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Pauropus dicastri Scheller 1968
<i>Pauropus dicastri</i> Scheller, 1968 <p>Chile, Osorno Prov., Parque Nacional Puyehue, moist rtemperate forest, 1 ad. 9 (♀), September 23, 1965, F. di Castri leg.</p>Published as part of <i>Scheller, Ulf, 2014, New records of Pauropoda (Myriapoda) with descriptions of new taxa, pp. 301-332 in Zootaxa 3866 (3)</i> on page 308, DOI: 10.11646/zootaxa.3866.3.1, <a href="http://zenodo.org/record/227447">http://zenodo.org/record/227447</a>
Stylopauropus plicatus Scheller, 2011, n. sp.
<i>Stylopauropus plicatus</i> n. sp. <p>Figs 12–20</p> <p> <b>Type locality.</b> U.S.A., Tennessee, Blount Co., Great Smoky Mountains National Park, Reed Creek.</p> <p> <b>Type specimens.</b> Holotype: ad. 9 (Ƥ), 27 June 1997, leg. P.J. Obenauer. Paratypes: 1 ad. 9(3), 1 subad. 8(Ƥ), 1 juv. 5, Tennessee, Blount Co., Great Smoky Mountains National Park, Cades Cove, Parsons Branch Road, 5 minutes from Jet Force Creek Road, hemlock log litter, 35º 37.75N, 83º 51.62W, 28 July 2004, leg. S.A. Gil & J. Hilten; 1 ad. 9(Ƥ), North Carolina, Swain Co., Lakeshore Trail, Pilky Creek, UTM 0 2 57 998E, 39 25 761N, 14 April 2004, leg. A.J. Mayor.</p> <p>Nontypes: Tennessee, Blount Co., lower Gregory Ridge trail, 1 mile from trail-head, litter, Berlese extraction, 1 subad. 8(Ƥ), 28 July 2004, leg. A.K. Tishechkin & B. Blitz; Cooper Road Trail, 0.8 miles from trail-head, elev. 1211 ft, 35º 37.021N, 83º 55.608W, 1 subad. (Ƥ), 31 July 2004, leg. J. Ceigier & S.A. Gil. Sevier Co., Cades Cove, Crooked Arm Ridge Trail, 35º 36.659N, 83º 46.796W, conifer cones, Berlese extraction, 2 subad. 8(Ƥ), 31 July 2004, leg. B. Blitz & V.M. Bayless. – 9 specimens.</p> <p> <b>Diagnosis</b>. <i>S. plicatus</i> <b>n. sp.</b> belongs to a group of Nearctic species related by similarities in the general shape of the anal plate, subsimilar shape and length of the pygidial setae <i>a</i> 1, <i>a</i> 2 and <i>a</i> 3 and antennal globulus with a strikingly high number of bracts: <i>S. californianus</i> Remy, <i>S. fratuelis</i> Remy, <i>S.</i> canadensis Scheller and <i>S. boreus</i> Scheller. The first is widely distributed, known from NE Siberia (Scheller 1986), California (Remy 1958), British Columbia (Scheller 1984) and the southern Appalachians. The other three have, as far as we know now, restricted ranges. Thus <i>S. boreus</i> Scheller is known only from Alaska (Scheller 1986), <i>S. fratuelis</i> Remy only from Tennessee (Remy 1958) and <i>S. canadensis</i> Scheller only from British Columbia (Scheller 1984). The new species can be distinguished from these species particularly by the shape of the styli of the pygidial tergum (thin and pointed, not more or less thick and blunt) and by the shape of the anal plate (posteriorly directed submedian branches parallel and very close to each other, not diverging). Similarities can be traced in the direction of the two Alaskan species <i>S. longipes</i> Scheller and <i>S. longitarsus</i> Scheller (Scheller 1986) but to a lesser degree.</p> <p> <b>Derivation</b> of name. From the Latin <i>plicatus</i> = fold (referring to the longitudinal folds of the anal plate). <b>Description.</b> <i>Length</i>. (0.89–) 0.99 mm.