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Eutarsopolipus hebronae Seeman 2021, sp. nov.
No full textEutarsopolipus hebronaesp. nov. (Figs 25–26) Diagnosis. Adult female: Gnathosomal length 61–65, width 60–67, cheliceral stylets 75–84; tarsi II-III setae tc′′ 3–4. Larval female: Gnathosomal length 50–55, width 50–53, cheliceral stylets 64–66; tarsi II-III setae tc′′ 2–3. Material examined. Holotype female, ex Castelnaudia cordata. Mt Mee SF, 6 km NNW barracks, 27° 2ʹ 53ʺ S 152° 40ʹ 49ʺ E, 8 Jan-3 Mar 1992, D. Cook, flight intercept trap, IN5148, HR# T 68607 (QMS 113463). Paratypes. All ex C. cordata. 6 females, 3 larvae, as follows: all ex. C. cordata. 3 females, 1 larva, same data as holotype (QMS113464–66, ANIC 52-003964); 1 larva, McIntosh Rd, Unumbar SF, 28° 25ʹ 19ʺ S 152° 42ʹ 14ʺ E, 25 Nov 1972, G. Monteith, IN6467, HR# T 64329 (ANIC 52-003965); 1 female, Lower Coomera, 28° 12ʹ 20ʺ S 153° 10ʹ 46ʺ E, 9 Jan-6 Apr 1995, G. Monteith, pitfall trap, IN5998, HR# T 68610 (QMS 113467); 2 females, 1 larva, Lever’s Plateau, 28° 18ʹ 57ʺ S 152° 50ʹ 53ʺ E, 22 Feb-8 May 1976, G. & S. R. Monteith, pitfall trap, rainforest, IN5485, HR# T 68619 (QMS 113468–69, ZMH-A0015194). 1 female, 1 larva in ANIC; 1 female in ZMH. Other material examined. 4 females from type locality, mounted with specimens of E. umbonatus. Not included as paratypes. Notes: 1 female from each of Mt Mee and Lever’s Plateau are E. hebronae but too poor to be considered paratypes. There is also a pharate female from McIntosh Rd. Description. FEMALE (Figs 25–26; n = 6). Gnathosoma (Fig. 25B). Length 61 (61–65), width 60 (60–67). Cheliceral stylets 78 (75–84), setae ch 35 (29–38), su 17 (13–15). Idiosoma. Length 420 (345–430), width 290 (255–340). Prodorsal plate eroded mid-anteriorly; setae slender; setae v1 6 (5–7), v2 alveolar remnant, sc1 10 (6–10), sc2 95 (85–105). Distance between setae v1–v1 56 (54–66), sc1–sc1 105 (93–105), v1–sc1 33 (21–30), v2–v2 64 (67–69), sc2–sc2 140 (135–150), sc1–sc2 40 (33–43). Opisthosomal setae slender. Plate C setae c1 9 (7–9), c2 12 (11–13), distance between setae c1–c1 82 (69–85). Plate D setae d 10 (7–10), d–d 145 (130–150), cupuli ia well apart from d, anterolaterad d. Plate EF setae f 10 (9–13), f–f 86 (79–95), cupuli im well apart from f, anterolaterad f. Plate H divided or fused weakly, seta h minute. Venter: coxisterna 1 well defined, coxisterna 2 with some folds, coxisterna 3 weakly defined; coxal setae slender, 1a 3 (2–3), 2a 5 (4–5), 3a 7 (6–7), 3b 5 (5). Alveoli of 1b, 2b present. Legs (Figs 26 D-E). Setal counts legs I–III, femur-tarsus: 3-2-6(+ φ)-8(+ ω), 0-1-4-6, 0-1-4-6. Leg I. Femur I, d minute, l’ 3 (3), vʺ 8 (6–7); genu I, l′ 3 (3), l″ 3 (3–4); tibia I, d 50 (50–65), l’ 11 (11–13), lʺ 3 (3–4), v’ 8 (5–7), vʺ 12 (10–15), k 5 (5–7), φ 10 (10–12); tarsus I, tc’ 22 (19–20), tcʺ 18 (16–20), pl’ 15 (12–16), pl’’ 23 (20–23), pv′ 2 (2), pvʺ 2 (2), ω 5 (4–5), s 5 (5–6), u′′ 2 (m–2). Leg II. Genu II, l′ 4 (4–6); tibia II, d 45 (45–50), l’ 11 (9–12), v’ 19 (17–19), vʺ 21 (22–29); tarsus II, tc′ 4 (3–4), tc″ 4 (3–4), pl″ 52 (38–50), pvʺ 2 (2), u’ 6 (6–7), u′′ 2 (m–2). Leg III. Genu III, l′ 5 (4–5); tibia III, d 40 (35–50), l’ 8 (8–10), v’ 15 (14–18), vʺ 30 (26–30); tarsus III, tc’ 3 (2–3), tc″ 4 (3–4), pl″ 40 (40–50), pvʺ 2 (2), u’ 6 (6), u′′ m (m). LARVIFORM MALE. Unknown. LARVAL FEMALE (n = 3). Gnathosoma. Length 50–55, width 50–53. Cheliceral stylets 64–66, setae ch 35–36, su 8–10. Idiosoma. Length 210–270, width 165–210. Prodorsal plate with setae v1 6–7, v2 alveolar remnant, sc1 7–9, sc2 90–120. Distance between setae v1–v1 33–40, sc1–sc1 74–86, v1–sc1 29–34, v2–v2 51–60, sc2–sc2 80–90, sc1–sc2 25–30. Plate C setae c1 7–9, c2 10–13, distance between setae c1–c1 50–59. Plate D setae d 8–10, d–d 61–74, cupuli ia anterolaterad d. Plate EF setae f 9–11, f–f 43–52, cupuli im anteriad to anterolaterad f. Plate H long and narrow, 16–20 wide, ca. 30–40 long bearing setae h1 ca. 150, h2 8–10. Venter: coxisterna 1-2 moderately well defined, with few folds, coxisterna 3 poorly defined, membranous; coxal setae slender, 1a 3, 2a 5, 3a 5, 3b 5–6. Alveoli of 1b, 2b present. Legs. Setal counts legs I–III, femur-tarsus: 3-2-6(+ φ)-8(+ ω), 0-1-4-6, 0-1-4-6. Leg I. Femur I, d minute, l′ minute, v″ 5–7; genu I, l′ 3, l″ 2–3; tibia I, d 75–80, l’ 3, l’’ 2–3, v’ 6–8, v’’ 9–12, k 3–5, φ 11–12; tarsus I, tc’ 15–16, tc’’ 16, pl’ 9–13, pl’’ 18–20, pv′ m–2, pv’’ 2, ω 3–4, s 5–6, u′′ minute. Leg II. Genu II, l′ 3–4; tibia II, d 45–50, l’ 9–10, v’ 13–18, v’’ 23–26; tarsus II, tc′ 2–3, tc ″ 2–3, pl ″ 45–50, pv’’ 2, u’ 5–6, u′′ minute. Leg III. Genu III, l′ 3–4; tibia III, d 40–42, l’ 9–13, v’ 12–15, v’’ 20–26; tarsus III, tc’ 2, tc ″ 2–3, pl ″ 40–50, pv’’ 2, u’ 5–6, u′′ m–2. Differential diagnosis. Eutarsopolipus hebronae sp. nov. differs from E. despoticus sp. nov. by having a much larger gnathosoma (61–65 × 60–67 versus 45–51 × 47–51) and cheliceral stylets (75–84 versus 50–55). The species also differ in the size of setae tc″ on tarsi II-III, being much larger in E. hebronae (3–4) than E. despoticus (2–3). Even though these setae are only about 1 μm longer, they are obviously larger, being around twice as massive in E. hebronae. These differences in gnathosomal size and tarsal setae are present in the adult and larval female. Etymology. It is with great pleasure that I name this species for Wendy Hebron, with whom I have shared the role of collection manager for 14 years. Not only has her welcome and proficient assistance in the Queensland Museum arachnology collection been greatly appreciated, but her fine efforts have allowed me to dedicate more time to acarology. Remarks. This species was found co-infesting its host species with E. umbonatus sp. nov. and an undescribed species of Eutarsopolipus close to E. echinatus Seeman, 2019a (Tables 1, 2). The ochoai species group - basiatus subgroup Diagnosis. Female with dorsal idiosomal setae fine, slender; setae h minute or alveolar vestiges. All life stages with gnathosomal apex with small paired lobes. The ochoai species group - basiatus subgroup Diagnosis. Female with dorsal idiosomal setae fine, slender; setae h minute or alveolar vestiges. All life stages with gnathosomal apex with small paired lobes.Published as part of Seeman, Owen D., 2021, Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae), pp. 1-74 in Zootaxa 4971 (1) on pages 39-41, DOI: 10.11646/zootaxa.4971.1.1, http://zenodo.org/record/477151
Tenuipalpus kermanicus Safdarkhan & Asad & Seeman 2018, sp. nov.
