196,469 research outputs found
Mecodema ponaiti Seldon & Leschen, 2011, sp. n.
Mecodema ponaiti, sp. n. Figures: 5, 9 b, 11 b, 13 a, 25 a. Diagnosis. Head broad but convex. Clypeus with a single setose puncture on each side, bearing 2 setae. Prothoracic carina broad, crenulation absent and extended to anterior angle. Pronotum fovea shallow and narrow. Elytral striae with regularly distributed, small asetose punctures. Setose punctures absent on 3 rd, 4 th and 5 th ventrites. Apical portion of aedeagus symmetrically rounded with a strong twist to right of vertical. Dorso-apical portion of central spicule, greatly expanded. Basal lobe of left paramere rectangular, arm and apical lobe relatively large, and with setation along apical ¾ of ventral edge. Description. Length 20–26 mm. Colour black. Head broad but convex. Microsculpture of frons and vertex absent. Vertexal groove broad laterally and narrow middle. Form of microsculpture on anterior area of clypeus, spatulate. Clypeus with single setose puncture on each side, bearing 2 setae (or 2 setose punctures on each side bearing a single seta each*). Form of lobes of the mentum, square. Submentum constriction broad, with 6 setae. Prothoracic carina broad, crenulation absent, and extended to anterior angle (or extending to beyond anterior angle*). Pronotum broad and flat, posterior lateral sinuation angle parallel, pronotal fovea shallow and narrow, microsculpture of pronotal disc absent. Elytra broad but convex, striae with regularly distributed small asetose punctures, intervals slightly convex. Setose punctures of mesocoxae present, absent metacoxae. Setose punctures absent 3 rd, 4 th and 5 th ventrites. Apical portion of aedeagus symmetrically rounded with a strong twist to right of vertical axis. Dorso-apical portion of central spicule, greatly expanded. Left paramere basal lobe rectangular, arm and apical lobe relatively large, and with setation along apical ¾ of ventral edge (or setation along apical ½ of ventral edge*). Comments. Britton (1949) placed specimens from the Poor Knights Islands and Whangarei area under M. atrox, which seemed to be the “default” species for Northland specimens. M. ponaiti is sister species to the remaining curvidens species and is endemic to the Poor Knights Islands. The similarities between the M. ponaiti and M. manaia species can be seen in one of the two variants of the island group. The variant from Aorangi Is. has similar external morphological features to those of M. manaia (Whangarei Heads), whereas the Tawhiti Rahi Is. variant exhibits similarities in external morphology to M. regulus (Three Kings Islands), especially the pronotal disc, which is more rounded laterally than the mainland species. In the description the difference in character states between the two variants are indicated by an asterisk (*). Several specimens of M. ponaiti have been collected in association with seabird burrows, either resting under logs in close proximity or from within the burrows themselves. Distribution. North Island: Northland (ND), Poor Knights Islands (only). Holotype: Male labelled (NZAC) NEW ZEALAND ND Poor Knights Is. Tawhiti Rahi Is. 18 Dec 2009 D.S. Seldon, R.A.B. Leschen & T. Buckley / HOLOTYPE Mecodema ponaiti n. sp. design. by Seldon & Leschen 2010 [red label]. Paratype material examined: 1, NEW ZEALAND ND Poor Knights Aorangi Is in rodent bait station 3 Aug 2002, I. Hogarth / AMNZ 54564 Auckland Museum New Zealand / PARATYPE Mecodema ponaiti design. by Seldon & Leschen 2010; 1, NEW ZEALAND. ND Poor Knights Is. Tawhiti Rahi 15 Apr 1996 I.A.N. Stringer / AMNZ 25709 Auckland Museum New Zealand / PARATYPE Mecodema ponaiti design. by Seldon & Leschen 2010; 1, NEW ZEALAND ND Poor Knights Is. Aorangi Is. 16 Apr 1996 Richard Parrish / AMNZ 33511 Auckland Museum New Zealand / PARATYPE Mecodema ponaiti design. by Seldon & Leschen 2010; 1, TAWHITI RAHI POOR KNIGHTS IS. 11.31. 45 Coll. Auk. Museum / MECODEMA n.sp. Det. J.I. Townsend 1969 / AMNZ 24110 Auckland Museum New Zealand / PARATYPE Mecodema ponaiti design. by Seldon & Leschen 2010; 1, N. Island Poor Knights in soil P. bulleri burrow 24 / 3 / 1954 E.G. Turbott / Mecodema sp. nr. atrox Det. J.C. Watt / MECODEMA n. sp. Det. J.I. Townsend 1969 / AMNZ 63295 Auckland Museum New Zealand / PARATYPE Mecodema ponaiti design. by Seldon & Leschen 2010; 4, NEW ZEALAND ND Poor Knights Is Tawhiti rahi 2 Dec 1980 G. Kuschel / PARATYPE Mecodema ponaiti design. by Seldon & Leschen 2010; 4, Tawhiti Rahi under rock 21 - 1-86 J.C. Watt / Mecodema atrox? E.B. Britton det. 1956 / J.I. Townsend Collection / PARATYPE Mecodema ponaiti design. by Seldon & Leschen 2010 (NZAC); 2, Tawhiti Rahi on ground 1 - 1-56 J.C. Watt / Mecodema atrox? E.B. Britton 1956 / J.I. Townsend Collection / PARATYPE Mecodema ponaiti design. by Seldon & Leschen 2010 (NZAC); 5, Tawhiti Rahi under rock 1 - 1-56 J.C. Watt / Collection J.C.Watt / J.I. Collection / PARATYPE Mecodema ponaiti design. by Seldon & Leschen 2010 (NZAC); 2, Forest Litter Tawhiti Rahi Poor Knights Is 10.Aug. 1970 B.M. May / PARATYPE Mecodema ponaiti design. by Seldon & Leschen 2010 (NZAC); 1, NEW ZEALAND ND Poor Knights Is Tawhiti Rahi Plateau 200m 3–10 Dec 1980 / J.C. Watt pit trap 80 / 179 / PARA- TYPE Mecodema ponaiti design. by Seldon & Leschen 2010 (NZAC); 1, NEW ZEALAND ND Poor Knights Is Tawhiti Rahi Plateau 190m 3–10 Dec 1980 / J.C. Watt pit trap 80 / 184 / PARATYPE Mecodema ponaiti design. by Seldon & Leschen 2010 (NZAC); 1, NEW ZEALAND ND Poor Knights Is Tawhiti Rahi Plateau 180m 3–10 Dec 1980 / J.C. Watt pit trap 80 / 185 / PARATYPE Mecodema ponaiti design. by Seldon & Leschen 2010 (NZAC); 2, NEW ZEALAND ND Poor Knights Is Tawhiti Rahi 200m 3–10 Dec 1980 / J.C. Watt pit trap 80 / 177 / PARATYPE Mecodema ponaiti design. by Seldon & Leschen 2010 (NZAC); 1, NEW ZEALAND ND Poor Knights Is Tawhiti Rahi 2–10 Dec 1980 / J.C. Watt pit trap 80 / 176 / PARATYPE Mecodema ponaiti design. by Seldon & Leschen 2010 (NZAC); 2, NEW ZEALAND ND Poor Knights Is Tawhiti Rahi Shag Bay 3–10 Dec 1980 / J.C. Watt pit trap 80 / 165 / PARATYPE Mecodema ponaiti design. by Seldon & Leschen 2010 (NZAC); 1, NEW ZEALAND ND Poor Knights Is Tawhiti Rahi Shag Bay 8–12 Dec 1980 / J.C. Watt / PARATYPE Mecodema ponaiti design. by Seldon & Leschen 2010 (NZAC); 1, Poor Knights Is, N.Z. 31.X. 65 – 1.XI. 65 K.G. Somerfield / NEW ZEALAND ND Poor Knights Is Tawhiti Rahi / Mecodema sp. Det. P.M. Johns Carabidae / PARATYPE Mecodema ponaiti design. by Seldon & Leschen 2010 (NZAC); 1, Poor Knights Is, N.Z. 1.XI. 65 K.G. Somerfield / PARATYPE Mecodema ponaiti design. by Seldon & Leschen 2010 (NZAC); 3, NEW ZEALAND ND Poor Knights Is Tawhiti Rahi 9 Sep 1980 J.C. Watt / under stones / PARATYPE Mecodema ponaiti design. by Seldon & Leschen 2010 (NZAC); 1, NEW ZEALAND ND Poor Knights Is Aorangi Urupa Point 11 Nov 1981 / J.C. Watt / PARATYPE Mecodema ponaiti design. by Seldon & Leschen 2010 (NZAC); 1, NEW ZEALAND ND Poor Knights Is Aorangi Urupa Point 11 Nov 1981 / J.C. Watt ex soil / PARATYPE Mecodema ponaiti design. by Seldon & Leschen 2010 (NZAC); 1, NEW ZEALAND ND Poor Knights Is Aorangi Puweto Valley 13 Nov 1981 / J.C. Watt under logs / PARATYPE Mecodema ponaiti design. by Seldon & Leschen 2010 (NZAC); 2, NEW ZEALAND ND Poor Knights Is Tawhiti Rahi Plateau 200m 3–10 Dec 1980 / J.C. Watt pit trap 80 / 178 / PARATYPE Mecodema ponaiti design. by Seldon & Leschen 2010 (NZAC); 1, NEW ZEALAND ND Poor Knights Is Aorangi Saddle 100m 16 Nov 1981 / J.S. Dugdale Litter 81 / 148 / Mecodema ponaiti n. sp. design. by Seldon & Leschen 2010; 1, NEW ZEALAND ND Poor Knights Is Tawhiti Rahi Summit Ridge 3 Dec 1984 / J.C. Watt under logs / PARATYPE Mecodema ponaiti design. by Seldon & Leschen 2010 (NZAC); 1, Tawhiti Rahi under rock 1 - 1-56 J.C. Watt / Mecodema sp. n. nr atrox Brit. Det. J.C. Watt 1957 / Collection J.C. Watt / J.I. Townsend Collection / PARATYPE Mecodema ponaiti design. by Seldon & Leschen 2010 (NZAC); 4, NEW ZEALAND ND Poor Knights Is. Aorangi Is. Jun 2006 D.S. Seldon & T. Buckley / PARATYPE Mecodema ponaiti design. by Seldon & Leschen 2010 (NZAC); 1, NEW ZEALAND ND Poor Knights Is. Tawhiti Rahi Is. 18 th Dec 2009, D.S. Seldon, T. Buckley & R.A.B. Leschen / PARATYPE Mecodema ponaiti design. by Seldon & Leschen 2010 (NZAC). Etymology. The Poor Knights Islands are within the Ngatiwai rohe and as Ngatiwai have no collective name for the Poor Knights Islands, they requested that the species be named ponaiti (pona-iti), which is a trans-literation of the name of the islands.Published as part of Seldon, David S. & Leschen, Richard A. B., 2011, Revision of the Mecodema curvidens species group (Coleoptera: Carabidae: Broscini), pp. 1-45 in Zootaxa 2829 on pages 36-38, DOI: 10.5281/zenodo.20285
Mecodema haunoho Seldon & Leschen, 2011, sp. n.
<i>Mecodema haunoho</i>, sp. n. <p>Figures: 5, 16d, 21b, 25a, 26.</p> <p> <b>Diagnosis.</b> Head broad and flattened. Clypeus with a single setose puncture on each side, bearing 2 setae. Prothoracic carina narrow, crenulated and extended to anterior angle. Pronotum fovea deep and broad. Elytral striae with regularly distributed, small asetose punctures. Setose punctures of 3rd, 4th and 5th ventrites absent. Apical portion of aedeagus symmetrically rounded with a moderate twist to right of vertical axis. Dorso-apical portion of central spicule, not greatly expanded. Basal lobe of left paramere rectangular, arm relatively large and apical lobe medium, and with setation along apical ¾ of ventral edge.</p> <p> <b>Description.</b> Length 20–26 mm. Colour black. Head broad and flattened. Microsculpture of frons may be present, absent on vertex. Vertexal groove narrow and defined entire length. Form of microsculpture on anterior area of clypeus, sulcate. Clypeus with a single setose puncture on each side bearing 2 setae. Form of lobe of the mentum, square. Submentum constriction narrow with 6 setae. Prothoracic carina narrow, crenulated and extended to anterior angle. Pronotum broad and flattened, posterior lateral sinuation slightly angled inward, pronotal fovea deep and broad, rugose microsculpture may be present on pronotal disc. Elytra broad but convex, striae with regularly distributed small asetose punctures, intervals slightly convex. Setose punctures of mesocoxae and metacoxae absent. Setose punctures of 3rd, 4th and 5th ventrites absent. Apical portion of aedeagus symmetrically rounded with a moderate twist to right of vertical axis. Dorso-apical portion of central spicule, not greatly expanded. Left paramere basal lobe rectangular, arm relatively large, apical lobe medium, and with setation along apical ¾ of ventral edge.</p> <p> <b>Comments.</b> <i>M. haunoho</i> is one of four species that form a monophyletic group (fig. 5), with its sister taxa <i>M. aoteanoho</i> (Great Barrier Island), <i>M. atrox</i> (Coromandel Ranges) and the widespread <i>M. occiputale</i>. This is the first described carabid species endemic to Little Barrier Island, the other 16 native carabid species on the island are also found on the mainland. Little is known of their ecology or biology; however, specimens have been mostly collected throughout the summer months in mature mixed kauri (<i>Agathis australis</i>) / broadleaf forest and usually in relatively close proximity (within 100 m) to freshwater. Additional specimens have been found in dry stream beds, and burrows of seabirds.</p> <p> <b>Distribution.</b> North Island: CL, Little Barrier Island (only).</p> <p> <b>Holotype:</b> Male labelled NEW ZEALAND CL, Little Barrier Island, Tirikakawa Stream, 2005, D.S. Seldon / HOLOTYPE Mecodema haunoho <b>n. sp.</b> design. by Seldon & Leschen 2010 [red label] (NZAC).</p> <p> <b>Paratype material examined:</b> 1, NEW ZEALAND AK Little Barrier I Summit Track 458 m on ground 10 Mar 1974 J.C. Watt / J.I. Townsend Collection / PARATYPE Mecodema haunoho <b>n. sp.</b> design. by Seldon & Leschen 2010 (NZAC); 1, NEW ZEALAND CL Little Barrier I Mt Hauturu Track 366m 26 Feb 1976 T.K. Crosby on ground / PARATYPE Mecodema haunoho <b>n. sp.</b> design. by Seldon & Leschen 2010 (NZAC); 1, NEW ZEA- LAND CL Little Barrier I 17 Dec 1983 J. Playfair / PARATYPE Mecodema haunoho <b>n. sp.</b> design. by Seldon & Leschen 2010 (NZAC); 1, NEW ZEALAND CL Little Barrier Is Waitoki Stm Track 7 1 Dec 1978 / R. Beever under rock / PARATYPE Mecodema haunoho <b>n. sp.</b> design. by Seldon & Leschen 2010 (NZAC); 9, NEW ZEA- LAND CL Little Barrier Is. Awaroa Stm 2007 D.S. Seldon / PARATYPE Mecodema haunoho <b>n. sp.</b> design. by Seldon & Leschen 2010 (NZAC); 1, NEW ZEALAND CL Little Barrier Is. Awaroa Stm 2006 D.S. Seldon & M. Rayner / PARATYPE Mecodema haunoho <b>n. sp.</b> design. by Seldon & Leschen 2010 (NZAC); 1, NEW ZEALAND CL Little Barrier Is. Valley Tk 2006 D.S. Seldon / PARATYPE Mecodema haunoho <b>n. sp.</b> design. by Seldon & Leschen 2010 (NZAC).</p> <p> <b>Etymology.</b> The name of this species was chosen from several names provided by the people of Ngati Manuhiri. The name <i>haunoho</i> is derived from hau, the site (Hauturu) and noho, which means ‘dwell, inhabit’.</p>Published as part of <i>Seldon, David S. & Leschen, Richard A. B., 2011, Revision of the Mecodema curvidens species group (Coleoptera: Carabidae: Broscini), pp. 1-45 in Zootaxa 2829</i> on pages 28-30, DOI: <a href="http://zenodo.org/record/202857">10.5281/zenodo.202857</a>
Mecodema aoteanoho Seldon & Leschen, 2011, sp. n.
