <i>la</i> on tergite IV. Urosternite I (Fig. 3B) with up to 30 macrochaetae (hypertrichia); urosternites II to VII with 5+5 macrochaetae, urosternite VIII with 2+2 macrochaetae. Apical setae of styli with basal tooth and 10 to15 distal barbs; subapical and medial sternal setae completely covered with thin barbs. Each cercus with a base divided into 2 or 3 secondary articles, followed by 8 or 12 primary articles.</p> <p> Male urosternite I without glandular setae on posterior margin (Fig. 3B). Subcylindrical appendages of urosternite I bearing 8 to10 glandular <i>a1</i> -setae on their apex and a lateral-internal field of 16 to 88 longer, thinner glandular <i>a2</i> -setae. Spiral of spermatozoid bundles 70 to 90 µm long with one and one-half turns; wider section 4.5 µm. Spiral filament broadly rounded on one end and pointed at other end; pointed end divided or entire (Fig. 3C, D).</p> <p> Female subcylindrical appendages on the urosternite I similar to those of male but narrower, shorter (12 to 20 µm) and with more glandular <i>a1</i> -setae (30‒40).</p> <p> <b>Material examined.</b> Nine males, 18 females 3 juveniles from Spain as follows:</p> <p> Valencia Community, Alicante, Bocairent, Avenc del Vinalopó (elev. 840 m), 1 ♂, 28.XII.1969, L. Auroux, O. Escolà & F. Español leg.; Cova de la Sarsa (Elev. 700 m), 1 ♂, 1.IX.1978, J.V. Gonzalez leg.; 3 ♂, 6 ♀, 19.V.2007, S. Teruel & A. Sendra leg.; Cocentaina, Cova de les Meravelles (elev. 1070 m), 1 ♂, 2.XII.1979, J.V. Gonzalez <b>FIGURE 3.</b> <i>Cestocampa iberica</i> <b>n. sp.</b> A. Pronotum. B. Urosternite I of the male (holotype). C. Spiral filament of spermatozoid bundles, from a male of Tunel dels Sumidors. D. Spiral filament of spermatozoid bundles, from a male of Cueva del Tornero. (<i>a1, a2</i> = glandular setae of the appendages of the urosternite I; <i>l</i> a = lateral anterior macrochaetae; <i>lp</i> = lateral posterior macrochaetae; <i>ma</i> = medial anterior macrochaetae). Scales: A and B, 50 m; C and D, 80 mm.</p> <p>leg.; 1 juvenile, 19.III.2003, V. Ortuño leg.; Onil, Cova de Galindo (elev. 1035 m), 2 ♀, 11.VIII.1983, J.A. Zaragoza & A. Sendra leg.; Valencia, Chera, Cova de l’Or (elev. 798 m), 1 ♀, 5.V.2009, S. Teruel & A. Sendra leg.; Millares, Cova Dones (elev. 550 m), 1 ♀, 4.VII.2005, A. Sendra leg.; Vallada, Túnel dels Sumidors (elev. 550 m), 1 ♂, 3 ♀, 13.VI.1982, J. Jornet leg.; 1 ♀, 4.I.1981, A. Sendra leg.; Túnel dels Sumidors, 1 ♀, III.1983, R. Lluch leg.; Tous, Sima del Campillo (elev. 430 m.), 1 ♂, 6.IX.1981, O. Escolà leg.; Cortés de Pallás, Cueva Hermosa (elev. 645 m), 1 ♀, 16.V.80, A. Sendra leg. Zarra, Cueva de la Hoz (elev. 550 m), 1 ♀, 2 juveniles, 5.V.2001, A. Sendra & S. Teruel leg.; Castilla-La Mancha, Guadalajara, Checa, Cueva del Tornero (elev. 1335 m), 1 ♂, 2 ♀, 29.IX.1971, J. Pallisé leg. Total: 9 males, 18 females and 3 immatures.</p> <p> <b>Type material.</b> Holotype, male “Cova de la Sarsa”, Bocairent, Alicante, Spain, 19-V-2007, S. Teruel & A. Sendra leg., preserved in 70 % ethanol, deposited in the Museu Valencià d’Història Natural. Paratypes, 2 males and 6 females collected with the holotype, preserved in 70% ethanol, deposited in the Museu Valencià d’Història Natura, Muséum d’Histoire naturelle de Genève.</p> <p> <b>Etymology.