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    FIGURE 8 in Pelagia benovici sp. nov. (Cnidaria, Scyphozoa): a new jellyfish in the Mediterranean Sea

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    FIGURE 8. Appearance (structure, colour) of oral arms and tentacles in (A) Pelagia noctiluca and (B) Pelagia benovici sp. nov.Published as part of Piraino, Stefano, Aglieri, Giorgio, Martell, Luis, Mazzoldi, Carlotta, Melli, Valentina, Milisenda, Giacomo, Scorrano, Simonetta & Boero, Ferdinando, 2014, Pelagia benovici sp. nov. (Cnidaria, Scyphozoa): a new jellyfish in the Mediterranean Sea, pp. 455-468 in Zootaxa 3794 (3) on page 465, DOI: 10.11646/zootaxa.3794.3.7, http://zenodo.org/record/491497

    FIGURE 8 in Pelagia benovici sp. nov. (Cnidaria, Scyphozoa): a new jellyfish in the Mediterranean Sea

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    FIGURE 8. Appearance (structure, colour) of oral arms and tentacles in (A) Pelagia noctiluca and (B) Pelagia benovici sp. nov.Published as part of Piraino, Stefano, Aglieri, Giorgio, Martell, Luis, Mazzoldi, Carlotta, Melli, Valentina, Milisenda, Giacomo, Scorrano, Simonetta & Boero, Ferdinando, 2014, Pelagia benovici sp. nov. (Cnidaria, Scyphozoa): a new jellyfish in the Mediterranean Sea, pp. 455-468 in Zootaxa 3794 (3) on page 465, DOI: 10.11646/zootaxa.3794.3.7, http://zenodo.org/record/491497

    Ulmaridae HAECKEL 1880

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    FAMILY ULMARIDAE HAECKEL, 1880 Semaeostomeae with simple or branched radial canals and a ring canal; with or without subgenital pits.Published as part of Scorrano, Simonetta, Aglieri, Giorgio, Boero, Ferdinando, Dawson, Michael N. & Piraino, Stefano, 2017, Unmasking Aurelia species in the Mediterranean Sea: an integrative morphometric and molecular approach, pp. 243-267 in Zoological Journal of the Linnean Society 180 (5) on page 254, DOI: 10.1111/zoj.12494, http://zenodo.org/record/571098

    Figure 6 in Unmasking Aurelia species in the Mediterranean Sea: an integrative morphometric and molecular approach

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    Figure 6. Ontogenetic variation in subgenital pore size (f11) and position (f12) in male specimens of Aurelia coerulea from: A, Empuriabrava Harbour (EH), bell diameter (BD) = 63 mm; B, Varano Lagoon (VL), BD = 120 mm.Published as part of Scorrano, Simonetta, Aglieri, Giorgio, Boero, Ferdinando, Dawson, Michael N. & Piraino, Stefano, 2017, Unmasking Aurelia species in the Mediterranean Sea: an integrative morphometric and molecular approach, pp. 243-267 in Zoological Journal of the Linnean Society 180 (5) on page 254, DOI: 10.1111/zoj.12494, http://zenodo.org/record/571098

    Aurelia LAMARCK 1816

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    GENUS AURELIA LAMARCK, 1816 Diagnosis: Ulmaridae with small marginal tentacles, and lappet-like structures arising from exumbrella slightly above umbrella margin. Umbrella margin divided by eight or 16 marginal clefts; four unbranched oral arms; anastomoses between a few or all of the radial canal branches. Type species: Aurelia aurita (Linnaeus, 1758).Published as part of Scorrano, Simonetta, Aglieri, Giorgio, Boero, Ferdinando, Dawson, Michael N. & Piraino, Stefano, 2017, Unmasking Aurelia species in the Mediterranean Sea: an integrative morphometric and molecular approach, pp. 243-267 in Zoological Journal of the Linnean Society 180 (5) on page 255, DOI: 10.1111/zoj.12494, http://zenodo.org/record/571098

    FIGURE 2 in Pelagia benovici sp. nov. (Cnidaria, Scyphozoa): a new jellyfish in the Mediterranean Sea

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    FIGURE 2. Pelagia benovici sp. nov. (A) Lateral view. (B) Sublateral view. (C) Aboral view, with prominent exumbrellar cnidocyst warts. (D) Oral view, showing the distinctive horse-shoe shaped white gonads, and overall subumbrellar morphology. rh: rhopalia, rs: radial septa (for clarity, only few labels added).Published as part of Piraino, Stefano, Aglieri, Giorgio, Martell, Luis, Mazzoldi, Carlotta, Melli, Valentina, Milisenda, Giacomo, Scorrano, Simonetta & Boero, Ferdinando, 2014, Pelagia benovici sp. nov. (Cnidaria, Scyphozoa): a new jellyfish in the Mediterranean Sea, pp. 455-468 in Zootaxa 3794 (3) on page 459, DOI: 10.11646/zootaxa.3794.3.7, http://zenodo.org/record/491497