<i>Mecodema aoteanoho</i>, sp. n. <p>Figures: 5, 14c, 25a, 26b.</p> <p> <b>Diagnosis.</b> Head broad but body convex. Clypeus with a single setose puncture on each side bearing 2 setae. Prothoracic carina narrow, crenulated and extended to beyond anterior angle. Pronotal fovea shallow and narrow. Elytra striae with regularly distributed, small asetose punctures. Setose punctures of 3rd, 4th and 5th ventrites absent. Apical portion of aedeagus symmetrically rounded with a moderate twist to right of vertical axis. Dorso-apical portion of central spicule, not greatly expanded. Basal lobe of left paramere rectangular, arm relatively large and apical lobe small, with setation along apical ½ of ventral edge.</p> <p> <b>Description.</b> Length 19–24 mm. Colour black. Head broad but body convex. Microsculpture of frons may be present, absent on vertex. Vertexal groove narrow, defined entire length. Form of microsculpture on anterior area of clypeus, sulcate. Clypeus with a single setose puncture on each side, bearing 2 setae. Form of lobes of mentum, square. Submentum constriction narrow with 6 setae. Prothoracic carina narrow, crenulated and extended to beyond anterior angle. Pronotum broad but moderately convex, posterior lateral sinuation angle slightly angled inward, pronotal fovea shallow and narrow, rugose microsculpture may be present laterally on pronotal disc. Elytra narrow and convex, striae with regularly distributed small asetose punctures, intervals slightly convex. Setose punctures of mesocoxae present, absent on metacoxae. Setose punctures of 3rd, 4th and 5th ventrites absent. Apical portion of aedeagus symmetrically rounded with a moderate twist to right of vertical axis. Dorso-apical portion of central spicule, not greatly expanded. Left paramere basal lobe rectangular, arm relatively large, apical lobe relatively small, with setation along apical ½ of ventral edge.</p> <p> <b>Comments.</b> The description of this species is based on 24 specimens that have been collected from several localities on Great Barrier Island. The holotype is male, with the genitalia contained in a silicon genitalic tube and pinned with specimen. Most specimens have been found in relatively undisturbed, mature native broadleaf forests that range from coastal to the highest points (600 m) of the island. <i>Mecodema aoteanoho</i> is the first described carabid species endemic to Great Barrier Island and is the sister species to <i>M. haunoho</i> (Little Barrier Island). The close relationship among these species is supported by the central spicule of the apical plate (fig. 26).</p> <p> <b>Distribution.</b> North Island: CL, Great Barrier Island (only).</p> <p> <b>Holotype:</b> Male labelled (NZAC) NEW ZEALAND CL Great Barrier I 23–28 Mar 1978 ESNZ Field Trip / Whangaparapara Track x under logs [hw] / HOLOTYPE Mecodema aoteanoho <b>n. sp.</b> design. Seldon & Leschen 2010 [red label].</p> <p> <b>Paratype material examined:</b> 1, Kaitoke, Great Barrier Is., on ground. 25-11-57 J.C. Watt. / Collection J.C. Watt / J.I. Collection / PARATYPE Mecodema aoteanoho <b>n. sp.</b> design. Seldon & Leschen 2010 (NZAC); 1, NEW ZEALAND CL Great Barrier I. Rosalie Bay Road, 15 Sep 1999, S.E. Thorpe / under stone in bush / AMNZ 29938 Auckland Museum New Zealand / PARATYPE Mecodema aoteanoho <b>n. sp.</b> design. Seldon & Leschen 2010; 1, NEW ZEALAND CL Great Barrier I Rosalie Bay Road 17 Sep 1999 S.E. Thorpe / under stone in bush / AMNZ 40770 Auckland Museum New Zealand / PARATYPE Mecodema aoteanoho <b>n. sp.</b> design. Seldon & Leschen 2010; 1, NEW ZEALAND Great Barrier I. Little Windy Hill, 260m, 17–21.ii 2002, J.W. Early / Forest. YPT L21104 / AMNZ 49087 Auckland Museum New Zealand / PARATYPE Mecodema aoteanoho <b>n. sp.</b> design. Seldon & Leschen 2010; 2, NEW ZEALAND CL Great Barrier I Little Windy Hill 180m 31 iii–30iv 2003 K. Parsons Pit trap. L12123 / AMNZ 54671 Auckland Museum New Zealand / PARATYPE Mecodema aoteanoho <b>n. sp.</b> design. Seldon & Leschen 2010; 1, NEW ZEALAND CL Great Barrier I Little Windy Hill viii 2002 K. Parsons L11750 / AMNZ 52228 Auckland Museum New Zealand / PARATYPE Mecodema aoteanoho <b>n. sp.</b> design. Seldon & Leschen 2010; 1, NEW ZEALAND CL Great Barrier I. Little Windy Hill xi 2002 K. Parsons / AMNZ 52943 Auckland Museum New Zealand / PARATYPE Mecodema aoteanoho <b>n. sp.</b> design. Seldon & Leschen 2010; 1, NEW ZEALAND CL Great Barrier I., Little Windy Hill, 18.vii.2002, K. Parsons. Under rock in forest. L11654 / AMNZ 51444 Auckland Museum New Zealand / PARATYPE Mecodema aoteanoho <b>n. sp.</b> design. Seldon & Leschen 2010; 1, NEW ZEALAND CL Great Barrier I. Little Windy Hill, 200m 20.xi–19.xii.2002 K. Parsons Pitfall trap. L11906 / AMNZ 52944 Auckland Museum New Zealand / PARATYPE Mecodema aoteanoho <b>n. sp.</b> design. Seldon & Leschen 2010; 1, NEW ZEALAND CL Great Barrier I. Little Windy Hill, 200m pitfall trap, 26.x–27.xi.2003 K. Parsons / AMNZ 56001 Auckland Museum New Zealand / PARATYPE Mecodema aoteanoho <b>n. sp.</b> design. Seldon & Leschen 2010; 1, NEW ZEALAND CL Great Barrier I. Little Windy Hill, 200m 8.vii– 25.viii.2003 K. Parsons Pitfall Trap. L12299 / AMNZ 55384 Auckland Museum New Zealand / PARATYPE Mecodema aoteanoho <b>n. sp.</b> design. Seldon & Leschen 2010; 3, NEW ZEALAND CL Great Barrier I. Little Windy Hill, 200m 18.ix–3.xi.2003 pitfall trap K. Parsons / AMNZ 55384, 55385 & 55376 Auckland Museum New Zealand / PARATYPE Mecodema aoteanoho <b>n. sp.</b> design. Seldon & Leschen 2010; 2, NEW ZEALAND CL Great Barrier I. Little Windy Hill, 200m pitfall trap, 27.xi–18.xii. 2003 K. Parsons / AMNZ 55999 & 56000 Auckland Museum New Zealand / PARATYPE Mecodema aoteanoho <b>n. sp.</b> design. Seldon & Leschen 2010; 2, NEW ZEA- LAND CL Great Barrier I Te Paparahi pitfall A 21 Jan 2002 A. Warren / AMNZ 61663 & 61769 Auckland Museum New Zealand / PARATYPE Mecodema aoteanoho <b>n. sp.</b> design. Seldon & Leschen 2010; 3, NEW ZEA- LAND CL Great Barrier I Te Paparahi pitfall A3 & A 25 Feb 2002 A. Warren / AMNZ 61657, 61770 & 63294 Auckland Museum New Zealand / PARATYPE Mecodema aoteanoho <b>n. sp.</b> design. Seldon & Leschen 2010; 2, NEW ZEALAND CL Great Barrier I Te Paparahi pitfall A3 & 23 Apr 2002 A. Warren / AMNZ 61584 & 61658 Auckland Museum New Zealand / PARATYPE Mecodema aoteanoho <b>n. sp.</b> design. Seldon & Leschen 2010. Additional material viewed from MONZ.</p> <p> <b>Etymology.</b> The name is based on its sister taxa <i>M. haunoho</i> and the close proximity of the two Barrier Islands. It is derived from ‘aotea’, the Māori name for Great Barrier Island and ‘noho’, which means ‘dwell, inhabit’.</p>Published as part of <i>Seldon, David S. & Leschen, Richard A. B., 2011, Revision of the Mecodema curvidens species group (Coleoptera: Carabidae: Broscini), pp. 1-45 in Zootaxa 2829</i> on pages 23-24, DOI: <a href="http://zenodo.org/record/202857">10.5281/zenodo.202857</a>
Mecodema parataiko Seldon & Leschen, 2011, sp. n.