</b> This species is named for the Iberian Peninsula, where this species is distributed along the Iberian and Prebetic Mountains..</p> <p> <b>Geographic distribution.</b> Caves located in the large karstic area occurring along the Castilian-Valencian Branch of the Iberian Mountain Range, which extends to the northwestern Prebetic Mountains and is bordered by Tertiary and Quaternary basins.</p> <p> <b>Affinities.</b> <i>Cestocampa iberica</i> <b>n. sp.</b> closely resembles <i>C. gasparoi</i> and <i>C. balcanica</i>. The three species share the presence of <i>mp</i> macrochaetae on the meso- and metanotum and the chaetotaxy of the notal and urotergal macrochaetae. In addition, <i>C. gasparoi</i> and <i>C. italica</i> lack a glandular field on the posterior border of urosternite I, but show a lateral-internal glandular setae field on the appendages of the males. <i>Cestocampa iberica</i> <b>n. sp.</b> differs from <i>C. gasparoi</i> by the number of hypertrichia on urosternite I (up to 30 macrochaetae <i>vs.</i> 7+7 or 7+1+ 7 in <i>C. gasparoi</i>) and the number of macrochaetae on urotergites VIII and IX i (7+7, 9+9, <i>vs.</i> 6+6, 8+ 8 in <i>C. gasparoi</i> and <i>C. balcanica</i>; character unknown in <i>C. italica</i>). Further differences are closely related to the more progressive troglomorphy (<i>sensu</i> Christiansen, 2005) of <i>C. iberica</i> n. sp. (longer appendages and increased number of antennomeres and cercal articles, and complexity and high number of sensilla in the cupuliform organ).</p>Published as part of <i>Sendra, Alberto, Arnedo, Miquel A. & Ribera, Carles, 2012, Revision of Cestocampa Condé (Diplura, Campodeidae), with description of a new species from caves in the eastern Iberian Peninsula, pp. 43-56 in Zootaxa 3252</i> on pages 47-50, DOI: <a href="http://zenodo.org/record/210286">10.5281/zenodo.210286</a&gt

    Cestocampa iberica Sendra & Conde, new species

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    <i>Cestocampa iberica</i> Sendra & Condé new species <p> <b>Diagnosis.</b> Epicuticle without ornamentation. Antennae with 34 to 41 antennomeres, cupuliform organ with 8‒10 sensilla. Sensillum of third antennomere in sternal position. Frontal process not developed. Notal macrochaetae: pronotum with 1+1 <i>ma</i>, 3+3 <i>la1,2,3</i>, 2+2 <i>lp2,3</i>; mesonotum with 1+1 <i>ma</i>, 3+3 <i>la1,2,3</i>, 2+2 <i>lp2,3</i> and 1+1 <i>mp</i>; metanotum with 1+1 <i>ma</i>, 2+2 <i>la1,2</i>, 2+2 <i>lp2,3</i>, 1+1 <i>mp</i>. Elongated legs, femur III with one tergal macrochaeta and tibiae with two sternal macrochaetae. Abdominal macrochaetae: urotergite I with 1+1 <i>post1</i>, II with 2+2 <i>post1,3</i>, III 1 +1 <i>la3</i>, 2+2 <i>post1,3</i>, IV 1 +1 <i>la3</i>, 4+4 <i>post1,3,4,5</i>, V-VII 3+3 <i>la1,2,3</i>, 5+5 <i>post1 to 5</i>, VIII 7 +7 <i>post</i> and IX 9 +9 <i>post</i>. Urosternite I with up to 30 macrochaetae (hypertrichia); urosternites II to VII with 5+5 and VIII with 2+2 macrochaetae. Urosternite I of male without glandular <i>g1-</i> setae; subcylindrical appendages with glandular <i>a1</i> -setae and <i>a2</i> -setae; female with glandular <i>a1</i> -setae. Cerci longer than the body, but less than twice as long.</p> <p> <b>Description.