No full textTenuipalpus kermanicus Khadem, Asadi & Seeman sp. nov. (Figs. 26–33) Diagnosis. Female: bOdy Ovate, withOut hysterOsOmal cOnstrictiOn; prOjectiOn On anteriOr margin Of prOdOrsum with twO well-develOped, pOinted lObes. Setae c1, d1, e1 present (f2 absent); dOrsum mOstly rugOse (cOmprising elOngate cells) except fOr medial prOpOdOsOmal shield mOstly smOOth; hysterOsOma with lateral ridge capturing hysterOsOmal pOres and setae e3, f3. DOrsal setae shOrt tO medium-length (10–27) and lanceOlate tO OblanceOlate, except filifOrm h2. One pair Of setae 3a, twO pairs Of setae 4a. Ventral cuticle between setae 1a–g1 with transverse striae; single ventrO-genital plate apparent. Palp three-segmented; palp femOrOgenu with lOng barbed seta; tarsus with One terminal sOlenidiOn. Leg chaetOtaxy: femOra 4-4-2-1; genua 3-3-1-0; tibiae 5-5-3-3; tarsi 9(1)-9(1)-5-5; seta d On femOra in dOrsal pOsitiOn. Material examined. HOlOtype female (SBUK) and five female paratypes (4 SBUC, 1 ACASΙ) ex Tamarix aphylla (Tamaricaceae), Ιran: Kerman PrOvince, Kuhbanan, 31°28’ N 56°01’ E, 6 May 2014, cOll. H. Khadem Safdarkhani. Description. FEMALE. ΙdiOsOma Ovate, withOut hysterOsOmal cOnstrictiOn. GnathOsOma: rOstrum just passing base Of femur Ι, setae m 7–9, barbed. Palpus three-segmented, femOrOgenu with a lOng barbed dOrsal seta, tibiOtarsus with a terminal sOlenidiOn (Fig. 28). Dorsum (Fig. 26) AnteriOr margin Of prOdOrsum with twO pOinted lObes. PrOdOrsum with a few irregular, elOngate cells anteriOrly and laterally, and a mOstly smOOth area medially; hysterOsOma with transverse elOngate cells mediOdOrsally, lOngitudinal cells laterally; a lateral ridge arOund hysterOsOma captures hysterOsOmal pOres and setae e3, f3. DOrsal setae ObOvate tO OblanceOlate, setae v2, sc1, c1, d1, e1 larger than Others. Lengths Of dOrsal setae: v2 26 (24–27), sc1 25 (23–25), sc2 12 (11–12), c1 25 (24–27), c3 13 (12–13), d1 18 (17–20), d3 13 (12–14), e1 18 (16–19), e3 10 (10–11), f3 15 (14–16), h1 12 (12–13), h2 105 (101–108); distances between dOrsal setae: v2 – h1 173 (169–178), v2–v2 18 (17–18), sc1–sc1 76 (74–77), sc2–sc2 110 (107–114), c1–c1 45 (44–47), c3–c3 119 (117–120), d1–d1 27 (26–28), d3–d3 103 (100–105), e1– e1 19 (18–21), e3–e3 74 (71–75), f3–f3 44 (42–45), h1–h1 19 (18–20), h2–h2 27 (25–28). Venter (Fig. 27). Transverse striae between setae 1a and anal plate; anteriOr area tO setae 1a and lateral hysterOsOma with lOngitudinal striae; ventrO-genital plate defined by transverse striae amOng arched striae. One shOrt intercOxal setae 3a; twO lOng pairs Of setae 4a. Ventral setae smOOth and fine. Lengths Of ventral setae: 1a 88 (85–89), 1b 16 (15–16), 1c 19 (17–19), 2b 51 (50–53), 2c 21 (20–22), 3a 17 (15–18), 3b 14 (14–15), 4a 1 67 (65–68), 4a 2 64 (61–65), 4b 16 (15–16), ag 24 (22–25), g1 16 (16–17), g2 14 (14–15), ps1 19 (18–20), ps2 17 (16–17). Spermatheca: NOt visible. Legs (Figs. 29–33). Tarsal claws pad-like and empOdium with tenent hairs. Setal fOrmula as fOllOws (cOxa tO tarsus; sOlenidia included in cOunt): Ι. 2–1–4–3–5–9(1ω), ΙΙ. 2–1–4–3–5–9(1ω), ΙΙΙ. 1–2–2–1–3–5, ΙV. 1–1–1–0–3– 5. DOrsal femOral setae On legs Ι–ΙΙ in dOrsal pOsitiOn; setae d On femur Ι–ΙΙ, genu Ι–ΙΙ and tibia Ι–ΙΙ Ovate tO OblanceOlate, serrate; setae lʹ On femur Ι–ΙΙ and trOchanter ΙΙΙ thickened, barbed. Male and immature stages. UnknOwn. Remarks. This new species belOngs tO the keiensis subgrOup Of the proteae grOup. Ιn additiOn tO the six species listed in Khanjani et al. (2013), the grOup alsO cOmprises: T. bassiae MOhanasundaram, 1988, T. clematidos Wang, 1983, T. ilocanus COrpuz-RarOs, 1978, T. isabelae Mesa et al., 2006, T. salicis Al-GbOOry, 1987 and T. tauricus (MitrOfanOv & StrunkOva, 1978). Tenuipalpus clematidos and T. salicis are tentatively included here, as they appear tO have setae f2 but lack setae d3; it is necessary tO nOte that setae d3 are sOmetimes OverlOOked, if they are small, situated marginally, and Obscured by leg ΙΙΙ. The brOad, ObOvate tO OblanceOlate dOrsal setae Of T. kermanicus sp. nov. distinguish it frOm mOst species in the proteae grOup. Ιn this regard, the new species is mOst similar tO T. berkheyae. These twO species are easily separated by numerOus features such as the three-segmented palps (twO-segmented in T. berkheyae), the lateral hysterOsOmal ridge (absent in T. berkheyae), dOrsal pOsitiOn Of seta d On femur Ι (lateral in T. berkheyae) and genual setal cOunt Of 3-3-1-0 (1-1-0-0 in T. berkheyae). Etymology. This species is named after the prOvince Of Kerman.Published as part of Safdarkhan, Khadem, Asad, Eh & Seeman, Owen D., 2018, Two new species of Tenuipalpus Donnadieu, 1875 (Acari: Trombidiformes: Tenuipalpidae) from Iran, pp. 511-524 in Zootaxa 4410 (3) on pages 519-522, DOI: 10.11646/zootaxa.4410.3.5, http://zenodo.org/record/122182
Jerry T. Seeman: Hall of Fame Inductee
No full textJerry T. Seeman: Hall of Fame Inductee
Jerry Seeman grew up and attended schools in Plainview and St. Charles, Minn. He was an outstanding quarterback on Winona State’s football teams in the 1950’s and also participated in basketball for three years. During his senior year he was assistant basketball coach under head coach Russ Owen.
He was active in several campus organizations including the Representative Council, W Club, Men’s PE Club, Science Club, intramural sports and Kappa Delta Ph honor society in education. Jerry was a class officer for three years and was named to Who\u27s Who Among American Colleges and Universities.
Jerry was a highly successful basketball coach in high schools at Whitehall, Wis., La Crosse, Wis. And Fridley, Minn. and compiled a .750 winning percentage with several championships in his eight years as a head basketball coach. His 1962 La Crosse Central team participated at the Wisconsin state tournament.
He served as a school administrator for 22 years and was Director of Finance and Personnel at Fridley Public Schools. Jerry was the recipient of a Shell Fellowship, Cornell University NSF Grant and received his M.S. degree from Western Washington State University.
Jerry has been known best for his sports officiating career having officiated contests at the high school, college and professional level for many years. He was a high school and collegiate basketball official for 22 years, and a referee for the National Football League since 1975. He was Director of Officials for the Minnesota State High School league for 14 years, a member of the National Federation of Basketball Rules Committee for four years and a NFL crew chief since 1979.https://openriver.winona.edu/halloffameinductees/1063/thumbnail.jp
Eutarsopolipus biuncatus Seeman 2021, sp. nov.
No full textEutarsopolipus biuncatussp. nov. (Figs 41–43) Diagnosis. Adult female: leg I with two claws; v1 8–10, sc2 15–20; genu I with two setae (l″ present); tarsus II setae tc″ 5–6, u′ 6–7. Adult male: setae sc2 2–3. Larval female: sc2 51–55. Material examined. Holotype female, ex. Notonomus transitus. Blue Mt, 0.6 km SE, 21° 36ʹ 19ʺ S 148° 58ʹ 21ʺ E, 3–4 Oct 1999, G. Monteith, D. Cook, C. Burwell, S. Evans, rainforest, IN7801, HR# T 89396 (QMS 113608). Paratypes. All ex. Notonomus transitus. 5 females, 5 males, 14 larvae, as follows: 2 females, 2 males, 10 larvae, same data as holotype (QMS 113609–18, ANIC 52-003975 – 77, ZMH-A0015201); 1 female, 1 male, 1 larva, 50 mi. W of Mackay, 21° 08ʹ 12ʺ S 148° 29ʹ 30ʺ E, 6 Jan 1973, J. Hammond, IN7864, HR# T 81753 (QMS 113621–23); 2 females, 2 males, 1 larva, Broken River, Eungella, 21° 10ʹ 5ʺ S 148° 30ʹ 30ʺ E, 4 Dec 1992 – 30 Apr 1993, P. Lawless, M. Shaw, R. Raven, IN 9781, HR# T110084 (QMS 113624–26, ZMH-A0015202–03); 2 larvae, Dalrymple Rd, Eungella NP, 21° 1ʹ 44ʺ S 148° 35ʹ 7ʺ E, 9 Nov 1991 – 29 Jul 1992, P. Lawless, R. Raven, M. Shaw, rainforest, IN7768, HR# T 81768 (QMS 113627–28). All in QM except 1 female, 1 male, 1 larva in ANIC and ZMH. Other material examined. 