<i>Mecodema parataiko</i>, sp. n. <p>Figures: 5, 6d, 7a, 13b, 18b, 25a.</p> <p> <b>Diagnosis.</b> Head and body broad, slightly convex. Clypeus with a single setose puncture on each side bearing 1 seta. Prothoracic carina broad, crenulated and extended to anterior angle. Pronotum fovea deep and broad. Elytral striae with large asetose punctures in a confused distribution. Setose punctures absent on 3rd, 4th and 5th ventrites. Apical portion of aedeagus symmetrically rounded with strong twist to right of vertical axis. Dorso-apical portion of central spicule, not greatly expanded. Basal lobe of left paramere rectangular, arm and apical lobe relatively large, and with setation along apical ¾ of ventral edge.</p> <p> <b>Description.</b> Length 20–22 mm. Colour black. Head broad and body slightly convex. Microsculpture of frons and vertex absent. Vertexal groove shallow laterally, narrow and defined middle. Form of microsculpture on anterior area of clypeus, poorly defined, clypeus with a single setose puncture bearing 1 seta. Form of lobes of the mentum, square. Submentum constriction broad, with 6 setae. Prothoracic carina broad, crenulated and extended to anterior angle. Pronotum broad and flat, posterior lateral sinuation angle parallel. Pronotal fovea deep and broad. Rugose microsculpture present on pronotal disc. Elytra narrow and highly convex, striae with large asetose punctures in a confused distribution. Setose punctures of mesocoxa and metacoxa absent. Setose punctures of 3rd, 4th and 5th ventrites absent. Apical portion of aedeagus symmetrically rounded with strong twist to right of vertical axis. Dorso-apical portion of central spicule, not greatly expanded. Left paramere basal lobe rectangular, arm and apical lobe relatively large, and with setation along apical ¾ of ventral edge.</p> <p> <b>Comments.</b> <i>Mecodema parataiko</i> and <i>M. pluto</i> are sister species, both are distinguishable from the other curvidens species by the strongly convex shape of the elytra and the confused distribution pattern of the asetose punctures along the elytra striae. The difference between the two species is that <i>M. parataiko</i> has larger and more strongly impressed asetose punctures distributed in a confused pattern along the elytral striae. These two species form a monophyletic group nested in the Northland basal elements of the group. <i>M. parataiko</i> is the most widespread of the species found in Northland.</p> <p> <b>Distribution.</b> New Zealand, North Island, Northland (ND), Parataiko Range, Warawara Forest, Tangihua Range, Omahuta F.P., Mangamuka S.R., Herekino Forest and Ahipara Plateau.</p> <p> <b>Holotype:</b> Male labelled (NZAC) NEW ZEALAND ND Mangamuka Gorge Wkwy 350m 351140S/1732725E 17.IX.–16.X.1999 Larivière. Larochelle / wet broadleaf forest pittraps / HOLOTYPE Mecodema parataiko <b>n. sp.</b> design. by Seldon & Leschen 2010 [red label].</p> <p> <b>Paratype material examined:</b> 3, NEW ZEALAND ND Mangataniwha Rd 400m 8–14 Dec 1976 B.M. May / PARATYPE Mecodema parataiko <b>n. sp.</b> design. by Seldon & Leschen 2010 (NZAC); 1, NEW ZEALAND ND Tangihua R Tangihua Lodge 6 Apr 1993, G. Hall / under stone / PARATYPE Mecodema parataiko <b>n. sp.</b> design. by Seldon & Leschen 2010 (NZAC); 2, NEW ZEALAND ND Omahuta For., Pukekohe Stm 27.XII.1992 A. Larochelle / Tree fern Mellcytus for. pit traps in daytime. / PARATYPE Mecodema parataiko <b>n. sp.</b> design. by Seldon & Leschen 2010 (NZAC); 1, NEW ZEALAND ND Omahuta For., Pukekohe Stm 27.XII.1992 A. Larochelle / Tree fern for. daily activ. rain / PARATYPE Mecodema parataiko <b>n. sp.</b> design. by Seldon & Leschen 2010 (NZAC); 1, NEW ZEALAND ND Mangamuka Walkway 500m 22.IX.1997 Larochelle. Larivière / wet broadleaf forest in burrows under stones. / PARATYPE Mecodema parataiko <b>n. sp.</b> design. by Seldon & Leschen 2010 (NZAC); 3, NEW ZEALAND ND Mangamuka Gorge Wkwy 350m 351140S/1732725E 17.IX.–16.X.1999 Larivière. Larochelle / wet broadleaf forest pittraps / PARATYPE Mecodema parataiko <b>n. sp.</b> design. by Seldon & Leschen 2010 (NZAC); 1, NEW ZEALAND ND Mangamuka Gorge Wlkwy 550m 351215S/1732652E 29.VI.–27.VII.1999 Larivière. Larochelle / wet broadleaf forest pittraps / PARATYPE Mecodema parataiko <b>n. sp.</b> design. by Seldon & Leschen 2010 (NZAC);1, NEW ZEALAND ND Mangamuka Gorge Wlkwy 350m 351140S/1732725E 27.V.– 29.VI.1999 Larivière. Larochelle / wet broadleaf forest pittraps / PARATYPE Mecodema parataiko <b>n. sp.</b> design. by Seldon & Leschen 2010 (NZAC); 1, NEW ZEALAND ND Mangamuka Gorge Wlkwy 625m 351228S/ 1732640E 27.VII.–14.VIII.1999 Larivière. Larochelle / PARATYPE Mecodema parataiko <b>n. sp.</b> design. by Seldon & Leschen 2010 (NZAC); 2, NEW ZEALAND ND Mangamuka Gorge Wlkwy 625m 351228S/1732640E 14.VIII.–17.IX.1999 Larivière. Larochelle / wet broadleaf forest pittraps / PARATYPE Mecodema parataiko <b>n. sp.</b> design. by Seldon & Leschen 2010 (NZAC); 1, Mangamuka Gorge R.A. Cumber Coll. 23/3/1959 / J.I. Townsend Collection / PARATYPE Mecodema parataiko <b>n. sp.</b> design. by Seldon & Leschen 2010 (NZAC); 2, Coll. E. Fairburn 26-3-1932 / Waimatenui Kaikohe N. Auckland. / A.E. Brookes Collection / J.I. Townsend Collection / PARA- TYPE Mecodema parataiko <b>n. sp.</b> design. by Seldon & Leschen 2010 (NZAC); 1, NEW ZAELAND ND Mangataniwha Ra 400m 8–14 Dec 1976 B.M. May / PARATYPE Mecodema parataiko <b>n. sp.</b> design. by Seldon & Leschen 2010; 1, N. of Donnelly’s Crossing N. Auck. 12-10-1936 C.E. Clarke / C. E. Clarke Collection / Mecodema sp.n. nr. atrox Brit. pro. ratio 1/1.26/1.29/1.68 width/length 1.08 Det. J.C. Watt 1959 / AMNZ 40842 Auckland Museum New Zealand / PARATYPE Mecodema parataiko <b>n. sp.</b> design. by Seldon & Leschen 2010; 1, Waimatenui 5-4-42 C.E. Clarke / C. E. Clarke Collection / AMNZ 26565 Auckland Museum New Zealand / PARATYPE Mecodema parataiko <b>n. sp.</b> design. by Seldon & Leschen 2010; 1, NEW ZEALAND ND. Waipoua FOREST, Yakas Tk S. end (approx. 35 39’ S 17333’E), under log. 26 Mar 2002 S.E. Thorpe / AMNZ 49742 Auckland Museum New Zealand / PARATYPE Mecodema parataiko <b>n. sp.</b> design. by Seldon & Leschen 2010; 1, Scrub on plateau N. side Hunahuna Stm Vy Ahipara Plateau 25.IV. 1984 K.A.J. Wise / AMNZ 40843 Auckland Museum New Zealand / PARATYPE Mecodema parataiko <b>n. sp.</b> design. by Seldon & Leschen 2010; 16, NEW ZEALAND ND Herekino Forest Herekino Saddle Tk 2010 D.S. Seldon & T. Buckley / PARATYPE Mecodema parataiko <b>n. sp.</b> design. by Seldon & Leschen 2010 (NZAC); 2, NEW ZEALAND ND Mangamuka Forest Mangamuka Ridge Tk 2010 D.S. Seldon & T. Buckley / PARATYPE Mecodema parataiko <b>n. sp.</b> design. by Seldon & Leschen 2010 (NZAC); 7, NEW ZEALAND ND Warawara Forest Hauturu Stm 2009 D.S. Seldon / PARATYPE Mecodema parataiko <b>n. sp.</b> design. by Seldon & Leschen 2010 (NZAC); 3, NEW ZEALAND, ND Trounson Kauri Park 21 viii 1996 G.L.F. Carlin pitfall traps in kauri forest / PARATYPE Mecodema parataiko <b>n. sp.</b> design. by Seldon & Leschen 2010 (LUNZ).</p> <p> <b>Etymology.</b> This species is named after the area (Parataiko Range) from which specimens were first collected.</p>Published as part of <i>Seldon, David S. & Leschen, Richard A. B., 2011, Revision of the Mecodema curvidens species group (Coleoptera: Carabidae: Broscini), pp. 1-45 in Zootaxa 2829</i> on pages 34-35, DOI: <a href="http://zenodo.org/record/202857">10.5281/zenodo.202857</a>
Mecodema tenaki Seldon & Leschen, 2011, sp. n.