</b> Lengths of body 3.9 to 5.6 mm (males), 3.6‒7.0 mm (females). Cercal length in adults 4.4 to 7.0 mm (n = 4), cercal and body lengths approximately correlated. Epicuticle without ornamentation; clothing setae sparse, glabrous or with short distal barbs.</p> <p> Antennae with 34 to 41 antennomeres: 34, 35, 36 (1 antenna each); 37 (3 antennae); 38 (7 antennae); 39 (2 antennae); 40, 41 (1 antenna each); regenerated antennae with 28, 31 and 33 antennomeres. Sensillum of third antennomere subcylindrical, short, located ventral between macrochaetae <i>d</i> and <i>e</i>. Central antennomeres 2.5× longer than wide. Apical antennomere about 2.3× longer than wide, its cupuliform organ occupying one-sixth of the length, containing 8 to 10 sensilla with 2 or 3 collarettes, each with a reticulated surface (Fig. 2 B). Sensilla well developed, in cross-section concave-convex (Fig. 2 A); one sensillum on the third antennomere, increasing gradually to about 6 sensilla in the medial antennomeres, to 10 sensilla in the distal antennomeres; on each antennomere the sensilla arranged in distal verticils. Frontal process not developed, with an barbed apical macrochaeta along the distal four-fifths and with several setae with a few barbs. Three macrochaetae along the line of insertion of the antennae and <i>x</i> setae with the following relative lengths: <i>a</i> = 0.53, <i>p</i> = 0.64, <i>i</i> = 0.79, <i>x</i> = 1. Labial palp suboval, with subcylindrical lateral-external sensilla similar to sensilla of maxillary palp and third antennomere, two guard setae, up to 11 setae on the anterior border and 150 neuroglandular setae.</p> <p> Thoracic nota elongated. Distribution of macrochaetae: pronotum with 1+1 <i>ma</i>, 3+3 <i>la1,2,3</i>, 2+2 <i>lp2,3</i> (Fig. 3A); mesonotum with 1+1 <i>ma</i>, 3+3 <i>la1,2,3</i> (3+2 on a female of 7 mm), 2+2 <i>lp2,3</i> and 1+1 <i>mp</i>; metanotum with 1+1 <i>ma</i>, 2+2 <i>la1,2</i>, 2+2 <i>lp2,3</i>, 1+1 <i>mp</i>. All macrochaetae thin and short with thin barbs along distal two-thirds. Pronotum with 1+1 well developed setae near center of the notum, <i>la</i> 1 and <i>la</i> 2 less differentiated than the other macrochaetae. Marginal setae thin, barbed along their distal half to two-thirds. Elongated legs, length of metathoracic leg reaching or exceeding abdominal segment X. Femur III with long dorsal macrochaeta with thin barbs except basally, inserted near middle of femur. Ventral femoral macrochaeta shorter and more distal than the dorsal macrochaetae. Usually two well-barbed ventral macrochaetae on tibiae I to III; in a few specimens tibia III with 3 or 4 ventral macrochaetae. All tarsal setae barbed, dorsal setae (<i>sda, sdp</i>) and the 3 lateral setae (<i>sla</i>) barbed in the middle, distal and proximal portions glabrous (Fig. 2 C, D), subapical ventral setae thinner than the others and completely barbed. Subequal claws with ridged lateral crests, without elongated talon. Telotarsal process laminar with dense sternal barbs (Fig. 2 C, D).