2 larvae, same data as holotype, in poor condition (QMS 113619–20). Description. FEMALE (Fig. 41; n = 6). Gnathosoma. Length 45 (44–46), width 39 (34–37). Palp with alveolar remnant of femoral seta. Cheliceral stylets 38 (35–36), setae ch 22 (14–16), su 11 (8–12). Idiosoma (Fig. 41A). Length ca. 500 (270–430), width ca. 300 (215–310), bigger specimens swollen, distorted. Stigmata apparent, large, in soft cuticle anteriad setae v1 to posterolaterad gnathosoma (varies according to mount), respiratory system weakly developed, tracheae not extending past small atrium, tracheae and atrium sometimes not visible. Prodorsal plate with setae v1 10 (8–9), v2 alveolar remnant, sc1 9 (7–8), sc2 20 (15–17). Distance between setae v1–v1 44 (39–51), sc1–sc1 69 (64–72), v1–sc1 22 (23–25), v2–v2 52 (53–57), sc2–sc2 84 (83–110), sc1–sc2 34 (24–32). Plate C appearing divided, each plate bearing its slender setae at their margins, c1 6 (5–7), c2 6 (4–5). Plate D setae d 5 (4–5), d–d 92 (85–95), cupuli ia anteriad d. Plate EF setae f 4 (4–5), f–f broken (49–60), cupuli im anterolaterad f. Plate H absent. Venter: coxisterna 1-3 well defined; coxal setae slender, 1a 5 (5–6), 2a 5 (4–5), 3a 6 (5–6), 3b 5 (4–5). Alveoli of 1b, 2b present. Legs (Figs 41 B-C). Setal counts legs I–III, femur-tarsus: 3-2-6(+ φ)-9(+ ω), 0-1-4-7(+ ω), 0-1-4-7. Leg I. Femur I, d 1 (1–3), l’ 5 (4–5), v″ 8 (9–10); genu I, l’ 3 (2–3), l″ 2 (2–3); tibia I, d 16 (14–19), l’ 3 (2–3), l’’ 5 (5–6), v’ 3 (3–5), v’’ 9 (8–14), k 3 (3–4), φ 6 (6–7); tarsus I, tc’ 9 (9–10), tc’’ 9 (9–10), pl’ 8 (8–10), pl’’ 10 (8–12), pv′ 2 (2–3), pv’’ 2 (2–3), ω 3 (3–4), s 6 (5–6), u′′ minute (m–2), p′ m (m–2). Leg II. Genu II, l’ 3 (2–3); tibia II, d 9 (7–9), l’ 5 (3–5), v’ 8 (8–11), v’’ 14 (13–16); tarsus II, tc′ 8 (6–8), tc ″ 5 (5–6), pl ″ 10 (7–9), pv′ 3 (3–4), pv’’ 4 (4–5), ω 3 (2–3), u’ 7 (6–7), u″ 2 (2). Leg III. Genu III, l’ 3 (2–3); tibia III, d 4 (5–6), l’ 4 (3–4), v’ 8 (6–9), v’’ 11 (13–15); tarsus III, tc’ 6 (6–7), tc ″ 5 (5–6), pl ″ 8 (9–10), pv′ 3 (2–3), pv’’ 2 (2–3), u’ 6 (6–7), u″ 2 (2). Claws on legs I-III well developed and paired. LARVIFORM ADULT MALE (Fig. 42; n = 5). Gnathosoma. Length 28–31, width 28–29. Palp with alveolar remnant of femoral seta. Cheliceral stylets 17–18, setae ch 1–2, su 4–6. Idiosoma. Length 155–165, width 110–120. Prodorsal plate with setae v 1 m–2, v2 alveolar remnant, sc1 minute, sc2 2–3, thickened. Distance between setae v1–v1 19–22, sc1–sc1 45–48, v1–sc1 19–21, v2–v2 34–36, sc2–sc2 54–56, sc1–sc2 22–23. Plate CD incised posterolaterally, setae c1, c2, d minute, distance between setae c1–c1 44–46, c1–c2 26–35, d–d 31–32; cupuli ia laterad to anterolaterad setae d. Plate EF with setae f minute, distance f–f 21–23; cupuli im anterolaterad setae f. Genital capsule length 22–25, width 34–36, setae h minute. Venter: coxal setae slender, 1a 2–3, 2a 2–3, 3a 2, 3b 2–3. Legs. Setal counts legs I–III, femur-tarsus: 2-0-6(+ φ)-8(+ ω), 0-0-4-7(+ ω), 0-0-4-7. Compared to female: Femur I seta v″ absent, genu I l′ absent, genua I-III l″ absent; tarsus I p′ absent. Leg I. Femur I, d minute, l’ 2; genu I, l′ minute; tibia I, d 4–5, l’ minute, l’’ m–2, v ’ 2, v ’’ 6–7, k minute, φ 4–5; tarsus I, tc’ 7–8, tc’’ 8–9, pl’ 4–6, pl’’ 6–9, pv ′ 2, pv’’ 2–3, ω 3, s 4–5, u ′′ 1–2. Leg II. Tibia II, d 3–5, l’ 2–3, v’ 6–7, v’’ 7–8; tarsus II, tc′ 3, tc ″ 4, pl ″ 5–6, pv ′ 2, pv’’ 2–3, ω 2–3, u’ 5–6, u″ minute. Leg III. Tibia III, d 2, l’ 2, v’ 6–8, vʺ 7–9; tarsus III, tc′ 2–3, tc ″ 4, pl ″ 4–5, pv ′ 2, pv’’ 2, u ’ 5, u″ minute. Claws on leg II-III present. Setal form similar to that in female, but with tibia I setae l′, l″, v′ as tiny spines. LARVAL FEMALE (Fig. 43; n = 14). Gnathosoma. Length 30–34, width 27–30. Palp with alveolar remnant of femoral seta. Cheliceral stylets 25–27, setae ch 19–23, su 8–10. Idiosoma (Fig. 43). Length 165–225, width 105–185; larger specimens swollen laterally between prodorsal shield and plate CD. Prodorsal plate with setae v1 12–14, v2 alveolar remnant, sc1 5–6, sc2 51–55. Distance between setae v1–v1 20–23, sc1–sc1 48–51, v1–sc1 25–28, v2–v2 29–35, sc2–sc2 49–54, sc1–sc2 27–33. Plates C and D fused medially; line of fusion visible. Plate CD setae c1 5–6, c2 3–4, d 4–5, cupuli ia anteriad to anteromesad d, distance between setae c1–c1 26–27, d–d 24–28. Plate EF setae f 4–5, f–f 23–26, cupuli im anteriad f. Plate H poorly expressed, caudal cone 22–30 wide, 8–13 long, bearing setae h1 ca. 65–70, h2 3–4. Venter: coxisterna 1-3 well-defined; coxal setae slender, 1a 2–3, 2a 3–4, 3a 4–5, 3b 3–4. Alveoli of 1b, 2b present. Legs. Setal counts legs I–III, femur-tarsus: 3-2-6(+ φ)-8(+ ω), 0-1-4-6(+ ω), 0-1-4-6 (setae pv′ absent on tarsi II- III). Leg I. Femur I, d minute, l’ 2–3, v’’ 5–7; genu I, l’ 2–3, l″ m–2; tibia I, d 17–21, l’ 6–7, l’’ 5–7, v’ 3–4, v’’ 7–9, k 2–3, φ 6–7; tarsus I, tc’ 8–9, tc’’ 9–10, pl’ 5–6, pl’’ 9–10, pv′ 2, pv’’ 2, ω 3–4, s 4–5, u′′ minute. Leg II. Genu II, l’ 2–3; tibia II, d 10–12, l’ 4–5, v’ 7–9, v’’ 11–13; tarsus II, tc′ 3–4, tc ″ 4–5, pl ″ 7–9, pv’’ 3–4, ω 3–4, u’ 5–6, u′′ minute. Leg III. Genu III, l’ 2; tibia III, d 10–11, l’ 4–5, v’ 7–8, v’’ 11–12; tarsus III, tc’ 4–5, tc ″ 4–5, pl ″ 8–10, u’ 5–6, pv’’ 5–6, u′′ minute. Differential diagnosis. Eutarsopolipus biuncatus sp. nov. and E. janus sp. nov. are the only members of the leytei species group to retain paired claws on tarsus I in females. The only other species of Eutarsopolipus with females retaining two claws on tarsus I is Eutarsopolipus diunculosus Eidelberg, 1994, a member of the pterostichi species group (lacking both a respiratory system and setae on genua II-III). In addition to the diagnostic features mentioned for E. janus, female specimens appear to have divided plate C. None of these specimens are of sufficient quality to be sure—they may have split when mounted—but the division of dorsal plates occurs elsewhere in Eutarsopolipus (e.g., Regenfuss 1968; Husband 2000). Etymology. The specific name biuncatus (two-hooked) refers to the two claws on tarsus I of females. Remarks. All females had waste matter that obscured morphological features. This was partially removed by clearing some specimens in 5% KOH, but no female was of high quality. The holotype was chosen on the basis of having the best quality prodorsum and legs I-II, where most diagnostic features occur. Both E. biuncatus sp. nov. and E. janus sp. nov. are unusual in retaining setae pv′ on tarsi II-III in the male, a seta usually absent in Tarsonemoidea (Lindquist 1986; Seeman 2019a). Another specimen of N. transitus from the type locality (HR# T95636) had an unusual male of another species of Eutarsopolipus (lacking all genual setae, large, blunt-tipped Y-shaped claws). Males are difficult to place in species groups as they always lack respiratory systems and sometimes lack genual setae that are present in the female.Published as part of Seeman, Owen D., 2021, Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae), pp. 1-74 in Zootaxa 4971 (1) on pages 59-63, DOI: 10.11646/zootaxa.4971.1.1, http://zenodo.org/record/477151
Eutarsopolipus umbonatus Seeman 2021, sp. nov.
No full textEutarsopolipus umbonatus sp. nov. (Fig. 11) Diagnosis. Adult female: setae sc1, c1, c2, d, f with strongly thickened bases, often blunt-tipped; anterior margin of prodorsal shield not or only slightly eroded, straight or weakly concave; setae v1 7–10. Larviform male: sc1 5–6. Larval female: Setae c1 8–11, d 7–10, f 7–10. Material examined. Holotype female ex. Castelnaudia cordata, Mt Mee SF, 6 km NNW barracks, 27° 2ʹ 53ʺ S 152° 40ʹ 49ʺ E, 8 Jan-3 Mar 1992, D. Cook, flight intercept trap, IN5148, HR# T 68607 (QMS113481). Paratypes. 16 females, 7 males, 15 larvae, as follows: all ex. C. cordata. 15 females, 6 males, 13 larvae, same data as holotype (QMS 113482–92, ANIC 52-003953 – 57, ZMH-A0015185–89); 1 female, 1 larva, McIntosh Rd, Unumbar SF, 28° 25ʹ 19ʺ S 152° 42ʹ 14ʺ E, 25 Nov 1972, G. Monteith, IN6467, HR# T 64329 (QMS 113493–94); 1 male, 1 larva, “ Glen Witheren ”, shelf scrub, 28° 2ʹ 58ʺ S 153° 6ʹ 58ʺ E, 30 Dec 1991 - 24 May 1992, G. Monteith, pitfall trap, IN5682, rainforest, HR#68606 (QMS 113495); 1 larva, Lower Coomera, 28° 12ʹ 20ʺ S 153° 10ʹ 46ʺ E, 9 Jan-6 Apr 1995, G. Monteith, pitfall trap, IN5998, HR# T 68610. All in QM except 3 females, 1 male, 1 larva in each of ANIC and ZMH. Other material examined. 