<i>Mecodema tenaki</i>, sp. n. <p>Figures: 5, 10d, 18d, 20b, 25a.</p> <p> <b>Diagnosis.</b> Head and body broad. Clypeus with 2 setose punctures on each side, anterior with single seta, posterior with 2 setae. Prothoracic carina broad, heavily crenulated, and extended to anterior angle, pronotum fovea deep and narrow. Elytral striae with regularly distributed, small asetose punctures. Setose punctures absent on 3rd ventrite, present on 4th and 5th ventrites. Apical portion of aedeagus symmetrically rounded with a moderate twist to right of vertical. Dorso-apical portion of central spicule, not greatly expanded. Basal lobe of left paramere rectangular, arm and apical lobe relatively small, and with setation along apical ½ of ventral edge.</p> <p> <b>Description.</b> Length 25–27 mm. Colour black. Head broad and flattened. Microsculpture of frons and vertex absent. Vertexal groove narrow, defined entire length with V-shape at midpoint. Form of microsculpture on anterior area of clypeus, poorly defined. Clypeus with 2 setose punctures on each side, anterior with single seta, posterior with 2 setae. Form of lobes of the mentum, square. Submentum constriction broad, with greater than 6 setae. Prothoracic carina broad, heavily crenulated and extended to anterior angle. Pronotum broad and flattened, posterior lateral sinuation angled slightly inward, pronotal fovea deep and narrow. Elytra broad and flattened, striae with small regularly distributed asetose punctures, intervals slightly convex. One setose puncture present on mesocoxa and metacoxa. Setose punctures absent on 3rd ventrite, present on 4th and 5th ventrites. Apical portion of aedeagus symmetrically rounded with a moderate twist to right of vertical axis. Dorso-apical portion of central spicule, not greatly expanded. Left paramere basal lobe rectangular, arm and apical lobe relatively small, and with setation along apical ½ of ventral edge.</p> <p> <b>Comments.</b> The presence of the ‘V-shape’ at the midpoint of the vertexal groove and the shape of the pronotum indicates that <i>M. tenaki</i> is closely related to <i>M. regulus</i> (Three Kings Islands). However, the presence of setose punctures on the 4th and 5th ventrites, crenulation of the prothoracic carina and pronotal posterior sinuation, and the anterior extention beyond the anterior angle show <i>M. tenaki</i> also shares similarities with <i>M. manaia</i> (Whangarei area). Presently, no specimens of either species have been collected from any of the eastern coastal native forests between the two localities, Cape Reinga and Whangarei area.</p> <p> <b>Distribution.</b> New Zealand, Northland (ND), restricted to the Cape Reinga / North Cape area.</p> <p> <b>Holotype:</b> Male labelled (NZAC) NEW ZEALAND ND Cape Reinga Te Paki RBB-7 13 Nov 2006 O. Ball / HOLOTYPE Mecodema tenaki <b>n. sp.</b> design. by Seldon & Leschen 2010 [red label].</p> <p> <b>Paratype material examined:</b> ND: 1 (card mounted) Unuwhau Spirits Bay 11/1957 R.A.C. / Mecodema of REGULUS pronotal margin less defined near hind angles 2 setiferous punctures on penultimate ventrite / J.I. Townsend Collection / PARATYPE Mecodema tenaki design. by Seldon & Leschen 2010 (NZAC); 1, Rd to Tom Bowling Bay / Spirits Bay 17-1-1956 A.K. Walker / Mecodema of Regulus / pronotal margin less defined near hind angles 2 setiferous punctures on penultimate ventrite. / J.I. Townsend Collection / PARATYPE Mecodema tenaki design. by Seldon & Leschen 2010 (NZAC); 1, NEW ZEALAND ND Cape Reinga Te Paki Aug 2007 pit trap O. Ball / PARATYPE Mecodema tenaki design. by Seldon & Leschen 2010 (NZAC); 1, NEW ZEALAND ND Unuwhau 1986 / PARATYPE Mecodema tenaki design. by Seldon & Leschen 2010 (NZAC); 1, In leaf litter / Forest remnant Whareana North Cape 6.XII.1967 K.A.J. Wise / AMNZ 27690 Auckland Museum New Zealand; 1, Forest remnant Whareana North Cape 6.XII.1967 K.A.J. Wise / AMNZ 27689 Auckland Museum New Zealand / PARA- TYPE Mecodema tenaki design. by Seldon & Leschen 2010.</p> <p> <b>Etymology.</b> This species <i>tenaki</i> is named in honour of the first Māori tribe (Te Naki) to settle in the North Cape area. The name was provided by Saana Murray of Ngati Kuri.</p>Published as part of <i>Seldon, David S. & Leschen, Richard A. B., 2011, Revision of the Mecodema curvidens species group (Coleoptera: Carabidae: Broscini), pp. 1-45 in Zootaxa 2829</i> on page 40, DOI: <a href="http://zenodo.org/record/202857">10.5281/zenodo.202857</a>
Mecodema ponaiti Seldon & Leschen 2011
Mecodema ponaiti Seldon & Leschen, 2011. Figures 16B and 48. Diagnosis: Differing from other North Island Mecodema species by: 1, the prothoracic carina with 4–6 setae along each side (curvidens species group); 2, submentum sclerite with 2 setae widely-spaced each side; 3, shape of the basal lobe of the left paramere (Fig. 48 LP). Description: Length 20–26 mm, pronotal width 6.1–6.9 mm, elytral width 7.2–8.6 mm. Colour of the entire body glossy black, ventrally and legs may be dark reddish-brown. Head: Narrow and flat, deflected laterally (Fig. 14C). Vertex smooth; vertexal groove medially narrow and broad laterally; large supraorbital puncture bearing 3–4 setae; 2–3 well-defined supraorbital grooves; frons smooth (Fig. 48); frontoclypeal suture well-defined, tentorial pits large; anterior area of clypeus with indistinct and broad grooves, 1 setose puncture each side bearing 2 setae (or 2 setose punctures on each side bearing a single seta each). Labrum rounded, anterior edge slightly emarginated with 2 proximate central setae, 2 setae each side evenly spaced. Mentum lobes squared (Fig. 12G), median process short and narrow, slightly angled upward (15°), weakly indentate (Fig. 12C); mentum setae present. Submentum sclerite constriction broad with 2 setae widely-spaced medially, 2 setae widely-spaced each side. Stipes with 2 basal setae. Gula pits large, suture defined, gula flat and smooth. Gena (Fig. 8) with a fine isodiametric pattern across entire area. Prothorax: Prothoracic carina broad the entire length, crenulations absent with 4–6 setae each side (Fig. 48), extended to anterior angle (or extending to beyond anterior angle); posterior sinuation evidently carinate, parallel; pronotum narrow and moderately deflected laterally, overall shape rounded (Fig. 48); midline well-defined, anterior medial impression absent, posterior medial diamond-shaped impression present, disc without microsculpture; pronotal foveae shallow and narrow; anterior edge strongly inwardly curved, posterior edge straight. Prosternum flat with evenly spaced, fine transverse lines; proepisternum without microsculpture. Procoxal setae absent; protibia distally expanded and shovel-like (Fig. 10B). Elytra: Narrow and moderately delected laterally, overall shape ovate (Fig. 48), posterior ⅓ steeply angled to apex; humeral angle anteriorly convergent (Fig. 6A) (enhancing ovate shape); basal margin moderately curved and bevelled to base, interval 1 extended to scutellum; lateral carina narrow the entire length, extended to humeral angle; humerus with 2 widely-spaced setose punctures; suture impressed; all striae with small, regularly distributed, asetose punctures (in some specimens striae punctures may be partially obsolescent); all intervals weakly convex, interval 9 truncated and broadened; interval microsculpture present as fine lines; 7 th strial setal pattern with 1–2 setose punctures in anterior ½, 4–5 setose punctures in posterior ½ (Fig. 48), setose punctures large. Ventral surface: Mesepisternum and metepisternum (Fig. 2) without microsculpture; setose punctures present on mesocoxae (1) and metacoxae absent. All abdominal ventrites finely lineate laterally; ventrites 3–5 without setose punctures each side of midline; ventrite 6 setae present: ♂ with 3 very proximate setose punctures grouped each side at junction of straight apical edge and curve anterad, ♀ with 2 setose punctures each side widely spaced on bluntly pointed apical edge; some ventrites laterally foveate (indistinct). Anterior metaventrite process (Fig. 2) a short, pointed triangle with a broad carina the entire length. Male genitalia: Apical portion