</p> <p> <i>Head</i> (Fig. 12). Tergal setae pubescent, median and submedian setae clavate, lateral and sublateral ones cylindrical, blunt, <i>a</i> 3 in 2nd row and <i>l</i> -group setae tapering pointed. Relative lengths of setae (holotype only), 1st row: <i>a</i> 1=10; <i>a</i> 2=11; 2nd row: <i>a</i> 1=11, <i>a</i> 2=12, <i>a</i> 3=18; 3rd row: <i>a</i> 1= <i>a</i> 2=10; 4th row: <i>a</i> 1=12, <i>a</i> 2=16, <i>a</i> 3=18, <i>a</i> 4 = 10; <i>l</i> 1= <i>l</i> 3=15, <i>l</i> 2=?. The ratio <i>a</i> 1 <i>/a</i> 1 <i>-a</i> 1 in 1st row 1.4, 2nd row 0.6, 3rd row 0.7, 4th row 1.5. Temporal organs mainly lateral, their length in tergal view 0.7 of shortest interdistance. Head cuticle glabrous.</p> <p> <i>Antennae</i> (Fig. 13). Segment 3 with 3 setae and rudimentary, almost cylindrical, globulus g´on distal part of tergal side. Segment 4 with at least 3 cylindrical striate setae, <i>r</i> thinnest. Relative lengths of setae (holotype only): <i>p</i> = 100, <i>p</i> ' = 73, <i>r</i> = 15. Tergal seta <i>p</i> 1.1 times as long as tergal branch <i>t</i>. The latter cylindrical, (4.5)4.9 times as long as its greatest diameter and as long as sternal branch <i>s</i> which is 2.3 times as long as its greatest diameter and with its posterodistal corner distinctly truncate. Seta <i>q</i> as tergal-anterior setae of 4th segment, 1.1 times as long as <i>s</i>. Relative lengths of flagella (with base segments included) and base segments: <i>F</i> 1=100, <i>bs</i> 1=6, <i>F</i> 2=90, <i>bs</i> 2=6, <i>F</i> 3=50, <i>bs</i> 3=5. <i>F</i> 1 2.7(2.9) times as long as <i>t</i>, <i>F</i> 2 and <i>F</i> 3 2.5 and 1.4 times as long as <i>s</i> respectively. Distal calyces of <i>F</i> 1 conical, longer than wide, glabrous, other calyces not studied. Globulus <i>g</i> 1.8 times as long as its greatest diameter; (16)19 bracts; capsule spherical. Diameter of <i>g</i> 2.7 times as long as greatest diameter of <i>t</i>. Antennae glabrous.</p> <p> <i>Trunk</i> (Figs 14, 15). Setae of collum segment (Fig. 14) leaf-shaped, blunt, shortly pubescent, furcate with rudimentary secondary branch, the latter blunt glabrous. Sternite process triangular, anterior part incised; appendages broadly conical with thick caps. Pubescence faint on appendages, distinct on anterior part of process.</p> <p> Setae on tergites only partly studied. 4+4 setae on tergite I, 6+6 setae on II–V, 4+2 on VI. Posterior setae on VI (Fig. 15) long tapering, 1.5 times as long as interdistance and 2.3 times as long as pygidial setae <i>a</i> 1. Tergites glabrous.</p> <p> <i>Genital papillae</i> (fig. 16). 1.7 and 2.0 times as long as greatest width, proximal half cylindrical, seta 0.4 of the length of papilla.</p> <p> <i>Bothriotricha</i>. Most bothriotricha lost or broken. Relative lengths: <i>T</i> 1 = 100, <i>T</i> 5 = 244(–324). Axes simple straight; pubescence consisting of simple straight hairs, oblique on most proximal parts, outwards erect.</p> <p> <i>Legs.</i> Setae on coxa (Fig. 17) and trochanter of legs 1–8 leaf-shaped blunt, very shortly pubescent, secondary branch rudimentary glabrous; these setae on leg 9 furcate with secondary branch clavate, shortly pubescent, 0.8 of the length of seta. Tarsus of leg 9 (Fig. 18) somewhat bow-shaped, tapering, (4.7)4.8 times as long as its greatest diameter. Proximal seta long tapering pointed, with short oblique pubescence, seta (0.6)0.7 of the length of tarsus and (4.2)4.3 times as long as distal seta, the latter cylindrical blunt striate. Cuticle of tarsus weakly pubescent.