5 females, 2 larvae, on same slide as specimens of E. hebronae, same data as holotype. Not included as paratypes. Description. FEMALE (Fig. 11; n = 17). Gnathosoma. Length 62 (56–61), width 57 (55–58). Cheliceral stylets 68 (68–70), setae ch 22 (25–30), su 10 (10–18). Idiosoma (Fig. 11). Length 330 (320–400), width 290 (250–325). Prodorsal plate slightly or not eroded midanteriorly, more substantially eroded anterolaterally; setae v1 7 (7–10), v2 alveolar remnant, sc1 usually thickened, thorn-like 7 (7–12) [14/ 16 in specimen #5, 12/ 15 in specimen #8, 9/ 15 in specimen #11], sc2 50–80. Distance between setae v1–v1 47 (41–49), sc1–sc1 105 (94–105), v1–sc1 38 (32–36), v2–v2 45 (41–50), sc2–sc2 130 (125– 130), sc1–sc2 27 (25–32). Opisthosomal setae c1, c2, d, f usually stout but sometimes with fine tips. Plate C setae c1 7 (6–12; 15 in specimen #8), c2 10 (10–15), distance between setae c1–c1 84 (82–100). Plate D setae d 10 (10–15), d–d 155 (160–185), cupuli ia laterad d. Plate EF setae f 7/10 (asymmetry) (7–12; 10/ 13 in specimen #8; 15 in specimen #15), f–f 115 (130–150), cupuli im laterad f. Plate H divided or fused weakly, seta h 7 (5–7). Venter: coxisterna 1 well defined, coxisterna 2 well defined but sometimes with folds, coxisterna 3 weakly defined; coxal setae slender, 1a 4 (3–4), 2a 6 (5–6), 3a thickened, 3–6, 3b 4–6. Alveoli of 1b, 2b present. Legs (holotype only, except if holotype different from E. hadros / mixtus). Setal counts legs I–III, femur-tarsus: 3-2-6(+ φ)-8(+ ω), 0-1-4-6, 0-1-4-6. Leg I. Femur I, d minute, l’ 2, v″ 14; genu I, l′ 3, l″ 3; tibia I, d 60, l’ 7, l’’ 10, v’ 5, v’’ 8, k 5, φ 8 (7–9); tarsus I, tc’ 14, tc’’ 15, pl’ 7, pl’’ 13, pv′ 3, pv’’ 4, ω 4, s 7, u′′ minute. Leg II. Genu II, l′ 4; tibia II, d 40, l’ 7, v’ 12, v’’ 23; tarsus II, tc′ 2, tc ″ 6, pl ″ 55, pv’’ 4, u’ 7, u′′ minute. Leg III. Genu III, l′ 3; tibia III, d 29, l’ 8, v’ 11, v’’ 21; tarsus III, tc’ 2, tc ″ 6, pl ″ 52, pv’’ 4, u’ 8, u′′ minute. LARVIFORM ADULT MALE (n = 7). Gnathosoma. Length 33–35, width 30–32. Cheliceral stylets 25–28, setae ch 6–7, su 5–7. Idiosoma. Length 170–185, width 120–130. Prodorsal plate with setae v1 3–5, v2 alveolar remnant, sc1 5–6, sc2 70–80. Distance between setae v1–v1 19–21, sc1–sc1 46–50, v1–sc1 22–26, v2–v2 19–22, sc2–sc2 57–63, sc1–sc2 20–21. Plate CD setae c1 5–7, c2 4–6, d 5–6, distance between setae c1–c1 42–45, c1–c2 27–31, d–d 40–43; cupuli ia anterolaterad setae d. Plate EF eroded medially, sometimes almost divided; with setae f 2–4, distance f–f 27–28; cupuli im anterolaterad to laterad setae f. Genital capsule length 28–31, width 31–32, setae h minute. Venter: coxal setae 1a 2–3, 2a 4, 3a 2–3, 3b 4–5. Legs. Setal counts legs I–III, femur-tarsus: 3-2-6(+ φ)-8(+ ω), 0-1-4-6, 0-1-4-6. Leg I. Femur I, d minute, l’ minute, v″ 7; genu I l′ 2, l″ 2; tibia I, d 44, l’ 2, l’’ 3, v ’ 3, v ’’ 7, k 4, φ 8; tarsus I, tc’ 12, tc’’ 12, pl’ 7, pl’’ 12, pv ′ 2, pv’’ 2, ω 3, s 5, u ′′ 2. Leg II. Genu II, l′ 2; tibia II, d 18, l’ 5, v’ 10, v’’ 17; tarsus II, tc′ 3, tc ″ 4, pl ″ 28, pv’’ 3, u’ 5, u″ 2. Leg III. Tibia III, d 14, l’ 5, v’ 11, vʺ 11; tarsus III, tc′ 3, tc ″ 4, pl ″ 22, pv’’ 3, u’ 5, u″ minute. LARVAL FEMALE (n = 15). Gnathosoma. Length 48–52, width 46–54. Cheliceral stylets 63–70, setae ch 27–29, su 7–9. Idiosoma. Length 195–280, width 135–215. Prodorsal plate with setae v1 4–7, v2 alveolar remnant, sc1 5–8, sc2 100–120. Distance between setae v1–v1 28–37, sc1–sc1 65–75, v1–sc1 23–26, v2–v2 32–37, sc2–sc2 71–85, sc1–sc2 32–37. Plate C setae c1 8–11, c2 5–10, distance between setae c1–c1 40–48. Plate D setae d 7–10, d–d 49–63, cupuli ia very close to setae d, anteriad to anterolaterad d. Plate EF setae f 7–10, f–f 48–50, cupuli im close to setae f, anteriad f. Plate H narrow, 13–15 wide, ca. 25–35 long bearing setae h1 ca. 170, h2 6–8 (Mt Mee specimens 12–15). Venter: coxisterna 1-2 well defined, coxisterna 3 poorly defined, membranous; coxal setae 1a 2–3, slender, 2a 4–5, slender, 3a 3–5 sometimes slightly thickened, 3b 4–5, slender. Alveoli of 1b, 2b present. Legs. Setal counts legs I–III, femur-tarsus: 3-2-6(+ φ)-8(+ ω), 0-1-4-6, 0-1-4-6. Leg I. Femur I, d minute, l′ 2, v″ 8; genu I, l′ 3, l″ 2; tibia I, d 62, l’ 9, l’’ 10, v’ 3, v’’ 11, k 5, φ 10; tarsus I, tc’ 12, tc’’ 13, pl’ 10, pl’’ 16, pv′ 2, pv’’ 3, ω 3, s 5, u′′ 2. Leg II. Genu II, l′ 3; tibia II, d 35, l’ 5, v’ 15, v’’ 15; tarsus II, tc′ 2, tc ″ 4, pl ″ 37, pv’’ 3, u’ 6, u′′ minute. Leg III. Genu III, l′ 3; tibia III, d 33, l’ 6, v’ 13, v’’ 15; tarsus III, tc’ 2, tc ″ 4, pl ″ 32, pv’’ 3, u’ 6, u′′ minute. Differential diagnosis. Eutarsopolipus umbonatus sp. nov. is similar to E. mixtus sp. nov. by having much stronger dorsal setae compared with E. hadros. However, like all species and particularly E. mixtus, the form of these setae differed intraspecifically: although the thicker nature of these setae are useful for separating E. umbonatus from E. hadros, they are too variable to separate E. umbonatus from E. mixtus. Larvae are more useful for distinguishing the latter two species. Larva of E. umbonatus have smaller idiosomal setae: setae c1 (8–11 versus 13–20), d (7–10 versus 11–12) and f (7–10 versus 14–19) are slightly to obviously shorter in E. umbonatus. Etymology. The specific name umbonatus (knob-like) refers to the short, more knob-like setae in females of this species. Remarks. In larvae, setae h2 were longer in the type series from Mt Mee (12–15) compared with the three specimens from other sites (6–8). No other features distinguished these populations. However, Mt Mee is part of a mountain range (the D’Aguilar Range) to the northwest of Brisbane, while all other collections were from the Border Range to the south to southwest of Brisbane. The geographical separation of these populations, combined with the difference in length of seta h, suggests that these may be separate species on isolated populations of Castelnaudia cordata. Furthermore, these differences also suggest that the host species deserves closer attention. The D’Aguilar Range has several endemic taxa, such as a unique assemblage of anamid spiders (Rix et al. 2020). Likewise, the beetle specimens from the D’Aguilar Range may be an endemic species closely related to C. cordata. The above three species, E. hadros, E. mixtus and E. umbonatus, are very closely related and it is tempting to consider all three as one species. However, each species does exhibit subtle differences, they occur on different host species, and are highly isolated geographically with the hosts C. setosiceps, C. mixta and C. cordata in north-east Queensland, middle-east Queensland and south-east Queensland, respectively. One paratype female has a fully developed larva within its body, with no egg, indicating vivipary. Two larvae of another species close to Eutarsopolipus echinatus Seeman, 2019a (i.e. same thorn-like setae, but clearly a different species) were also collected from C. cordata: one from the type host (on a separate slide) and one from Lower Coomera and on the same slide as the larva of E. umbonatus (QMS 113496). This species was found co-infesting its host with E. hebronae sp. nov. and E. osculum sp. nov. (Tables 1, 2). The ochoai species group - nahmani subgroup Diagnosis. Female with dorsal idiosomal setae fine, slender; setae h small but developed. All life stages with gnathosomal apex evenly rounded. The ochoai species group - nahmani subgroup Diagnosis. Female with dorsal idiosomal setae fine, slender; setae h small but developed. All life stages with gnathosomal apex evenly rounded.Published as part of Seeman, Owen D., 2021, Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae), pp. 1-74 in Zootaxa 4971 (1) on pages 21-24, DOI: 10.11646/zootaxa.4971.1.1, http://zenodo.org/record/477151
Eutarsopolipus janus Seeman 2021, sp. nov.
No full textEutarsopolipus janussp. nov. (Figs 44–48) Diagnosis. Adult female: leg I with two claws; v1 19–24, sc2 36–40; genu I with two setae (l″ present); tarsus II setae tc″ 8–9, u′ 8–10. Adult male: setae sc2 27–34. Larval female: sc2 70–80. Material examined. Holotype female, ex. Notonomus flos. Mt Lewis Rd, 29 km from Highway, Cow Bay, 16° 13ʹ 51ʺ S 145° 27ʹ 43ʺ E, 18 Nov 1997 – 9 Feb 1998, G. Monteith & D. Cook, pitfall trap, rainforest, IN1702, HR# T 65790 (QMS 113527). Paratypes. All ex Notonomus flos. 7 females, 7 males, 9 larvae, as follows: 7 females, 7 males, 7 larvae, same data as holotype (QMS 113528–42, ANIC 52-003978 – 80, ZMH-A0015204–05); 2 larvae, same data except different beetle, HR# T 65789 (QMS 113543, ZMH-A0015206). All in QM except 1 female, 1 male, 1 larva in each of ANIC and ZMH. Other material examined. Ex N. aurifer: 7 females, 5 males, 7 larvae, as follows: 6 females, 2 males, 4 larvae, Lamb’s Head, 20 km SW Cairns, 17° 1ʹ 23ʺ S 145° 38ʹ 33ʺ E, 10–12 Dec 1989, G. Monteith, G. Thompson, H. Janetzki, HR# T 20079 (QMS 113544–54, 1 female ANIC); 2 males, 1 larva, same data except different beetle, HR# T 20078 (QMS 113555–57); 1 female, 1 male, 2 larvae, same data except different beetle, HR# T 20081 (QMS 113558–61). Ex N. spurgeoni: 5 females, 3 males, 4 larvae, as follows: 1 female, Mossman Bluff Track site 8, 16° 27ʹ 36ʺ S 145° 17ʹ 12ʺ E, 17–19 Dec 1988, G. Monteith, G. Thompson, ANZSES, HR# T 20799 (QMS 113591); 1 female, Mossman Bluff Track general, 16° 28ʹ S 145° 18ʹ E, 20 Dec 1989 – 15 Jan 1990, G. Monteith, G. Thompson, ANZSES, HR# T 20863 (QMS 113592); 1 male, 4 larvae, same data except different beetle, HR# T 20849 (QMS 113598–602); 3 females, 2 males, Mt Spurgeon, 7 km N, camp 2, 16° 22ʹ 31ʺ S 145° 12ʹ 49ʺ E, 17–19 Oct 1991, G. Monteith, H. Janetzki, D. Cook, L. Roberts, HR# T 20957 (QMS 113593-97). Ex N. dimorphicus: 1 female, 1 male, 3 larvae, as follows: 1 female, 1 larva, Mt Lewis, barracks, 16° 35ʹ 39ʺ S 145° 16ʹ 28ʺ E, no date, G. Wood, ex HR# T 