of penis lobe symmetrically rounded with a moderate deflection to right of vertical axis (VV) (Fig. 15B); shaft of penis lobe narrow and straight the entire length (VV) (Fig. 48 PL); overall length of penis lobe slightly curved ventrally (LV) (Fig. 16A). Structures of the endophallus (Fig. 4): lateral form of the apex of the central spicule rounded; dorsal form of the apex of the central spicule short and narrow; setal (scales) coverage of the apical plate moderate (26–75%); size of left setose flange small; size of right setose flange, large. Left paramere narrowly rectangular without dorsal hump (Fig. 48 LP), moderate slope (45°) to arm; arm long and broad, gradually narrowed to terminus; terminal lobe indistinct from arm, apical tuft of sparse long setae, sparsely distributed, short-lengthed setae extended along apical ½ of ventral edge (Fig. 48 LP); ventral edge straight. Right paramere narrowly triangular, gradually narrowed to terminus with a double row of mediumlengthed setae extended along apical ¾ along of ventral edge (Fig. 48 RP). Female genitalia: Basal gonocoxite 1 long and narrow, apically broadened, ventral surface with a few rugose wrinkles, internal dorso-lateral carina with 2 setae. Gonocoxite 2 (Fig. 5) ovate. Ramus long and broad. Comments: Mecodema ponaiti is endemic to the Poor Knights Islands and is more closely related to M. regulus from the Three Kings Islands, than the nearest mainland species (Seldon & Leschen 2011). Distribution: New Zealand, North Island, Northland, Poor Knights Islands (only). Holotype: NZAC male labelled. NEW ZEALAND ND Poor Knights Is. Tawhiti Rahi Is. 18 Dec 2009 D.S. Seldon, R.A.B. Leschen & T. Buckley / HOLOTYPE Mecodema ponaiti n. sp. design. by Seldon & Leschen 2010 [red label]. Paratypes: (see Seldon & Leschen 2011). Material examined: 1♂, NEW ZEALAND, ND, Tawhiti Rahi, Poor Knights Isl., 10 Dec 2013, T.R. Buckley, S. Myers, J. Stavert, TB781, ex rotting log, 35 26.990 S, 174 44.154 E, 145 m / CAR127 (NZAC); 1♀, NEW ZEALAND, ND, Tawhiti Rahi, Poor Knights Isl., 31 Oct 2004 F. Brook (NZAC).Published as part of Seldon, David S. & Buckley, Thomas R., 2019, The genus Mecodema Blanchard 1853 (Coleoptera: Carabidae: Broscini) from the North Island, New Zealand, pp. 1-148 in Zootaxa 4598 (1) on pages 95-97, DOI: 10.11646/zootaxa.4598.1.1, http://zenodo.org/record/266806
Mecodema manaia Seldon & Leschen, 2011, sp. n.
<i>Mecodema manaia</i>, sp. n. <p>Figures: 5, 6a, 12a, 14b, 16c, 18a, 25a.</p> <p> <b>Diagnosis.</b> Head broad and flattened. Clypeus with a single setose puncture on each side bearing 2 setae. Prothoracic carina broad, crenulated and extended to anterior angle, pronotum fovea deep and broad. Elytral striae with regularly distributed, small asetose punctures. Setose punctures on 3rd ventrite absent, present 4th and 5th ventrites. Apical portion of aedeagus symmetrically rounded with a strong twist to right of vertical axis. Dorso-apical portion of central spicule, not greatly expanded. Basal lobe of left paramere ovate, arm and apical lobe medium, and with setation along apical ¾ of ventral edge.</p> <p> <b>Description.</b> Length 26–29 mm. Colour black. Head broad and flattened. Microsculpture of frons and vertex absent. Vertexal groove narrow, well defined for entire length and is angled posteriorly at midpoint. Form of microsculpture on anterior area of clypeus, spathate. Clypeus with a single setose puncture on each side bearing 2 setae. Form of lobes of the mentum, rounded. Submentum constriction broad, with less than or equal to 6 setae.</p> <p>Prothoracic carina is broad, crenulated and extended to anterior angle. Pronotum broad and flattened, posterior lateral sinuation parallel (or square), pronotal fovea deep and broad. Elytra broad and flat, striae with regularly distributed, small asetose punctures, intervals slightly convex. A single setose puncture present on mesocoxa, setose punctures of metacoxa absent. Setose punctures of 3rd ventrite absent, present on 4th and 5th ventrites. Apical portion of aedeagus symmetrically rounded with a strong twist to right of vertical axis. Dorso-apical portion of central spicule, not greatly expanded. Left paramere basal lobe ovate, arm and apical lobe medium, and with setation along apical ¾ of ventral edge.</p> <p> <b>Comments.</b> This mainland species (<i>M. manaia</i>) is the sister species to all remaining species in the curvidens group with the exception of <i>M. ponaiti</i> (Poor Knights Islands). <i>Mecodema manaia</i> shares close morphological characters with <i>M. tenaki</i> (Cape Reinga) and <i>M. regulus</i> (Three Kings Islands) and although there is suitable forest habitat along the eastern coastline to the north, no specimens have been collected from these areas.</p> <p> <b>Distribution.</b> North Island: Northland (ND), Whangarei area.</p> <p> <b>Holotype:</b> Male labelled NEW ZEALAND ND Whangarei Heads Mt Lion 2005 F. Brook / HOLOTYPE Mecodema manaia <b>n. sp.</b> design. by Seldon & Leschen 2010 [red label] (NZAC).</p> <p> <b>Paratype material examined:</b> ND: 3 (card mounted) Whangarei Heads 22.9.1945 E. Fairburn [hw on underside of card] / Mecodema atrox mihi E.B. Britton. det.1960 / E. S. Gourlay Acc. 1970 Ent. Div. / J.I. Townsend Collection / NZ Arthropod Collection NZAC 04009616 / PARATYPE Mecodema manaia <b>n. sp.</b> design. by Seldon & Leschen 2010 (NZAC); 1, Coll. E. Fairburn 9-11-1929 / Kara Forest Whangarei N. Auckland / A.E. Brookes Collection / J.I. Townsend / PARATYPE Mecodema manaia <b>n. sp.</b> design. by Seldon & Leschen 2010 (NZAC); 3, Coll. E. Fairburn 17-12-1932 / Whangarei Heads N. Auckland / n. sp / A.E. Brookes Collection / J.I. Townsend Collection (NZAC); 4, Coll. E. Fairburn 22.9.1945. / Whangarei Heads N. Auckland / A.E. Brookes Collection / J.I. Townsend Collection / PARATYPE Mecodema manaia <b>n. sp.</b> design. by Seldon & Leschen 2010 (NZAC); 1, Illustrated D.E. Helmore / NEW ZEALAND ND Bream Head Whangarei 9 April 1999 R. Parrish / under rock in broadleaf forest / AMNZ 82681 Auckland Museum New Zealand / PARATYPE Mecodema manaia <b>n. sp.</b> design. by Seldon & Leschen 2010.</p> <p> <b>Etymology.</b> The species’ name, <i>manaia</i>, was provided by the people of Ngatiwai and is named after the type locality, Mt Manaia, Whangarei.</p>Published as part of <i>Seldon, David S. & Leschen, Richard A. B., 2011, Revision of the Mecodema curvidens species group (Coleoptera: Carabidae: Broscini), pp. 1-45 in Zootaxa 2829</i> on pages 30-31, DOI: <a href="http://zenodo.org/record/202857">10.5281/zenodo.202857</a>
Mecodema haunoho Seldon & Leschen 2011
Mecodema haunoho Seldon & Leschen, 2011. Figures 4, 15B and 30. Diagnosis: Distinguished from other North Island Mecodema species by having: 1, the prothoracic carina narrow the entire length, moderately crenulated with 3 setae along each side (curviden s species group); 2, distinctive shape of the penis lobe (Fig. 30 PL). Description: Length 20.5–24.1 mm, pronotal width 5.1–5.9 mm, elytral width 6.3–6.8 mm. Colour of entire body matte to glossy black. Head: Narrow and flat (Fig. 14C). Vertex with very fine wrinkles forming an isodiametric pattern (may be indistinct); vertexal groove narrow and well-defined the entire length; small supraorbital puncture bearing 2–3 setae; 1–2 well-defined supraorbital grooves (especially the one that encompasses the supraorbital puncture); frons with a large and shallow depression each side of midline (Fig. 30); frontoclypeal suture well-defined, tentorial pits absent; anterior area of clypeus (Fig. 9) with shallow and narrow grooves, very faint in some specimens; clypeus with 1 setose puncture each side bearing 2 setae. Labrum rectangular, anterior edge outwardly curved with 2 proximate central setae, 2 setae each side evenly spaced. Mentum lobes squared (Fig. 12G), median process short and narrow, slightly angled upward (15°), notched indentation (Fig. 12D); mentum setae variably present or absent. Submentum sclerite constriction narrow with 6–8 setae, medial setae pair distantly spaced, 2 lateral setae pairs anterad the others. Stipes with 2 basal setae. Gula pits (Fig. 