</p> <p> <i>Pygidium</i> (Figs 19, 20). <i>Tergum</i>. Posterior part broadly triangular, distinct semicircular posterior lobe between <i>st</i>. Setae thin with short oblique pubescence; their relative lengths: <i>a</i> 1=10, <i>a</i> 2=9(10), <i>a</i> 3=10(11), <i>st</i> =5; <i>a-</i> setae directed posteriorly, somewhat curved inwards, subcylindrical, in distal half tapering, <i>st</i> straight tapering pointed, distinctly converging. Distance <i>a</i> 1 <i>-a</i> 1 0.7 of interdistance, distance <i>a</i> 1 <i>-a</i> 2 1.7(2.0) times as long as distance <i>a</i> 2- <i>a</i> 3; distance <i>st-st</i> (1.7)1.8 times as long as <i>st</i> and (1.2)1.3 times as long as distance <i>a</i> 1 <i>-a</i> 1. Cuticle glabrous.</p> <p> <i>Sternum</i>. Setae <i>b</i> 1 on small lobes, posterior margin in between almost straight. Relative lengths of setae (pygidial <i>a</i> 1=10): <i>b</i> 1=20. <i>b</i> 1 subcylindrical blunt, with short oblique pubescence, 1.3 times as long as interdistance.</p> <p> <i>Anal plate</i> narrowest anteriorly, as broad as long, lateral margins convex, posterior corners lengthened into short tapering, posteriorly directed, points; posterior margin with two submedian subcircular indentations and between them two long appendages protruding backwards close to each other; appendages as long as plate, cylindrical tapering. Plate and appendages glabrous.</p>Published as part of <i>Scheller, Ulf, 2011, Pauropoda (Myriapoda) from Great Smoky Mountains National Park, U. S. A., with descriptions of four new species, pp. 36-48 in Zootaxa 2962</i> on pages 40-42, DOI: <a href="http://zenodo.org/record/202418">10.5281/zenodo.202418</a>
Decapauropus arcuatilis Scheller, 2011, n. sp.
Decapauropus arcuatilis n. sp. Figs 1–11 Type locality. U.S.A., Tennessee, Great Smoky Mountains National Park, Sevier Co., Rainbow Falls. Type specimens. Holotype: ad. 9 (3), 4 May 1999, leg. E.C. Bernard & P.L. Jennings. — 1 specimen. Diagnosis. D. arcuatilis n. sp. is well defined by the shape of the anal plate, with long cylindrical appendages which also are curved inwards, the T-shaped st and the downwards directed appendages of the collum segment. Its relationships can not be traced at present. Derivation of name. From the Latin arcuatilis = shaped like a bow (referring to the appendages of the anal plate). Description. Length. 0.68 mm. Head (Fig. 1). Tergal setae striate, median and submedian setae somewhat clavate, lateral and sublateral ones cylindrical, a 3 in 2 nd row and l -group setae tapering, pointed. Relative lengths of setae, 1 st row: a 1 = 10; a 2 = 9; 2 nd row: a 1 = 18, a 2 = 19, a 3 = 16; 3 rd row: a 1 = 10, a 2 =?; 4 th row: a 1 = 14, a 2 = 21, a 3 = 13, a 4 = 17; l 1 = 14, l 2 = 18, l 3 =?. Ratio a 1 /a 1 -a 1 in 1 st row 0.5, 2 nd and 3 rd rows 0.8, 4 th row 1.0. Temporal organs large, their length in tergal view 2.2 times as long as shortest interdistance; small pistill in posterior part. Head cuticle glabrous. Antennae (Fig. 2). Segment 4 with at least 3 cylindrical striate blunt setae. Relative lengths of setae: p = 10, p ʹ= 15, p ʺ= 3. Tergal seta p as long as tergal branch t. The latter fusiform, distally cut obliquely, 2.5 times as long as its greatest diameter and as long as sternal branch s, the latter 2.1 times as long as its greatest diameter and with its anterodistal corner truncate. Seta q as p ' of 4 th segment, 0.7 of the length of s. Relative lengths of flagella (with base segments included) and base segments: F 1 = 100, bs 1 = 8, F 2 = 73, bs 2 = 5, F 3 = 75, bs 3 = 8. F 1 4.0 times as long as t, F 2 and F 3 2.9 and 3.0 times as long as s respectively, F 3 tapering. Distal calyces subhemispherical, largest on F 1, axis of flagella widened below calyx only in F 1. Globulus g subspherical, 9 bracts; capsule somewhat flattened. Diameter of g 0.9 of greatest diameter of t. Antennae glabrous. Trunk (Figs 3, 4). Setae on collum segment (Fig. 3) furcate, branches cylindrical blunt, main branch striate, secondary branch rudimentary glabrous, sublateral setae 2.6 times as long as submedian setae. Sternite processes very thin anteriorly; appendages directed downwards. Collum segment glabrous. Tergite I partly divided transversally, setae on tergites only partly studied. 