20722 (QMS 113603); 1 male, 1 larva, Mt Lewis Rd, 22 km from Highway, ANZSES site 3, 16° 32ʹ 48ʺ S 145° 16ʹ 53ʺ E, 29 Nov 1997 – 9 Feb 1998, G. Monteith, D. Cook, rainforest, IN1701, HR# T 65741 (QMS 113605–06); 1 larva, Mt Lewis, 2.5 km N, via Julatten, 16° 33ʹ S 145° 15ʹ E, 3 Nov 1983, D. Yeates, G. Thompson, HR# T 20721 (QMS 113607). Note: One male is fully developed within a broken egg and is included as a paratype. Description (excluding specimens from N. aurifer). FEMALE (Figs 44–46; n = 21). Gnathosoma. Length 55 (52–57), width 51 (45–51). Palp with alveolar remnant of femoral seta. Cheliceral stylets 45 (45–49), setae ch 25 (25–35), su 18 (15–20). Idiosoma (Figs 44–45). Length ca. 340 (290–600), width ca. 240 (230–450), bigger specimens swollen, distorted. Stigmata apparent, large, in soft cuticle anteriad setae v1 to posterolaterad gnathosoma (varies according to mount), respiratory system not visible in most mounts but sometimes narrow trachea and small atrium apparent. Prodorsal plate with setae v1 20 (19–24), v2 alveolar remnant, sc1 10 (9–11), sc2 38 (36–40). Distance between setae v1–v1 56 (53–60), sc1–sc1 88 (87–96), v1–sc1 31 (34–37), v2–v2 69 (68–75), sc2–sc2 95 (91–95), sc1–sc2 37 (32–36). Plate C entire, setae c1 8 (8–9), c2 8 (8–10). Plate D setae d 9 (8–10), d–d 105 (110–125), cupuli ia anterolaterad d. Plate EF setae f 9 (7–8), f–f 71 (80–87), cupuli im anterolaterad to laterad f. Plate H present as small remnant. Venter: coxisterna 1-2 poorly defined, slightly broader than apodemes; coxisterna 3 well-defined; coxal setae slender, 1a 6 (6–7), 2a 6 (6–7), 3a 6 (7–9), 3b 6 (6–7). Alveoli of 1b, 2b present. Legs (Fig. 46). Setal counts legs I– III, femur-tarsus: 3-2-6(+ φ)-9(+ ω), 0-1-4-7(+ ω), 0-1-4-7. Leg I. Femur I, d 2 (2), l’ 13 (13–16), v″ 14 (12–15); genu I, l’ 5 (5–6), l″ 3 (3); tibia I, d 25 (27–30), l’ 10 (7–9), l’’ 6 (6–8), v’ 7 (7–8), v’’ 14 (11–12), k 4 (4–5), φ 6 (7–8); tarsus I, tc’ 13 (11–12), tc’’ 13 (13–15), pl’ 15 (14–17), pl’’ 18 (15–20), pv′ 3 (3–4), pv’’ 3 (2–3), ω 4 (4–5), s 6 (6–7), u′′ minute (m–2), p′ 2 (m–2). Leg II. Genu II, l’ 8 (5–6); tibia II, d 16 (15–20), l’ 9 (8–9), v’ 15 (14–16), v’’ 21 (20–23); tarsus II, tc′ 9 (9–11), tc ″ 8 (8–9), pl ″ 18 (16–18), pv′ 2 (2–3), pv’’ 4 (3–4), ω 4 (4–5), u’ 9 (9–10), u″ m (m), difficult to discern next to large u′. Leg III. Genu III, l’ 5 (4–5); tibia III, d 14 (15–20), l’ 9 (7–9), v’ 12 (12–15), v’’ 17 (20–21); tarsus III, tc’ 8 (8–10), tc ″ 8 (8–9), pl ″ 20 (18–20), pv′ 4 (3–5), pv’’ 3 (3–5), u’ 8 (8–9), u″ m (m), difficult to discern next to large u′. Claws on legs I-III well developed and paired. Tarsi I-III setae pvʹ, pvʺ slender. Large blunt spur-like setae: tarsus I s; tarsi II– III u’, tcʺ. Setae u′ pointed, not bifid. LARVIFORM ADULT MALE (Fig. 47; n = 16). Gnathosoma. Length 28–37, width 29–33. Cheliceral stylets 25–28, setae ch 3–4, su 8–9. Idiosoma (Fig. 47). Length 175–190, width 135–155. Prodorsal plate with setae v1 4–9, v2 alveolar remnant, sc 1 m–2, sc2 27–34. Distance between setae v1–v1 24–30, sc1–sc1 61–66, v1–sc1 24–26, v2–v2 42–49, sc2–sc2 64–72, sc1–sc2 24–28. Plate CD eroded posterolaterally, setae c1 2–3, c2 2, d m–2, distance between setae c1–c1 56–65, c1–c2 32–35, d–d 42–47; cupuli ia laterad to anterolaterad setae d. Plate EF with setae f m–2, distance f–f 22–25; cupuli im laterad to anterolaterad setae f. Genital capsule length 25–27, width 35–38, setae h minute. Venter: coxal setae slender, 1a 4–5, 2a 4–5, 3a 5–6, 3b 5–6. Legs. Setal counts legs I– III, femur-tarsus: 2-0-6(+ φ)-8(+ ω), 0-0-4-7(+ ω), 0-0-4-7. Compared to female: Femur I seta v″ absent, genua I-III l′ and l″ absent. Leg I. Femur I, d 2, l’ 2–4; tibia I, d 19–25, l’ 2, l’’ 2–3, v ’ 2–3, v ’’ 9–10, k 2–3, φ 6–7; tarsus I, tc’ 8–9, tc’’ 10–11, pl’ 8–11, pl’’ 10–12, pv ′ 2, pv’’ 2, ω 3–4, s 5, u ′′ minute. Leg II. Tibia II, d 6–11, l’ 3–4, v’ 9–12, v’’ 11–14; tarsus II, tc′ 6–8, tc ″ 5–6, pl ″ 11–12, pv ′ 2, pv’’ 3–4, ω 3–4, u’ 6, u″ minute, difficult to discern. Leg III. Tibia III, d 2, l’ 3–5, v’ 8–11, vʺ 10–14; tarsus III, tc′ 3–4, tc ″ 6–7, pl ″ 9–11, pv ′ 2–3, pv’’ 3–5, u ’ 6–7, u″ minute, difficult to discern. Claws on leg I single, claws on leg II-III paired. Setal form similar to that in female, but with tibia I setae l′, l″ tiny spines and v′ slightly thickened. LARVAL FEMALE (Fig. 48). Gnathosoma. Length 38–46, width 31–34. Palp without alveolar remnant of femoral seta. Cheliceral stylets 36–42, setae ch 26–33, su 12–15. Idiosoma (Fig. 48). Length 180–225, width 135–240; larger specimens swollen evenly around body. Prodorsal plate with setae v1 15–18, v2 alveolar remnant, sc1 8–10, sc2 70–80. Distance between setae v1–v1 29–31, sc1–sc1 69–72, v1–sc1 31–33, v2–v2 50–54, sc2–sc2 72–78, sc1–sc2 33–37. Plates C and D fused medially; line of fusion visible. Plate C setae c1 7–10, c2 7–10, distance between setae c1–c1 31–35. Plate D setae d 7–9, d–d 30–35, cupuli ia anterolaterad d. Plate EF setae f 6–8, f–f 29–33, cupuli im anteriad to anterolaterad f. Plate H small, 18–22 wide, 13–18 long, bearing setae h1 ca. 100, h2 4–6. Venter: coxisterna 1-3 well-defined; coxal setae slender, 1a 4–5, 2a 4–5, 3a 7–8, 3b 5–6. Alveoli of 1b, 2b present. Legs. Setal counts legs I– III, femur-tarsus: 3-2-6(+ φ)-8(+ ω), 0-1-4-6(+ ω), 0-1-4-6 (setae pv′ absent on tarsi II- III). Leg I. Femur I, d m–2, l’ 2–3, v’’ 9–13; genu I, l’ 3, l″ 2–3; tibia I, d 25–32, l’ 7–8, l’’ 7–9, v’ 4–5, v’’ 8–12, k 3–4, φ 7–8; tarsus I, tc’ 8–9, tc’’ 10–11, pl’ 8–10, pl’’ 13–15, pv′ 2–3, pv’’ 2–3, ω 4–5, s 5–6, u′′ 2. Leg II. Genu II, l’ 4–5; tibia II, d 13–17, l’ 8–9, v’ 9–12, v’’ 13–16; tarsus II, tc′ 7–8, tc ″ 5–6, pl ″ 13–17, pv’’ 4–5, ω 4–5, u’ 6–7, u′′ minute. Leg III. Genu III, l’ 3–4; tibia III, d 15–17, l’ 5–6, v’ 10–11, v’’ 16–17; tarsus III, tc’ 7–9, tc ″ 5–7, pl ″ 16–20, u’ 6–7, pv’’ 5–6, u′′ minute. Setal fprm similar to that in female. Note: one larva (#2) has an unusual asymmetrical reversal of tcʺ and plʺ. Differential diagnosis. Eutarsopolipus janus sp. nov. and E. biuncatus sp. nov. both have the unusual feature of paired claws on leg I in females. Female Eutarsopolipus janus differ by having longer setae sc2 (36–40 versus 15–20) and tarsi II-III with longer setae tc′ (8–11 versus 3–7) and tc′′ (8–9 versus 5–6). Males are even more distinctive, with seta sc2 being well-developed in E. janus (27–34) but greatly reduced in E. biuncatus (2–3). Etymology. The specific name janus refers to the god Janus, who symbolised beginnings and transitions, alluding to the questions regarding the evolution and speciation of this possible complex of cryptic species on several host species. Remarks. This species was found on four host species: the type host, N. flos, as well as N. aurifer, N. dimorphicus and N. spurgeoni. All these species are found in the Wet Tropics region of north Queensland and at least N. aurifer, N. dimorphicus and N. flos are closely related (Will 2015). Thus, these species may not have speciated when their hosts speciated, or they have done so cryptically. In support of the latter hypothesis, some minor differences were observed in N. aurifer. Females of this species differed by having: slightly longer cheliceral stylets (50–56 versus 43–50), slightly longer tibia I solenidion (6–8 versus 8–10), slightly longer seta tc′ on tarsus II (10–13 versus 8–10) and tarsus III (11–13 versus 8–10), slightly longer setae tcʺ on tarsus II (9–10 versus 8–9) and tarsus III (10–11 versus 8–9) and u′ on tarsus III (9–11 versus 8–9). Several of these differences are also apparent in the larvae. Larvae from N. aurifer have longer cheliceral stylets (43–49 versus 36–42), longer tarsal seta tc′ on tarsus III (10–12 versus 7–9) and tarsal seta tc″ on tarsus II (6–7 versus 5–6). The species N. aurifer is also geographically separated from the other three species: the former is from a large fragment of the Wet Tropics south-west of Cairns, while the others are from a larger fragment southwest to west of Mossman comprising Mt Lewis (N. flos), Mt Spurgeon (N. spurgeoni) and Lamb’s Head (N. dimorphicus). These regions are separated by a biogeographical barrier, the Black Mountain Corridor, which separates radiations of other endemic rainforest animals (e.g., Krosch et al. 2009; Edwards & Melville 2010) and plants (e.g., Burke et al. 2013). Furthermore, N. dimorphicus and N. flos are more closely related to each other than they are to N. aurifer (Will 2015).Published as part of Seeman, Owen D., 2021, Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae), pp. 1-74 in Zootaxa 4971 (1) on pages 64-70, DOI: 10.11646/zootaxa.4971.1.1, http://zenodo.org/record/477151
Eutarsopolipus mixtus Seeman 2021, sp. nov.