    Pelagiidae Gegenbaur 1856

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    Family Pelagiidae Gegenbaur 1856 Diagnosis of the genus Pelagia Péron & Lesueur 1809 (redefined, after Gershwin & Collins 2002) Pelagiidae with exumbrella covered by conspicuous cnidocyst warts; eight marginal tentacles alternating with eight marginal sense-organs located on shallow sensory pits; 16 marginal lappets; 16 unbranched, simple radial septa terminating between sense organs and tentacles, dividing the gastrovascular sinus into 8 tentacular and 8 rhopalial separate pouches that do not communicate with neighbouring pouches at the umbrella margin. Direct life cycle without polyp stage. Type species: Pelagia noctiluca (Forskål 1775)Published as part of Piraino, Stefano, Aglieri, Giorgio, Martell, Luis, Mazzoldi, Carlotta, Melli, Valentina, Milisenda, Giacomo, Scorrano, Simonetta & Boero, Ferdinando, 2014, Pelagia benovici sp. nov. (Cnidaria, Scyphozoa): a new jellyfish in the Mediterranean Sea, pp. 455-468 in Zootaxa 3794 (3) on page 458, DOI: 10.11646/zootaxa.3794.3.7, http://zenodo.org/record/491497

    FIGURE 5 in Pelagia benovici sp. nov. (Cnidaria, Scyphozoa): a new jellyfish in the Mediterranean Sea

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    FIGURE 5. Pelagia benovici sp. nov. (A–B) Cnidocyst warts (or cnidocyst clubs) of different sizes scattered over the subumbrellar gonads. Oocytes (oo) of different sizes and maturity are shown. (C) Discharged microbasic eurytele (eu). (D) Discharged holotrichous isorhiza (is) and microbasic eurytele (eu). (E) Discharged holotrichous isorhiza (o-is). (F) Microbasic eurytele (eu) and third type of cnidocyst (birh), provisionally identified as heterotrichous microbasic birhopaloid II type (sensu Ostman 2000).Published as part of Piraino, Stefano, Aglieri, Giorgio, Martell, Luis, Mazzoldi, Carlotta, Melli, Valentina, Milisenda, Giacomo, Scorrano, Simonetta & Boero, Ferdinando, 2014, Pelagia benovici sp. nov. (Cnidaria, Scyphozoa): a new jellyfish in the Mediterranean Sea, pp. 455-468 in Zootaxa 3794 (3) on page 462, DOI: 10.11646/zootaxa.3794.3.7, http://zenodo.org/record/491497

    FIGURE 4 in Pelagia benovici sp. nov. (Cnidaria, Scyphozoa): a new jellyfish in the Mediterranean Sea

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    FIGURE 4. Pelagia benovici sp. nov. (A) Rhopaliar pit (rp). (B) Portion of gastrovascular sinus showing radial septa (rs), subumbrellar perradial warts (sw), tentacular (tp) and rhopaliar (rhp) pouches, rhopalia (rh), and rounded marginal lappets. (C) Gastric filaments (gf), horse-shoe shaped gonads (g), and oral arm (oa) covered by transparent warts. (D) Enlargement of ribbon-like gonad (g) covered by cnidocyst warts (nw) bordering the stomach wall with emergent gastric filaments (gf).Published as part of Piraino, Stefano, Aglieri, Giorgio, Martell, Luis, Mazzoldi, Carlotta, Melli, Valentina, Milisenda, Giacomo, Scorrano, Simonetta & Boero, Ferdinando, 2014, Pelagia benovici sp. nov. (Cnidaria, Scyphozoa): a new jellyfish in the Mediterranean Sea, pp. 455-468 in Zootaxa 3794 (3) on page 461, DOI: 10.11646/zootaxa.3794.3.7, http://zenodo.org/record/491497

    Unmasking Aurelia species in the Mediterranean Sea: an integrative morphometric and molecular approach

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    Molecular analyses have led to an increased knowledge of the number and distribution of morphologically cryptic species in the world's oceans and, concomitantly, to the identification of non-indigenous species (NIS). Traditional taxonomy and accurate delimitation of species’ life histories and autecology lag far behind, however, even for the most widely distributed taxa, such as the moon jellyfish Aurelia (Cnidaria, Scyphozoa) species complex. Here we analysed mitochondrial cytochrome c oxidase subunit I (COI) and nuclear 28S ribosomal RNA (28S) gene sequences to assign polyps, ephyrae, and medusae collected in the Mediterranean Sea to different phylogenetic species. We find evidence for three Aurelia species, none of which are referable to the type species of the genus, Aurelia aurita (Linnaeus, 1758), and describe the anatomical, morphometric, and developmental variation within and between them. We identify Aurelia coerulea von Lendenfeld, 1884 and Aurelia solida Browne, 1905 as established non-indigenous species in the Mediterranean Sea. We describe Aurelia relicta sp. nov., an endemic species currently unique to a population in the marine lake of Mljet (Croatia). These results demonstrate the usefulness of integrative approaches in resolving taxonomic uncertainty surrounding cryptic species complexes, identifying patterns of marine biodiversity, and recognizing non-indigenous species in marine ecosystems
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