</p> <p> Usual distribution of abdominal macrochaetae on tergites as follows: tergite I 1 +1 <i>post1</i>, II 2 +2 <i>post1,3</i>, III 1 +1 <i>la3</i>, 2+2 <i>post1,3</i>, IV 1 +1 <i>la3</i>, 4+4 <i>post1,3,4,5</i>, V-VII 3+3 <i>la1,2,3</i>, 5+5 <i>post1 to 5</i>, VIII 7 +7 <i>post</i>, IX 9 +9 <i>post</i>. In some specimens the number of macrochaetae show some variations; for instance, two of the males of Sima del Campillo and Avenc Vinalopó with 2+1 and 2+2 <i>post</i> on urotergite I; one female from Cueva del Tornero with 0+1 <i>la</i> and 2+1 <i>post</i> on urotergite II; one female from Cueva Hermosa with 4+4 <i>post</i> on urotergites V and VI; one male from Avenc Vinalopó and one female of Cova de Galindo with 2+2 <i>la</i> on tergite IV. Urosternite I (Fig. 3B) with up to 30 macrochaetae (hypertrichia); urosternites II to VII with 5+5 macrochaetae, urosternite VIII with 2+2 macrochaetae. Apical setae of styli with basal tooth and 10 to15 distal barbs; subapical and medial sternal setae completely covered with thin barbs. Each cercus with a base divided into 2 or 3 secondary articles, followed by 8 or 12 primary articles.</p> <p> Male urosternite I without glandular setae on posterior margin (Fig. 3B). Subcylindrical appendages of urosternite I bearing 8 to10 glandular <i>a1</i> -setae on their apex and a lateral-internal field of 16 to 88 longer, thinner glandular <i>a2</i> -setae. Spiral of spermatozoid bundles 70 to 90 µm long with one and one-half turns; wider section 4.5 µm. Spiral filament broadly rounded on one end and pointed at other end; pointed end divided or entire (Fig. 3C, D).</p> <p> Female subcylindrical appendages on the urosternite I similar to those of male but narrower, shorter (12 to 20 µm) and with more glandular <i>a1</i> -setae (30‒40).</p> <p> <b>Material examined.</b> Nine males, 18 females 3 juveniles from Spain as follows:</p> <p> Valencia Community, Alicante, Bocairent, Avenc del Vinalopó (elev. 840 m), 1 ♂, 28.XII.1969, L. Auroux, O. Escolà & F. Español leg.; Cova de la Sarsa (Elev. 700 m), 1 ♂, 1.IX.1978, J.V. Gonzalez leg.; 3 ♂, 6 ♀, 19.V.2007, S. Teruel & A. Sendra leg.; Cocentaina, Cova de les Meravelles (elev. 1070 m), 1 ♂, 2.XII.1979, J.V. Gonzalez <b>FIGURE 3.</b> <i>Cestocampa iberica</i> <b>n. sp.</b> A. Pronotum. B. Urosternite I of the male (holotype). C. Spiral filament of spermatozoid bundles, from a male of Tunel dels Sumidors. D. Spiral filament of spermatozoid bundles, from a male of Cueva del Tornero. (<i>a1, a2</i> = glandular setae of the appendages of the urosternite I; <i>l</i> a = lateral anterior macrochaetae; <i>lp</i> = lateral posterior macrochaetae; <i>ma</i> = medial anterior macrochaetae). Scales: A and B, 50 m; C and D, 80 mm.</p> <p>leg.; 1 juvenile, 19.III.2003, V. Ortuño leg.; Onil, Cova de Galindo (elev. 1035 m), 2 ♀, 11.VIII.1983, J.A. Zaragoza & A. Sendra leg.; Valencia, Chera, Cova de l’Or (elev. 798 m), 1 ♀, 5.V.2009, S. Teruel & A. Sendra leg.; Millares, Cova Dones (elev. 550 m), 1 ♀, 4.VII.2005, A. Sendra leg.; Vallada, Túnel dels Sumidors (elev. 550 m), 1 ♂, 3 ♀, 13.VI.1982, J. Jornet leg.; 1 ♀, 4.I.1981, A. Sendra leg.; Túnel dels Sumidors, 1 ♀, III.1983, R. Lluch leg.; Tous, Sima del Campillo (elev. 430 m.), 1 ♂, 6.IX.1981, O. Escolà leg.; Cortés de Pallás, Cueva Hermosa (elev. 645 m), 1 ♀, 16.V.80, A. Sendra leg. Zarra, Cueva de la Hoz (elev. 550 m), 1 ♀, 2 juveniles, 5.V.2001, A. Sendra & S. Teruel leg.; Castilla-La Mancha, Guadalajara, Checa, Cueva del Tornero (elev. 1335 m), 1 ♂, 2 ♀, 29.IX.1971, J. Pallisé leg. Total: 9 males, 18 females and 3 immatures.</p> <p> <b>Type material.