No full textEutarsopolipus mixtussp. nov. (Figs 9–10) Diagnosis. Adult female: setae sc1, c1, c2, d, f with strongly thickened bases, often blunt-tipped; anterior margin of prodorsal shield eroded, concave; setae v1 10–13. Larviform male: sc1 3–6. Larval female: Setae c1 13–20, d 11–12, f 14–19. Material examined. Holotype female ex. Castelnaudia mixta. Broken River, Eungella, 21° 10′ 05″ S 148° 30′ 30″E, 17 Nov 1992 - 15 Apr 1993, G. Monteith & D. Cook, flight intercept & pitfall trap, HR# T 82585 (QMS 113035). Paratypes. 12 females, 4 males, 4 larvae, as follows: 11 females, 3 males, 1 larva, same data as holotype (QMS 113036-43, 113045, 113047, ANIC 52-003950 – 51, ZMH-A0015182–83); 1 male, 1 larva, same data except HR# T 82586 (QMS 113046, ZMH-A0015184). 1 female, same data except HR# T 82582 (QMS 113044); 1 larva, same data except HR# T 82584 (ANIC 52-003952); 1 larva, Pearse’s Lookout, Eungella, 21° 06′ 53″ S 148° 31′ 06″S, 17 Nov 1992 - 15 Apr 1993, G. Monteith & D. Cook, IN5934, HR# T 82526 (QMS 113048). All in QM except 1 female, male, larva in ANIC and ZMH. Description. FEMALE (Figs 9–10; n = 13). Gnathosoma. Length 59 (57–65), width 55 (53–60). Cheliceral stylets 75 (67–78), setae ch 18 (16–25), su 17 (11–18). Idiosoma (Fig. 9). Length 395 (365–400), width 280 (270–330). Prodorsal plate eroded mid-anteriorly and anterolaterally; setae v1 13 (10–13), v2 alveolar remnant, sc1 thickened, 11/16 (asymmetry) (11–17), sc2 75 (70–95) (Fig. 10). Distance between setae v1–v1 48 (41–52), sc1–sc1 94 (82–100), v1–sc1 33 (26–40), v2–v2 43 (41–51), sc2–sc2 130 (130–140), sc1–sc2 35 (24–34). Opisthosomal setae c1, c2, d, f thickened, sometimes appearing stout by lacking fine tips, especially setae c2. Plate C setae c1 12/20 (asymmetry) (10–20), c2 9 (10–15), distance between setae c1–c1 100 (78–95). Plate D setae d 17 (12–21), d–d 145 (135–150), cupuli ia laterad d. Plate EF setae f 13/18 (asymmetry) (11–19), f–f 135 (110–125), cupuli im laterad f. Plate H divided into two small plates, or fused weakly, often ventral, seta h 5 (5–6). Venter: coxisterna 1 and 2 well defined, coxisterna 3 weakly defined; coxal setae slender, 1a 4 (3–4), 2a 8 (6–8), 3a thickened, 5 (5–7), 3b 9 (7–9). Alveoli of 1b, 2b present. Legs (holotype only, except if holotype different from E. hadros). Setal counts legs I–III, femur-tarsus: 3-2- 6(+ φ)-8(+ ω), 0-1-4-6, 0-1-4-6. Leg I. Femur I, d minute, l’ 4 (3–5), v″ 11; genu I, l′ 2, l″ 3; tibia I, d 65, l’ 8, l’’ 10, v’ 5, v’’ 10, k 5, φ 7 (7–9); tarsus I, tc’ 16, tc’’ 15, pl’ 9, pl’’ 13, pv′ 3, pv’’ 3, ω 3, s 7, u′′ minute. Leg II. Genu II, l′ 3; tibia II, d 57, l’ 7, v’ 13, v’’ 19; tarsus II, tc′ 2, tc ″ 6, pl ″ 60, pv’’ 4, u’ 7, u′′ minute. Leg III. Genu III, l′ 3; tibia III, d 42, l’ 7, v’ 11, v’’ 15; tarsus III, tc’ 2, tc ″ 5, pl ″ 39, pv’’ 3, u’ 6, u′′ minute. LARVIFORM ADULT MALE (n = 4). Gnathosoma. Length 32–35, width 30–34. Cheliceral stylets 23–27, setae ch 5–7, su 5–10. Idiosoma. Length 160–205, width 125–150. Prodorsal plate with setae v1 4–5, v2 alveolar remnant, sc1 3–6, sc2 60–70. Distance between setae v1–v1 21–26, sc1–sc1 40–52, v1–sc1 20–25, v2–v2 22–27, sc2–sc2 57–77, sc1–sc2 16–22. Plate CD setae c1 4–6, c2 4–5, d 4–5, distance between setae c1–c1 46–59, c1–c2 22–28, d–d 36–52; cupuli ia anterolaterad setae d. Plate EF eroded medially, almost divided; with setae f 2–3, distance f–f 24–30; cupuli im anterolaterad to laterad setae f. Genital capsule length 28–30, width 34–38, setae h minute. Venter: coxal setae 1a 2–3, 2a 4–5, 3a 3–4, 3b 5–6. Legs. Setal counts legs I–III, femur-tarsus: 3-2-6(+ φ)-8(+ ω), 0-1-4-6, 0-1-4-6. Leg I. Femur I, d minute, l′ 2, v″ 7; genu I l′ 2, l″ 2; tibia I, d 38, l’ 3, l’’ 2, v’ 3, v’’ 10, k 4, φ 6–8; tarsus I, tc’ 11–14, tc’’ 11–14, pl’ 7, pl’’ 11, pv ′ 2, pv’’ 3, ω 3, s 4, u ′′ minute. Leg II. Genu II, l′ 2; tibia II, d 16, l’ 5, v’ 10, v’’ 13; tarsus II, tc′ 2, tc ″ 4, pl ″ 27, pv’’ 3, u’ 5, u″ 2. Leg III. Tibia III, d 16, l’ 4, v’ 12, v″ 12; tarsus III, tc′ 2, tc ″ 3–5, pl ″ 20, pv’’ 3, u’ 5, u″ present. LARVAL FEMALE (n = 4). Gnathosoma. Length 46–51, width 43–49. Cheliceral stylets 61–64, setae ch 23–27, su 7–10. Idiosoma. Length 205–260, width 165–215. Prodorsal plate overlapping folded membranous cuticle, with setae v1 6–8, v2 alveolar remnant, sc1 7–10, sc2 120–135. Distance between setae v1–v1 31–35, sc1–sc1 59–63, v1–sc1 23–25, v2–v2 30–34, sc2–sc2 72–83, sc1–sc2 31–33. Plate C setae c1 13–20, c2 8–11, distance between setae c1–c1 42–48. Plate D setae d 11–12, d–d 53–61, cupuli ia very close to setae d, anteriad to anterolaterad d. Plate EF setae f 14–19, f–f 46–52, cupuli im close to setae f, anterolaterad f. Plate H narrow, 15–20 wide, ca. 25–30 long bearing setae h1 ca. 150, h2 13–17. Venter: coxisterna 1-2 well defined, coxisterna 3 poorly defined, membranous; coxal setae slender, 1a 3, 2a 4–5, 3a 4–5, 3b 4–5. Alveoli of 1b, 2b present. Legs. Setal counts legs I–III, femur-tarsus: 3-2-6(+ φ)-8(+ ω), 0-1-4-6, 0-1-4-6. Leg I. Femur I, d minute, l′ 2, v″ 7; genu I, l′ 2, l″ 3; tibia I, d 54, l’ 9, l’’ 10, v’ 4, v’’ 8–12, k 6, φ 10; tarsus I, tc’ 13, tc’’ 15, pl’ 8, pl’’ 16, pv′ 2, pv’’ 3, ω 3, s 5, u′′ minute. Leg II. Genu II, l′ 4; tibia II, d 40, l’ 7, v’ 14, v’’ 16; tarsus II, tc′ 2, tc ″ 4, pl ″ 35, pv’’ 3, u’ 5, u′′ 2. Leg III. Genu III, l′ 4; tibia III, d 35, l’ 7, v’ 13, v’’ 14; tarsus III, tc’ 2, tc ″ 4, pl ″ 21–40, pv’’ 3, u’ 6, u′′ minute. Differential diagnosis. Eutarsopolipus mixtus sp. nov. is similar to E. hadros sp. nov. but is most easily distinguished by having thicker dorsal setae. This feature is useful for species separation, but is complicated by intraspecific variation in setal form, which is often expressed asymmetrically. However, setae sc1 are always thicker in E. mixtus, even if they taper. Likewise, other dorsal idiosomal setae tend to be thicker in E. mixtus, exemplified by setae f (Fig. 10). The prodorsal shield of E. umbonatus is weakly eroded anteriorly, in contrast to the deep erosion of E. hadros and lack of erosion in E. umbonatus. However, the intermediate state of this feature reduces its effectiveness. In larvae, E. mixtus have longer setae f (14–19 versus 10–11 in E. hadros and 7–10 in E. umbonatus). Etymology. The species name mixtus (Latin, mixed) refers both to the host species C. mixta but also the variation in form of the dorsal setae. Remarks. Three paratype females have fully developed larvae within their bodies, with no egg, suggestive of vivipary. Two additional specimens of C. mixta from Finch Hatton Gorge (HR# T82535, T82529) had a male and a larva (one per host) of an undermined species of Eutarsopolipus (Table 1).Published as part of Seeman, Owen D., 2021, Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae), pp. 1-74 in Zootaxa 4971 (1) on pages 19-21, DOI: 10.11646/zootaxa.4971.1.1, http://zenodo.org/record/477151
Eutarsopolipus savatus Seeman 2021, sp. nov.