4 + 4 setae on tergite I, 6 + 6 setae on II–IV, 4 + 2 on VI. Posterior setae on VI (Fig. 4) tapering, length 0.8 of interdistance and 1.7 times as long as pygidial setae a 1. Tergites glabrous. Bothriotricha (Figs 5, 6). Axes thin straight, thickest in the middle of T 3 (Fig. 6). Relative lengths: T 1 = 100, T 2 ≈ 100, T 3 = 118, T 4 =?, T 5 = 187. They have thin simple straight axes; pubescence consisting of simple straight hairs, oblique on most proximal parts, outwards erect, on distal 1 / 3 of T 1 (Fig. 5) and on T 2 branched erect hairs. Genital papillae (Fig. 7). Conical, twice longer than greatest diameter, base segment well developed, seta 0.3 of the length of papilla. Legs (Figs 8, 9). Legs short, setae on coxa and trochanter (Fig. 8) of leg 9 furcate, branches cylindrical blunt striate, secondary branch somewhat thinner than main branch. These setae on more anterior legs with rudimentary secondary branches. Tarsus of leg 9 (Fig. 9) straight tapering, 4.8 times as long as its greatest diameter. Setae striate, proximal seta tapering pointed, 0.3 of the length of tarsus and 2.2 times as long as cylindrical blunt distal seta. Cuticle of tarsus glabrous. Pygidium (Figs 10, 11). Tergum. Posterior margin almost straight but with low broad lobe behind st. a -setae thin cylindrical blunt, a 1 straight diverging, a 2 and a 3 somewhat curved inwards, converging, st in sternal/tergal view straight converging T-shaped glabrous, distal broad part bent downwards. Relative lengths of setae: a 1 = 10, a 2 = 14, a 3 = 21, st ≈ 7. Distance a 1 -a 1 1.4 times as long as interdistance, distance a 1 -a 2 almost as long as distance a 2 - a 3; distance st-st about 3 times longer than st and 0.7 of distance a 1 -a 1. Cuticle glabrous. Sternum. Posterior margin between b 1 with low bulge below anal plate. Setae tapering, striate distally, b 1 blunt, b 2 pointed, their relative lengths (pygidial a 1 = 10): b 1 = 51, b 2 = 13. b 1 1.5 times as long as interdistance, b 2 0.8 of distance b 1 - b 2. Anal plate subquadrate, lateral margins concave, posterior corners each with a long cylindrical blunt appendage, this curved inwards and somewhat longer than plate itself. Plate and appendages glabrous. FIGURES 1–11. Decapauropus arcuatilis n. sp., holotype (3). 1, head, submedian and right part, tergal view. 2, left antenna, sternal view. 3, collum segment, median and left part, sternal view. 4, tergite VI, right posterior corner and posteriomedian part. 5, T 1. 6, T 3. 7, genital papilla, lateral view. 8, seta on trochanter of leg 9. 9, tarsus of leg 9. 10, pygidium, posteriomedian and left posteror corner, sternal view. 11, anal plate, lateral view. Scale a: 5, 6, 9; b: 1, 4, 7, 8; c: 2, 3; d: 10, 11.Published as part of Scheller, Ulf, 2011, Pauropoda (Myriapoda) from Great Smoky Mountains National Park, U. S. A., with descriptions of four new species, pp. 36-48 in Zootaxa 2962 on pages 38-39, DOI: 10.5281/zenodo.20241
Donzelotauropus dividuus Scheller, 2011, n. sp.