</b> Holotype, male “Cova de la Sarsa”, Bocairent, Alicante, Spain, 19-V-2007, S. Teruel & A. Sendra leg., preserved in 70 % ethanol, deposited in the Museu Valencià d’Història Natural. Paratypes, 2 males and 6 females collected with the holotype, preserved in 70% ethanol, deposited in the Museu Valencià d’Història Natura, Muséum d’Histoire naturelle de Genève.</p> <p> <b>Etymology.</b> This species is named for the Iberian Peninsula, where this species is distributed along the Iberian and Prebetic Mountains..</p> <p> <b>Geographic distribution.</b> Caves located in the large karstic area occurring along the Castilian-Valencian Branch of the Iberian Mountain Range, which extends to the northwestern Prebetic Mountains and is bordered by Tertiary and Quaternary basins.</p> <p> <b>Affinities.</b> <i>Cestocampa iberica</i> <b>n. sp.</b> closely resembles <i>C. gasparoi</i> and <i>C. balcanica</i>. The three species share the presence of <i>mp</i> macrochaetae on the meso- and metanotum and the chaetotaxy of the notal and urotergal macrochaetae. In addition, <i>C. gasparoi</i> and <i>C. italica</i> lack a glandular field on the posterior border of urosternite I, but show a lateral-internal glandular setae field on the appendages of the males. <i>Cestocampa iberica</i> <b>n. sp.</b> differs from <i>C. gasparoi</i> by the number of hypertrichia on urosternite I (up to 30 macrochaetae <i>vs.</i> 7+7 or 7+1+ 7 in <i>C. gasparoi</i>) and the number of macrochaetae on urotergites VIII and IX i (7+7, 9+9, <i>vs.</i> 6+6, 8+ 8 in <i>C. gasparoi</i> and <i>C. balcanica</i>; character unknown in <i>C. italica</i>). Further differences are closely related to the more progressive troglomorphy (<i>sensu</i> Christiansen, 2005) of <i>C. iberica</i> n. sp. (longer appendages and increased number of antennomeres and cercal articles, and complexity and high number of sensilla in the cupuliform organ).</p>Published as part of <i>Sendra, Alberto, Arnedo, Miquel A. & Ribera, Carles, 2012, Revision of Cestocampa Condé (Diplura, Campodeidae), with description of a new species from caves in the eastern Iberian Peninsula, pp. 43-56 in Zootaxa 3252</i> on pages 47-50, DOI: <a href="http://zenodo.org/record/210286">10.5281/zenodo.210286</a&gt

    Figures 3-4 from: Sendra A, Sket B, Stoev P (2017) A striking new genus and species of troglobitic Campodeidae (Diplura) from Central Asia. Subterranean Biology 23: 47-68. https://doi.org/10.3897/subtbiol.23.14631

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    Figures 3-4 - Turkmenocampa mirabilis Sendra & Stoev, sp. n. 3 Pro-, meso- and metanotum, left side, holotype 4 Urotergites I-IX, right side, holotype. Scale bars: 0.2 m

    Moore–Penrose approach in the Hough transform framework

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    Let F(x, a) be a real polynomial in two sets of variables, x and a, that is linear with respect to one of the variable sets, say a. In this paper, we deal with two of the main steps of the Hough transform framework for the pattern recognition technique to detect loci in images. More precisely, we present an algorithmic process, based on the Moore–Penrose pseudo-inverse, to provide a region of analysis in the parameter space. In addition, we state an upper bound for the sampling distance of the discretization of the parameter space region