No full text<i>Eutarsopolipus savatus</i> sp. nov. <p>(Figs 39–40)</p> <p> <b>Diagnosis.</b> Adult female: <i>sc1</i> 11–19, <i>c1</i> 11–19, <i>c2</i> 14–20, <i>d</i> 12–19, <i>f</i> 8–12, <i>v2-v2</i> 48–58, <i>sc1-sc1</i> 76–88, <i>d-d</i> 120– 160, <i>f-f</i> 95–120; genua II-III setae <i>l′</i> 2–4, tibia I setae <i>l′</i> 8–10, <i>v′</i> 4–5, tarsi II-III setae <i>tc′′</i> 4–5. Adult male: leg I with one claw; <i>c1</i> 4–5, <i>c2</i> 5–6, <i>v2-v2</i> 34–41; tarsi II-III setae <i>tc″</i> 3–4. Larval female: <i>c1</i> 6–9, <i>c2</i> 9–12, <i>d</i> 7–10, <i>f</i> 7, <i>sc2-sc2</i> 78–100, <i>v1-sc1</i> 22–24; tarsi II-III setae <i>tc′′</i> 3.</p> <p> <b>Material examined</b>. Holotype female, ex. <i>Castelnaudia eungella.</i> Pease’s Lookout, Eungella, 21° 6ʹ 53ʺ S 148° 31ʹ 6ʺ E, 17 Nov 1992 - 15 Apr 1993, D. Cook, G. Monteith, flight intercept & pitfall trap, IN5934, HR# T 82502 (QMS 113497). Paratypes. All ex. <i>C. eungella</i>. 9 females, 4 males, 11 larvae, as follows: 1 female, 4 larvae same data as holotype (QMS 113498, 113500–503); 2 larvae, same data except another beetle, HR# T 82503 (QMS 113504, 113578); 2 females, 1 male, 2 larvae, same data except another beetle, HR# T 82506 (QMS 113577, 11379–80, ANIC 52-003973, ZMH-A0015199); 2 females, 1 male, 1 larva, Mt Dalrymple, 21° 1ʹ 37ʺ S 148° 38ʹ 2ʺ E, 21 Dec 1992 – 10 Jan 1993, ANZSES, HR# T 82501 (QMS 113581–83, ZMH-A0015200); 1 female, 2 larvae, Eungella National Park, Mt Dalrymple track, 1200 m site, 21.026° S 148.627°E, 23 Nov 2013, C. Burwell, rainforest, sampcode 25933, beetle CB6 (QMS 113584–85, ANIC 52-003974); 1 female, 1 male, Mt William, 21° 1ʹ 5ʺ S 148° 35ʹ 57ʺ E, 21 Dec 1992 – 10 Jan 1993, ANZSES, HR# T 82489 (QMS 113586–87); 2 females, 1 male, Upper Cattle Creek, Eungella, 21° 1ʹ 41ʺ S 148° 36ʹ 11ʺ E, 17 Nov 1992, G. Monteith, G. Thompson, H. Janetzki, HR# T 82513 (QMS 113588–90). All in QM except 1 female and 1 larva in each of ANIC and ZMH.</p> <p> <b>Other material examined</b>. 1 pharate female ex HR# T 82503 (QMS 113505).</p> <p> <b>Description.</b> FEMALE (Figs 39–40; n = 10).</p> <p> <i>Gnathosoma.</i> Length 43 (43–48), width 41 (40–45), apex indented, terminating in small but distinct lobes. Cheliceral stylets 44 (38–40), setae <i>ch</i> 31 (24–39), <i>su</i> 14 (13–16).</p> <p> <i>Idiosoma</i> (Fig. 39). Length 360 (305–385), width 405 (250–350). Prodorsal plate not eroded anteriorly; setae slender, <i>v1</i> 9 (6–9), <i>v2</i> alveolar remnant, <i>sc1</i> 16 (11–19; one specimen asymmetrical 10/15), <i>sc2</i> 80 (80–110). Distance between setae <i>v1–v1</i> 45 (37–45), <i>sc1–sc1</i> 82 (73–88), <i>v1–sc1</i> 26 (often asymmetrical, 23–32), <i>v2–v2</i> 58 (48–58), <i>sc2–sc2</i> 130 (125–135), <i>sc1–sc2</i> 38 (often asymmetrical, 31–46). Opisthosomal setae slender. Plate C setae <i>c1</i> 19 (9–16), <i>c2</i> 19 (14–20), distance between setae <i>c1–c1</i> 85 (80–115). Plate D setae <i>d</i> 19 (12–19), <i>d–d</i> 140 (120–160), cupuli <i>ia</i> anterolaterad <i>d</i>. Plate EF setae <i>f</i> 10 (8–12), <i>f–f</i> 115 (95–120), cupuli <i>im</i> anteriad to anterolaterad <i>f</i>. Plate H divided, seta <i>h</i> minute. Venter: coxisterna 1-2 moderately well defined, with few folds, coxisterna 3 membranous; coxal setae slender, <i>1a</i> 4 (3–4), <i>2a</i> 5 (4–5), <i>3a</i> 6 (5–6), <i>3b</i> 6 (6–7). Alveoli of <i>1b, 2b</i> present.</p> <p> <i>Legs</i> (Fig. 40). Setal counts legs I–III, femur-tarsus: 3-2-6(+ <i>φ</i>)-8(+ <i>ω</i>), 0-1-4-6, 0-1-4-6. Paratype measurement given only when holotype measurement different from <i>E. basiatus.</i> Leg I. Femur I, <i>d</i> minute, <i>l’</i> 3, <i>v″</i> 8 (7–10); genu I, <i>l′</i> 3, <i>l″</i> 3; tibia I, <i>d</i> 60, <i>l’</i> 10 (8–10), <i>l’’</i> 9 (8–9), <i>v’</i> 5 (4–5), <i>v’’</i> 11 (10–17), <i>k</i> 5, <i>φ</i> 9; tarsus I, <i>tc’</i> 15, <i>tc’’</i> 13 (13–15), <i>pl’</i> 9, <i>pl’’</i> 16 (15–18), <i>pv′</i> 2 (2), <i>pv’’</i> 2, <i>ω</i> 4, <i>s</i> 6 (5), <i>u′′</i> m. Leg II. Genu II, <i>l′</i> 4; tibia II, <i>d</i> 48 (48–55), <i>l’</i> 7, <i>v’</i> 17 (14–15), <i>v’’</i> 17 (20–27); tarsus II, <i>tc′</i> 2, <i>tc</i> ″ 4 (5), <i>pl</i> ″ 59 (50), <i>pv’’</i> 2, <i>u’</i> 6, <i>u′′</i> minute. Leg III. Genu III, <i>l′</i> 3; tibia III, <i>d</i> 55 (40–49), <i>l’</i> 7, <i>v’</i> 16, <i>v’’</i> 17 (22–29); tarsus III, <i>tc’</i> 2, <i>tc</i> ″ 5 (4–5), <i>pl</i> ″ 39 (37–40), <i>pv’’</i> 2, <i>u’</i> 7 (6–7), <i>u′′</i> minute.</p> <p>LARVIFORM ADULT MALE (n = 4).</p> <p> <i>Gnathosoma</i>. Length 31–39, width 28–36, apex indented, terminating in small but distinct lobes. Cheliceral stylets 20–22, setae <i>ch</i> 6–7, <i>su</i> 6–7.</p> <p> <i>Idiosoma</i>. Length 150–180, width 125–150. Prodorsal plate with setae <i>v1</i> 2–3, <i>v2</i> alveolar remnant, <i>sc1</i> 3–5, <i>sc2</i> 60–70. Distance between setae <i>v1–v1</i> 30–36, <i>sc1–sc1</i> 43–50, <i>v1–sc1</i> 15–16, <i>v2–v2</i> 34–41, <i>sc2–sc2</i> 63–77, <i>sc1–sc2</i> 19–25. Plate CD setae <i>c1</i> 4–5, <i>c2</i> 5–6, <i>d</i> 5–6, distance between setae <i>c1–c1</i> 46–54, <i>c1–c2</i> 32–38, <i>d–d</i> 40–46; cupuli <i>ia</i> anterolaterad setae <i>d</i>. Plate EF eroded medially; with setae <i>f</i> 4–5, distance <i>f–f</i> 25–29; cupuli <i>im</i> anterolaterad setae <i>f</i>. Genital capsule length 27–28, width 38–40, setae <i>h</i> minute. Venter: coxal setae <i>1a</i> 2–3, <i>2a</i> 3–4, <i>3a</i> 2–3, <i>3b</i> 4–5.</p> <p> <i>Legs</i>. Setal counts legs I–III, femur-tarsus: 3-2-6(+ <i>φ</i>)-8(+ <i>ω</i>), 0-1-4-6, 0-1-4-6. Leg I. Femur I, <i>d</i> minute, <i>l′</i> minute, <i>v″</i> 4–6; genu I, <i>l′</i> minute, <i>l″</i> minute; tibia I, <i>d</i> 42–52, <i>l’</i> 2–3, <i>l’’</i> 3–4, <i>v</i> ’ 3, <i>v</i> ’’ 10, <i>k</i> 3, <i>φ</i> 8; tarsus I, <i>tc’</i> 12–14, <i>tc’’</i> 12–14, <i>pl’</i> 9, <i>pl’’</i> 10–14, <i>pv</i> ′ m–2, <i>pv’’</i> m–2, <i>ω</i> 3–4, <i>s</i> 4–5, <i>u</i> ′′ minute. Leg II. Genu II, <i>l′</i> minute; tibia II, <i>d</i> 26–30, <i>l’</i> 5, <i>v’</i> 11, <i>v’’</i> 18–21; tarsus II, <i>tc′</i> 2, <i>tc</i> ″ 3–4, <i>pl</i> ″ 25–28, <i>pv’’</i> minute, <i>u’</i> 4, <i>u″</i> minute. Leg III. Genu III, <i>l′</i> minute; tibia III, <i>d</i> 26–28, <i>l’</i> 4–6, <i>v’</i> 8–12, <i>v’’</i> 16; tarsus III, <i>tc′</i> 2, <i>tc</i> ″ 4, <i>pl</i> ″ 29–31, <i>pv’’</i> 2, <i>u’</i> 4, <i>u″</i> minute.</p> <p>LARVAL FEMALE (n = 11).</p> <p> <i>Gnathosoma.</i> Length 38–41, width 33–36, apex indented, terminating in small but distinct lobes. Cheliceral stylets 30–35, setae <i>ch</i> 22–27, <i>su</i> 6–7.</p> <p> <i>Idiosoma</i>. Length 185–260, width 140–220. Prodorsal plate with setae <i>v1</i> 4–5, <i>v2</i> alveolar remnant, <i>sc1</i> 5–6, <i>sc2</i> 105–110. Distance between setae <i>v1–v1</i> 32–35, <i>sc1–sc1</i> 59–65, <i>v1–sc1</i> 22–24, <i>v2–v2</i> 44–46, <i>sc2–sc2</i> 78–100, <i>sc1–sc2</i> 22–34. Plate C setae <i>c1</i> 6–9, <i>c2</i> 9–10, distance between setae <i>c1–c1</i> 50–71. Plate D setae <i>d</i> 7–10, <i>d–d</i> 54–72, cupuli <i>ia</i> anteriad to anteromesad <i>d</i>. Plate EF setae <i>f</i> 7–10, <i>f–f</i> 47–59, cupuli <i>im</i> anteriad to anterolaterad <i>f</i>. Plate H narrow, large, 18–21 wide, 31–36 long, bearing setae <i>h1</i> ca. 160, <i>h2</i> 6–8. Venter: coxisterna 1-2 moderately well defined, with some folds, coxisterna 3 membranous; coxal setae slender, <i>1a</i> 3, <i>2a</i> 4, <i>3a</i> 2–4, <i>3b</i> 5 (one specimen lacking setae <i>3a</i> and <i>3b</i> expressed as alveoli). Alveoli of <i>1b</i>, <i>2b</i> present.</p> <p> <i>Legs.</i> Setal counts legs I–III, femur-tarsus: 3-2-6(+ <i>φ</i>)-8(+ <i>ω</i>), 0-1-4-6, 0-1-4-6. Ranges given only when first measurement different from <i>E. basiatus.</i> Leg I. Femur I, <i>d</i> minute, <i>l′</i> minute, <i>v″</i> 4–6; genu I, <i>l′</i> m–2, <i>l″</i> 2; tibia I, <i>d</i> 55–65, <i>l’</i> 7–11, <i>l’’</i> 8–11, <i>v’</i> 4–5, <i>v’’</i> 8–10, <i>k</i> 4, <i>φ</i> 10; tarsus I, <i>tc’</i> 15, <i>tc’’</i> 15, <i>pl’</i> 6–7, <i>pl’’</i> 14–17, <i>pv′</i> minute, <i>pv’’</i> minute, <i>ω</i> 4, <i>s</i> 5, <i>u′′</i> minute. Leg II. Genu II, <i>l′</i> m–2; tibia II, <i>d</i> 40, <i>l’</i> 6, <i>v’</i> 10–14, <i>v’’</i> 21; tarsus II, <i>tc′</i> minute, <i>tc</i> ″ 3, <i>pl</i> ″ 28, <i>pv’’</i> minute, <i>u’</i> 5, <i>u′′</i> minute. Leg III. Genu III, <i>l′</i> 3; tibia III, <i>d</i> 36, <i>l’</i> 7, <i>v’</i> 10–15, <i>v’’</i> 21; tarsus III, <i>tc’</i> m–2, <i>tc</i> ″ 3, <i>pl</i> ″ 38–43, <i>pv’’</i> minute, <i>u’</i> 5, <i>u′′</i> minute.</p> <p> <b>Differential diagnosis.</b> <i>Eutarsopolipus savatus</i> <b>sp. nov.</b> is most similar to <i>E. labiatus</i> and the distinguishing features are discussed under the <i>Differential Diagnosis</i> for the latter species.</p> <p> <b>Etymology.</b> The name <i>savatus</i> (a tender kiss) alludes to both the lip-like protrusion of the gnathosoma but also alludes to the larger size of this species.</p> <p> <b>Remarks.</b> One larva similar to <i>E. echinatus</i> Seeman, 2019a was found on HR# T82502; one male closest to the <i>nahmani</i> subgroup group was found on HR# T82502 (QMS 113499) (Tables 1, 2).</p> <p> <b> The <i>leytei</i> species group</b> </p>Published as part of <i>Seeman, Owen D., 2021, Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae), pp. 1-74 in Zootaxa 4971 (1)</i> on pages 56-59, DOI: 10.11646/zootaxa.4971.1.1, <a href="http://zenodo.org/record/4771514">http://zenodo.org/record/4771514</a>
Eutarsopolipus teuceri Seeman 2021, sp. nov.