8) large, suture poorly defined, gula flat and smooth. Gena with fine wrinkles forming an isodiametric pattern across entire area. Prothorax: Prothoracic carina narrow the entire length, moderately crenulated with 3–4 setae along each side (Fig. 30), extended beyond anterior angle; posterior lateral sinuation evidently carinate, slightly angled inward; pronotum broad and laterally deflected, overall shape squared; midline weakly impressed, medial impressions absent, disc with very fine wrinkles laterally; pronotal foveae shallow and narrow; anterior edge slightly inwardly curved, posterior edge slightly inwardly curved medially. Prosternum (Fig. 2) flat and smooth; proepisternum with indistinct wrinkles. Procoxal setae absent; protibia distally expanded and shovel-like (Fig. 10B). Elytra: Narrow and slightly deflected; humeral angle evenly convex (Fig. 6B), basal margin slightly curved and bevelled to base, interval 1 extended to margin; lateral carina narrow the entire length, extended to humeral angle; humerus with 2 setose punctures within angle (Fig. 30); suture impressed but no more so than striae; striae with small, regularly spaced asetose punctures; intervals 1–4 slightly convex, 5–8 moderately convex, interval 9 strongly convex; interval microsculpture absent; 7 th strial setal pattern with 0–1 setose puncture in anterior ½, 4 setose punctures in posterior ½ (Fig. 30), setose punctures small. Ventral surface: Mesepisternum and metepisternum with corrugose wrinkles; setose punctures of mesocoxae (Fig. 2) and metacoxae absent. All abdominal ventrites finely lineate; ventrites 3–5 setose punctures absent each side of midline; ventrite 6 setae present: ♂ with 2 setose punctures each side of straight portion of apical edge, ♀ with 2 setose punctures each side, distantly spaced at edge of bluntly rounded apical edge; lateral foveae absent. Anterior metaventrite process a short rounded triangle with a very broad (apically) and distinctly defined carina. Male genitalia: Apical portion of penis lobe symmetrically rounded with a moderate deflection to right of vertical axis (VV) (Fig. 15B), dorsal process slightly more flattened than ventral apical curve (Fig. 30 PL); overall length of penis lobe shaft with curve to right absent (VV); overall length of penis lobe shaft short and broad, moderately curved ventrally (LV) (Fig. 16B). Structures of the endophallus (Fig. 4): lateral form of the apex of the central spicule pointed; dorsal form of the apex of the central spicule short and narrow; setal (scales) coverage of the apical plate moderate (26–75%); size of left setose flange small; size of right setose flange large. Left paramere basal lobe a very slender rectangle (only slightly broader than right paramere), short slope to arm (Fig. 30 LP); arm relatively long and broad, gradually narrowed to terminal lobe; short and narrow terminal lobe with a tuft of medium length setae (Fig. 30 LP); sparse shorter setae extend along apical ¾ of ventral edge; ventral edge straight. Right paramere narrow and triangular with setae extending along apical ¾ of ventral edge (Fig. 30 RP). Female genitalia: Basal gonocoxite 1 short and broad, apically flattened with a few grooves present on ventral surface, internal dorso-lateral carina with 1 seta. Gonocoxite 2 (Fig. 5) a short, broadly rounded triangle, apical edge strongly reflexed. Ramus long and relatively broad. Comments: Mecodema haunoho is endemic to Te Hauturu-o-Toi/Little Barrier Island, and its very close sister taxa, M. aoteanoho is endemic to Aotea. Distribution: New Zealand, North Island, Coromandel, Te Hauturu-o-Toi/Little Barrier Island (only). Holotype: NZAC, male labelled: NEW ZEALAND CL, Little Barrier Island, Tirikakawa Stream, 2005, D.S. Seldon ♂ / HOLOTYPE Mecodema haunoho n. sp. design. by Seldon & Leschen 2010 [red label]. Paratypes: (see Seldon & Leschen 2011). Material examined: 1♀, NEW ZEALAND CL, Little Barrier Is., Awaroa Stream PTs 22.11.06 – 22.1.07, D.S. Seldon / NZAC04127983.Published as part of Seldon, David S. & Buckley, Thomas R., 2019, The genus Mecodema Blanchard 1853 (Coleoptera: Carabidae: Broscini) from the North Island, New Zealand, pp. 1-148 in Zootaxa 4598 (1) on pages 53-55, DOI: 10.11646/zootaxa.4598.1.1, http://zenodo.org/record/266806
Mecodema manaia Seldon & Leschen 2011
Mecodema manaia Seldon & Leschen, 2011. Figures 12A, 15B, 16C and 37. Diagnosis: Distinguished from other North Island Mecodema species by having: 1, the pronotal carina with 3–4 setae along each side (curvidens species group); 2, pronotum overall shape squared, midline poorly defined, medial impressions and microsculpture absent; 3, elytral interval 7 curved inwardly to parallel humeral angle; 4, ventrites 3 and 4 without setose punctures, ventrite 5 with 1 setose puncture each side of midline. Description: Length 24.5–28.7 mm, pronotal width 6.5–7.7 mm, elytral width 7.8–9.2 mm. Colour of entire body matte to glossy black. Head: Broad and convex (Fig. 14B). Vertex smooth; vertexal groove narrow, well-defined the entire length, curved posterad at midpoint; small supraorbital puncture bearing 3 setae; 3 moderately impressed supraorbital grooves; frons with a large and shallow depression each side of midline; frontoclypeal suture (Fig. 9) well-defined, tentorial pits large; anterior area of clypeus with broad and shallow grooves, 1 setose puncture each side bearing 2 setae (Fig. 37). Labrum rounded, anterior edge outwardly curved with 2 proximate central setae, 2 setae each side evenly spaced. Mentum lobes rounded (Fig. 12F), median process narrow and short, slightly angled upward (15°), moderately indentate (Fig. 12B); mentum setae present. Submentum sclerite constriction narrow with 4–6 evenly spaced setae. Stipes with 2 basal setae. Gula pits large, suture indistinct, gula flat with transverse lines present. Gena (Fig. 8) with very fine lines forming an isodiametric pattern across entire area. Prothorax: Prothoracic carina narrow the entire length, slightly crenulated with 3–4 setae each side (Fig. 37), extended to anterior angle; posterior lateral sinuation indistinctly carinate, parallel; pronotum broad and laterally deflected, overall shape squared; midline poorly defined, medial impressions and microsculpture absent; pronotal foveae deep and narrow; anterior edge (Fig. 11) inwardly curved, posterior edge curved, emarginated medially. Prosternum flat with fine transverse lines; proepisternum without microsculpture. Procoxal setae absent; protibia distally expanded and shovel-like (Fig. 10B). Elytra: Broad and moderately deflected; humeral angle evenly convex (Fig. 6B); basal margin moderately curved and bevelled to base, interval 1 extended to margin; lateral carina narrow the entire length, extended to humeral angle; humerus with 2–3 setose punctures (Fig. 37); suture poorly impressed; striae with small, regularly distributed, asetose punctures that slightly increased in size laterally; intervals 1–4 weakly convex, intervals 5–9 moderately convex, interval 7 curved inwardly to parallel humeral angle; interval microsculpture absent; 7 th strial setal pattern with 2 setose punctures in anterior ½, 5 setose punctures in posterior ½ (Fig. 37), setose punctures large. Ventral surface: Mesepisternum and metepisternum (Fig. 2) without microsculpture; setose punctures present on mesocoxa (1) and metacoxa (1). All abdominal ventrites finely lineate laterally, ventrites 3 and 4 without setose punctures, ventrite 5 (Fig. 2) with 1 setose puncture each side of midline; ventrite 6 setae present: ♂ with 2 evenly spaced setose punctures each side, most lateral pair at junction of straight apical edge and curve anterad, ♀ with 2–3 setose punctures proximate, but distantly separated on bluntly rounded apical edge; lateral foveae absent. Anterior metaventrite process (Fig. 2) a bluntly rounded triangle with a well-defined, narrow carina the entire length. Male genitalia: Apical portion of penis lobe symmetrically rounded (Fig. 37 PL) with a distinct deflection to right of vertical axis (VV) (Fig. 15A); shaft of penis lobe straight and relatively broad entire (VV), overall lenght