<i>Donzelotauropus dividuus</i> n. sp. <p>Figs 21–28</p> <p> <b>Type locality.</b> U.S.A., Great Smoky Mountains National Park, North Carolina, Swain Co., Balsam Mountain Road, 4 km from south entrance, 3537,445N, 8310.761W, west-facing slope, deciduous forest with <i>Acer rubrum, Halesia carolina, Fagus grandiflora, Hamamelis virginiana,</i> under bark on dead tree, 24 September 2002, leg. U. Scheller, E.Bernard, I. Stocks.</p> <p> <b>Type material.</b> Holotype: ad. 9(3), data as above. Paratypes: 3 ad. 9(Ƥ), 2 subad. 8(Ƥ), 1 juv. 6, 1 juv. 5, 2 juv. 3, Tennessee, Sevier Co., above Laurel Falls, in soil under large poplars (<i>Lirodendron tulipifera</i>), 11 November 2004, leg. W. Reynolds & E. Gileh.— 11 specimens.</p> <p> <b>FIGURES 21–28.</b> <i>Donzelotauropus dividuus</i> <b>sp. n.</b>, holotype, 3. 21, head, submedian and right part, tergal view. 22, right antenna, tergal view. 23, collum segment, median and left part, sternal view. 24, genital papillae and seta on coxa of leg 2, anterior view. 25, seta on trochanter of leg 9. 26, tarsus of leg 9. 27, posterior part of tergite VI and median and right part of pygidial tergum, tergal view. 28, posterior part of pygidial sternum showing the placing of setae <i>b</i> 1 and <i>b</i> 3. Scale a: 24; b: 21, 25–27; c: 22, 23, 28.</p> <p> <b>Diagnosis</b>. Though this species has only one pair of setae on the pygidial sternum, <i>b</i> 1, and thus has to be placed in <i>Donzelotauropus</i>, it shows many similarities with a species in <i>Stylopauropus</i> earlier described from Great Smoky Mountains, <i>Stylopauropus sulcatoidus</i> Scheller (Scheller & Bernard, 2005). They resemble each other in the shape of the antennal globulus <i>g,</i> the bothriotricha, the general plan of the anal plate and the shape of the posterior part of the pygidial tergum and its setae. However, besides the presence of the <i>b</i> 1 <i>D. dividuus</i> is easily distinguished from <i>S. sulcatoidus</i> by the proportion <i>q/s</i>, 0.9, not 0.5 and the shape of the posterior part of the anal plate, incision deep U-shaped and appendages curved inwards and distinctly pubescent, not incision shallow, broadly Vshaped and appendages straight, faintly pubescent.</p> <p> <b>Derivation of name.</b> From the latin <i>dividuus</i> = divided into two parts (referring to the shape the anal plate).</p> <p> <b>Description.</b> <i>Length</i> (0.71–) 1.06 mm.</p> <p> <i>Head</i> (Fig. 21). Tergal setae mainly clavate blunt, with pubescence in whorls, <i>a</i> 3 in 2nd row and the <i>l</i> -group tapering, pointed pubescent, <i>a</i> 4 in 4th row cylindrical. Relative lengths of setae, 1st row: <i>a</i> 1=10; <i>a</i> 2=12; 2nd row: <i>a</i> 1=10(13), <i>a</i> 2=12(–14), <i>a</i> 3=15(–17); 3rd row: <i>a</i> 1=10(–11), <i>a</i> 2=?(–13); 4th row: <i>a</i> 1=(11–)16, <i>a</i> 2=(14–)20, <i>a</i> 3=15(–16), <i>a</i> 4=(10–)12; <i>l</i> 1=18(–21), <i>l</i> 3=17(–18), <i>l</i> 3=?