    Campodea (Dicampa) azkarraga Sendra 2006

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    Campodea (Dicampa) azkarraga Sendra, 2006 Material examined. 1 Ψ, Duzce, close to Yıġılca, 36 T 3753404536201, 304 m, 13 -VIII- 2005, S. Teruel leg.; 1 Ψ, Gümüshane, Çayra, alder leaf litter, 19 -VIII- 2005, A. Moreno leg. These new collections strengthen the status of this species, which was recently described (Sendra et al. 2006) from Pontic Mountains, in the Turkish Black Sea region.Published as part of Tusun, Sadreddin & Özbay, Cengizhan, 2010, New species, new records, and distribution of Campodeidae (Diplura) in Anatolia, pp. 40-52 in Zootaxa 2639 on page 49, DOI: 10.5281/zenodo.19853

    Imazighenjapyx Sendra & Sanchez-Garcia 2023, gen. nov.

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    Genus <i>Imazighenjapyx</i> Sendra & Sánchez-García gen. nov. <p>urn:lsid:zoobank.org:act: ADDB00E9-BDE5-4007-82AB-2A2A8B3F5572</p> Type species <p> <i>Imazighenjapyx marocanus</i> Sendra & Sánchez-García gen. et sp. nov.</p> Diagnosis <p>Body large and elongate; epicuticle reticulate; abdominal segment X with micropores at high magnification. Antennae with 41 antennomeres; medial and distal antennomeres with a few ms and abundant s setae, plus two whorls of micro-barbed sM setae; apical antennomere with abundant placoid sensilla (up to 24). Pronotum, mesonotum, and metanotum with 5+5 M1−5. Prosternum with up to 72 M; meso-poststernum with up to 19 M; meso-intersternum with up to 22 M; mesosternum with up to 84 M; meta-poststernum with up to 26 M; meta-intersternum with up to 21 M; and metasternum with up to 99 M; scutum urotergite I with 1+1 M5; urotergite II with 10 M; urotergites III‒VII with 12 M; urite X with 17 M, ventral side with 52 M setae; urosternite I with up to 120 M setae on scutum, plus 200 sM setae on posterior position. Median glandular organ with abundant pseudospores; each lateral subcoxal organ with one row of glandular setae and one row of sensory setae; urosternites II‒III with about 140 M setae; urosternites IV‒VII with about 160 M setae; urosternite VIII with about 50 M setae between two well-defined carinae; cerci asymmetric with subsymmetric teeth, rectilinear along the proximal half, and curved in the distal half. Cerci with concave top side and with distal end up; right cercus with pointed medial tooth, predental margin bearing two rows of denticles; very protruded postdental margin looking like a scraper shape with a row of denticles; left cercus predental margin with three rows, postdental margin with small round denticles ending before the hook.</p> Etymology <p> The genus name is a combination of the prefix <i>Imazighen</i> and the suffix <i>Japyx</i>. Berbers call themselves Imazighen, which means ‘free men’ or ‘noble men’.</p>Published as part of <i>Sendra, Alberto, Sánchez-García, Alba, Hoch, Hannelore, Jiménez-Valverde, Alberto, Selfa, Jesús, Moutaouakil, Soumia, Preez, Gerhard Du, Millar, Ian & Ferreira, Rodrigo Lopes, 2023, Life in darkness: an overview of cave-adapted japygids (Hexapoda, Diplura), pp. 1-54 in European Journal of Taxonomy 894 (1)</i> on page 12, DOI: 10.5852/ejt.2023.894.2287, <a href="http://zenodo.org/record/8388995">http://zenodo.org/record/8388995</a&gt
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