No full textEutarsopolipus teucerisp. nov. (Figs 21–23) Diagnosis. Adult female: Gnathosomal length 58–61, width 51–54; anterior margin of prodorsal shield slightly eroded medially; v1 8–10, sc1 18–20, c1 17–22, c2 15–16, d 18–19, f 15–18, h 7–10, v1-v1 42–48, d-d 130–140, f-f 125–130; genua II-III setae l′ 2–4, tibia I seta l′ 6–7, tarsi II-III setae tc′ 3–4, tc′′ 5–6, pv′′ large, 4. Larval female: Unknown. Material examined. Holotype female, ex Castelnaudia porphyriaca. 1.5 km from Mt Cabinet, via Jimna, Sunday Ck, 26° 42ʹ 39ʺ S 152° 33ʹ 49ʺ, 9 Sep-30 Nov 1974, G. & S. Monteith, pitfall trap, IN5370, HR# T 57070 (QMS 113506). Paratypes. 2 females, 3 males, ex C. porphyriaca, Conondale Range, 27 Nov. 1974, G. Monteith, unregistered specimen (QMS 113507, 113510-12, ANIC 52-003962). All in QM except one female in ANIC. Description. FEMALE (Figs 21–22; n = 3). Gnathosoma. Length 58 (60–61), width 54 (51–54). Cheliceral stylets 77 (75–82), setae ch 26 (20–27), su 8 (10). Idiosoma (Fig. 21). Length 360 (310–325), width 250 (225–250). Prodorsal plate not eroded anteriorly; setae slender, v1 10 (8–9), v2 alveolar remnant, sc1 20 (18–20), sc2 broken (85–105). Distance between setae v1–v1 48 (42–44), sc1–sc1 92 (80–86), v1–sc1 31 (26–30), v2–v2 48 (41–43), sc2–sc2 105 (100–105), sc1–sc2 28 (23–29). Opisthosomal setae slender. Plate C setae c1 17 (17–22), c2 17 (15–16), distance between setae c1–c1 80 (70–75). Plate D setae d 19 (18–19), d–d 140 (130–135), cupuli ia anterolaterad d. Plate EF setae f 18 (15–16), f–f 130 (125), cupuli im anterolaterad f. Plate H divided or fused weakly, seta h 7 (7–10). Venter: coxisterna 1-3 moderately well defined, with few folds, coxisterna 3 weaker than 1-2; coxal setae slender, 1a 4 (3–4), 2a 5 (5–6), 3a 4 (5), 3b 7 (5–6). Alveoli of 1b, 2b present. Legs (Fig. 22). Setal counts legs I–III, femur-tarsus: 3-2-6(+ φ)-8(+ ω), 0-1-4-6, 0-1-4-6. Leg I. Femur I, d minute, l’ m (2), v″ 8 (7-9); genu I, l′ 2 (2–3), l″ 2 (2–3); tibia I, d broken (60–68), l’ 7 (5–6), l’’ 6 (8), v’ 4 (4–5), v’’ 9 (8–10), k 4 (3–4), φ 9 (8); tarsus I, tc’ 15 (13–14), tc’’ 14 (15), pl’ 8 (11–12), pl’’ broken (9–15), pv′ 3 (3–4), pv’’ 3 (4), ω 2 (3), s 6 (6–7), u′′ m (m). Leg II. Genu II, l′ 2 (3–4); tibia II, d 40 (35–43), l’ 6 (7), v’ 10 (10–14), v’’ 17 (18–20); tarsus II, tc′ 3 (3–4), tc ″ 5 (5–6), pl ″ 50 (41–43), pv’’ 4 (4), u’ 6 (7), u′′ m (m). Leg III. Genu III, l′ 2 (3); tibia III, d obscured (32–35), l’ 6 (5–6), v’ obscured (11–13), v’’ obscured (14–15); tarsus III, tc’ 4 (3), tc ″ 5 (5), pl ″ broken (38–42), pv’’ 4 (4), u’ 7 (6–7), u′′ m (m). LARVIFORM ADULT MALE (Fig. 23; n = 3). Gnathosoma. Length 32–34, width 21–25. Cheliceral stylets 21–25, setae ch 5–6, su 5–6. Idiosoma. Length 155–165, width 110–125. Prodorsal plate with setae v1 3–4, v2 alveolar remnant, sc1 5–6, sc2 62–70. Distance between setae v1–v1 16–19, sc1–sc1 41–42, v1–sc1 15–19, v2–v2 18–21, sc2–sc2 49–50, sc1–sc2 21–25. Plate CD setae c1 6–7, c2 5–6, d 6–7, distance between setae c1–c1 42–45, c1–c2 24–29, d–d 35–37; cupuli ia anterolaterad setae d. Plate EF slightly eroded medially; with setae f 2–4 (thickened in one specimen), distance f–f 22–24; cupuli im anterolaterad setae f. Genital capsule length 26–29, width 30–32, setae h minute. Venter: coxal setae 1a 2–3, 2a 3–4, 3a 3–4, 3b 4–5. Legs. Setal counts legs I–III, femur-tarsus: 3-2-6(+ φ)-8(+ ω), 0-1-4-6, 0-1-4-6. Leg I. Femur I, d minute, l′ minute, v″ 3–4; genu I l′ 2, l″ m–2; tibia I, d 32–37, l’ 3–4, l’’ 3–4, v ’ 2–3, v ’’ 6–10, k 2–3, φ 6–7; tarsus I, tc’ 13, tc’’ 13–14, pl’ 6–7, pl’’ 9–12, pv ′ m, pv’’ 2, ω 2, s 4–5, u ′′ m. Leg II. Genu II, l′ 2; tibia II, d 16–17, l’ 5–6, v’ 9–12, v’’ 10–15; tarsus II, tc′ 2, tc ″ 4, pl ″ 27–33, pv″ 2–3, u’ 4–5, u″ minute. Leg III. Genu III, l′ 2–3; tibia III, d 12–16, l’ 4, v’ 8–10, v″ 12–15; tarsus III, tc′ 2, tc ″ 4–5, pl ″ 26–32, pv″ 3, u ’ 5–6, u″ minute. LARVAL FEMALE. Unknown. Differential diagnosis. Eutarsopolipus teuceri sp. nov. is close to E. raveni. Females are most easily separated by the length of seta h, which is short (2–4) in E. raveni but longer (7–10) in E. teuceri. Other small differences are the size of setae tc′ (3–4 versus 2) and pv′ (4 versus 2–3) on tarsus III, which are larger in E. teuceri, and the longer cheliceral stylets (75–82 versus 68–72). Males are almost identical, differing only by the length of setae sc2 (45–50 vs 62–70 in E. raveni), although the smaller tc′ is also apparent (2 versus minute in E. raveni). Eutarsopolipus raveni and E. teuceri both share a small gnathosoma with E. rutherfordae, which is also a member of the nahmani subgroup but its host is Trichosternus subvirens. These two species are most easily distinguished from E. rutherfordae by the strong, stiff setae d and f in E. rutherfordae (very fine, not stiff in E. raveni and E. teuceri). Eutarsopolipus raveni also has small setae h (2–4 compared to 7–10 in E. rutherfordae and E. teuceri), while E. teuceri has longer cheliceral stylets (75–82 versus 60), shorter cheliceral setae (20–27 versus 42), shorter seta v′′ on femur II (7–9 versus 16) and much larger setae pv′′ on tarsi II-III (4 versus 2). Etymology. The specific name teuceri refers to the legendry archer Teucer of the Odyssey, who in battle hid behind the glittering shield of Ajax, alluding to the spectacular metallic purple sheen of the elytra of the host beetle E. porphyriaca. Remarks. Castelnaudia porphyriaca hosted three species described herein, but also had one larva of another species collected at Conondale. The larva belongs to the pterostichi species group and has an enlarged tibia I seta v′, the only femoral seta is d, and the only setae on all genua is on leg I (seta l′). The ochoai species group - despoticus subgroup Diagnosis. Female with dorsal idiosomal setae fine, slender; setae h minute. All life stages with gnathosomal apex evenly rounded.Published as part of Seeman, Owen D., 2021, Contrasting species diversification of Eutarsopolipus (Acariformes: Podapolipidae) on Castelnaudia and Notonomus (Coleoptera: Carabidae), pp. 1-74 in Zootaxa 4971 (1) on pages 34-37, DOI: 10.11646/zootaxa.4971.1.1, http://zenodo.org/record/477151
On a Regev-Seeman conjecture about Z_2-graded tensor products
No full textIn the Theory of Polynomial Identities of algebras, superalgebras play a key role, as emphasized by the celebrated Kemer’s results on the structure of T -ideals of the free associative algebra. Kemer succeeded in classifying the T –prime algebras over a field of characteristic zero, and all of them possess a natural superalgebra structure. In a celebrated work Regev proved that the tensor product of PI-algebras is again a PI-algebra, and the so-called Kemer’s
Tensor Product Theorem shows that the tensor product of T -prime algebras is again PI-equivalent to a T -prime algebra, explicitly described.
When dealing with superalgebras, however, it is possible to define an alternative tensor product, sometimes called super, or graded, or signed tensor product. In a recent paper Regev and Seeman studied graded tensor products, and they proved that the graded tensor product of PI-algebras is again PI, as for the ordinary case. Then natural questions
arise: is the graded tensor product of T -prime algebras again T -prime? If so, do the graded and ordinary tensor products of T -prime algebras give the “same results” up to PI-equivalence? Among their results, Regev and Seeman noticed cases for which a graded version of Kemer’s Tensor Product Theorem does hold. More precisely, the re-
sulting algebra is still a T -prime algebra, possibly “different” from the “natural” one. Then they conjectured this should be true in general.
The present paper positively solves the conjecture. More precisely, we can prove that, in zero characteristic, the graded tensor product of T -prime algebras “is” again T -prime, and we describe the resulting algebra up to PI-equivalence
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