of penis lobe without curve ventrally (LV). Structures of the endophallus (Fig. 4): lateral form of the apex of the central spicule rounded; dorsal form of the apex of the central spicule short and narrow; relative extent of setae (scales) of the apical plate sparse (1–25%); size of left setose flange small; size of right setose flange large. Left paramere basal lobe narrowly rectangular, short slope to arm; arm long and relatively broad, gradually narrowed to terminal lobe; terminal lobe indistinct from arm, apical tuft of long setae; medium-lengthed setae continued along apical ¾ of ventral edge (Fig. 37 LP); ventral edge slightly curved. Right paramere narrowly rectangular, gradually narrowed apically with medium to long setae extended along apical ¾ ventral edge (Fig. 37 RP). Female genitalia: Basal gonocoxite 1 (Fig. 5) long and broad, flattened apically, internal dorso-lateral carina with 2 relatively long setae, ventral surface smooth. Gonocoxite 2 bluntly rounded and triangular. Ramus short and narrow. Comments: Mecodema manaia shares similar morphological characters with M. tenaki (Cape Reinga) and M. regulus (Three Kings Islands), and although there is suitable forest habitat along the eastern coastline to the north, no specimens have been collected from these areas. Distribution: New Zealand, North Island, Northland, Whangarei Head’s area. Holotype: NZAC male labelled. NEW ZEALAND ND Whangarei Heads Mt Lion 2005 F. Brook / HOLOTYPE Mecodema manaia n. sp. design. by Seldon & Leschen 2010 [red label]. Paratypes: (see Seldon & Leschen 2011).Published as part of Seldon, David S. & Buckley, Thomas R., 2019, The genus Mecodema Blanchard 1853 (Coleoptera: Carabidae: Broscini) from the North Island, New Zealand, pp. 1-148 in Zootaxa 4598 (1) on pages 69-71, DOI: 10.11646/zootaxa.4598.1.1, http://zenodo.org/record/266806
Mecodema parataiko Seldon & Leschen 2011
<i>Mecodema parataiko</i> Seldon & Leschen, 2011. <p>Figures 6A, 12D and 45.</p> <p> <b>Diagnosis:</b> Distinguishable from other North Island <i>Mecodema</i> species by having: <b>1</b>, 4–6 setae along the prothoracic carina (<i>curvidens</i> species group); <b>2</b>, elytral humerus without setose punctures; <b>3</b>, all elytral striae with large, irregularly spaced asetose punctures (Fig. 45).</p> <p> <b>Description:</b> Length 19.3–22 mm, pronotal width 5–5.7 mm, elytral width 5.8–6.9 mm. Colour of entire body matte to glossy black, coxae may be dark red.</p> <p> <i>Head:</i> Narrow and flat, deflected laterally (Fig. 14C). Vertex with a few rugose wrinkles laterally; vertexal groove narrow and well-defined medially, shallowly impressed laterally; large supraorbital puncture bearing 3–4 setae; 2–3 well-defined supraorbital grooves; frons (Fig. 9) convex medially with a small and shallow depression each side, shallow grooves may be present laterally; frontoclypeal suture defined, tentorial pits small; anterior area of clypeus with indistinct broad grooves, 1 deep setose puncture each side bearing 2 setae (Fig. 45). Labrum rounded, anterior edge straight with 2 proximate central setae, 2 setae each side evenly spaced. Mentum lobes squared (Fig. 12G), median process long and narrow, upward angle absent, notched indentation (Fig. 12D); mentum setae present. Submentum sclerite constriction broad with 3 setae clustered laterally (very wide gap medially). Stipes with 2 basal setae. Gula pits small, suture poorly defined, gula flat and smooth. Gena (Fig. 8) with rugose wrinkles forming an isodiametric pattern laterally.</p> <p> <i>Prothorax:</i> Prothoracic carina broad the entire length, moderately crenulated with 4–6 setae each side (Fig. 45), extended to anterior angle; posterior lateral sinuation evidently carinate, inwardly angled; pronotum broad and moderately deflected laterally, overall shape squared (Fig. 45); midline impressed, anterior medial and posterior medial impressions absent, disc with transverse wrinkles laterally (Fig. 45); pronotal foveae deep and narrow; anterior edge distinctly inwardly curved, posterior edge straight. Prosternum flat and smooth; proepisternum with corrugose ridges. Procoxal setae absent; protibia distally expanded and shovel-like (Fig. 10B).</p> <p> <i>Elytra:</i> Narrow and moderately deflected; humeral angle anteriorly convergent (Fig. 6A); basal margin moderately curved and bevelled to base, interval 1 extended to scutellum; lateral carina narrow the entire length but broadened in posterior ⅓, extended to humeral angle; humerus without setose punctures (Fig. 45); suture welldefined; all striae with large, asetose punctures, irregularly spaced and of confused distribution; intervals 1–4 weakly convex, intervals 5–9 moderately convex; interval without microsculpture; 7 th strial setal pattern (Fig. 1) without setose punctures in anterior ½, 2–3 setose punctures in posterior ½, setose punctures large.</p> <p> <i>Ventral surface:</i> Mesepisternum and metepisternum with wrinkles; setose punctures absent on mesocoxa and metacoxa. All abdominal ventrites (Fig. 2) lineate; ventrites 3–5 without setose punctures each side of midline; ventrite 6 setae present: ♂ and ♀ with 1–3 setae each side of midline along a bluntly rounded apical edge; ventrites foveate laterally (very small and shallow). Anterior metaventrite process a short triangle with a broad carina the entire length.</p> <p> <i>Male genitalia:</i> Apical portion of penis lobe symmetrically rounded with a strong deflection to right of vertical axis (VV) (Fig. 15A), apex is a slightly flattened curve (Fig. 45 PL), shaft of penis lobe straight (VV), shaft of penis lobe narrowed apically but gradually broadened toward base, overall length of the penis lobe slightly curved ventrally (LV) (Fig. 16A). Structures of the endophallus (Fig. 4): lateral form of the apex of the central spicule pointed; dorsal form of the apex of the central spicule short and narrow; seta (scales) coverage of the apical plate extensive (76–100%); size of left setose flange small; size of right setose flange large. Left paramere basal lobe narrowly rectangular with a small dorsal hump, short slope to arm (Fig. 45 LP); arm short and broad, gradually narrowed to terminus; apical tuft of very long setae; setae extended along apical ½ (just past arm junction) of ventral edge; ventral edge moderately curved. Right paramere narrowly triangular, gradually narrowed to terminus with a double row of long setae extended along apical ¾ of ventral edge (Fig. 45 RP).</p> <p> <i>Female genitalia:</i> Basal gonocoxite 1 short and broad apically, ventral surface smooth, internal dorso-lateral carina with 1 seta near edge of gonocoxite 2. Gonocoxite 2 short and rounded triangle, slightly reflexed along apical edge. Ramus (Fig. 5) long and narrow.</p> <p> <b>Comments:</b> <i>Mecodema parataiko</i> is the most geographically widespread species of the <i>curvidens</i> species group found in Northland.</p> <p> <b>Distribution:</b> New Zealand, North Island, Northland, Parataiko Range, Warawara Forest, Tangihua Range, Omahuta-Puketi F.P., Mangamuka S.R., Herekino Forest and Ahipara Plateau.</p> <p> <b>Holotype:</b> NZAC male labelled. NEW ZEALAND ND Mangamuka Gorge Wkwy 350m 351140S/1732725E 17.IX.–16.X.1999 Larivière. Larochelle / wet broadleaf forest pit traps / HOLOTYPE Mecodema parataiko <b>n. sp.</b> design. Seldon & Leschen 2010 [red label].</p> <p> <b>Paratypes:</b> (see Seldon & Leschen 2011).</p> <p> <b>Material examined:</b> 1♀, NEW ZEALAND ND, Maungataniwha Range, Mangamuka River S.R., PTs 13–21 Jan 2010 D.S. Seldon, T.R. Buckley / Kauri-broadleaf forest, pitfall traps (NZAC); 1♂, 1, NEW ZEALAND ND, Herekino Forest Saddle Tk, PTs, 14–21 Jan 2010, D.S. Seldon, T.R. Buckley / 35°12.393, 173°11.487 223 m, CAR39B (NZAC).</p>Published as part of <i>Seldon, David S. & Buckley, Thomas R., 2019, The genus Mecodema Blanchard 1853 (Coleoptera: Carabidae: Broscini) from the North Island, New Zealand, pp. 1-148 in Zootaxa 4598 (1)</i> on pages 88-90, DOI: 10.11646/zootaxa.4598.1.1, <a href="http://zenodo.org/record/2668063">http://zenodo.org/record/2668063</a>
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