(13). The ratio <i>a</i> 1 <i>/a</i> 1 <i>-a</i> 1 is in 1st row 1.1(–1.2), in 2nd row 0.5, in 3rd row 0.5(–0.6), in 4th row (0.9–)1.1. Temporal organs large, their length in tergal view 1.1(–1.2) times as long as shortest interdistance; pore with canal posteriorly. Head cuticle glabrous.</p> <p> <i>Antennae</i> (Fig. 22). Segment 3 with 2 subcylindrical setae and rudimentary <i>g'</i>, the latter oblique distally. Segment 4 with 5 cylindrical setae, all but <i>r</i> pointed, <i>u</i> rudimentary. Relative lengths of setae: <i>p</i> =100, <i>p</i> ΄=75(–87), <i>p"</i> =(45–)47, <i>r</i> =25(–31). Tergal seta <i>p</i> 0.7 of the length of tergal branch <i>t</i>. The latter fusiform, 3.7(–4.0) times as long as its greatest diameter and 1.2(–1.3) times as long as sternal branch <i>s,</i> the latter 2.1(–2.5) times as long as its greatest diameter and with its posterodistal corner distinctly truncate. Seta <i>q</i> as tergal-anterior setae of 4th segment, 0.9 of the length of <i>s</i>. Relative lengths of flagella (with base segments included) and of base segments alone: <i>F</i> 1=100, <i>bs</i> 1=17, <i>F</i> 2=78(80), <i>bs</i> 2=17, <i>F</i> 3≈100, <i>bs</i> 3=18(19). The <i>F</i> 1 (2.5–)2.7 times as long as <i>t</i>, <i>F</i> 2 and <i>F</i> 3 3.1 and (1.2–)1.4 times as long as <i>s</i> respectively. Distal calyces glabrous, on <i>F</i> 1 conical, longer than wide, other calyces with rounded tips. Globulus <i>g</i> with thick stalk, (1.9–)2.1 times as long as its greatest diameter; ≈10 bracts, these partly bifurcate; capsule subspherical. Diameter of <i>g</i> as long as greatest diameter of <i>t</i>. Antennae glabrous.</p> <p> <i>Trunk</i>. Setae of collum segment (Fig. 23) leaf-shaped blunt, indistinctly pubescent, furcate with rudimentary secondary branches; the latter blunt glabrous; sublateral seta 1.2 times as long as submedian seta. Sternite process proportionately broad, anterior part divided into two rounded lobes by a shallow median incision; appendages stump-like, caps flat low. Process with distinct pubescence, appendages glabrous.</p> <p> Setae on anterior tergites as submedian setae on tergal side of head, posteriorly growing subcylindrical; 4+4 setae on tergite I, 6+6 setae on II–V, 4+2 on VI (Fig. 27). Posterior setae on VI somewhat clavate pubescent, 0.3(– 0.4) of interdistance and 1.3 times as long as the length of pygidial setae <i>a</i> 1. Tergites glabrous.</p> <p> <i>Bothriotricha</i>. Relative lengths: <i>T</i> 1=100, <i>T</i> 2=(96–)105, <i>T</i> 3=104(–111), <i>T</i> 4=110(–126), <i>T</i> 5 =126(–158). Axes thin simple straight, <i>T</i> 3 thickest; pubescence of simple straight hairs, oblique on most proximal parts, outwards erect on <i>T</i> 1 and <i>T</i> 2.</p> <p> <i>Genital papillae</i> (Fig. 24). Longish, proximal half cylindrical, roundly conical distally, base two-parted, setae thin, 0.5 of the length of papilla.</p> <p> <i>Legs</i> (Figs 25, 26). Setae on coxa and trochanter (Fig. 25) of leg 9 furcate, main branch folioform, secondary branch clavate, both faintly pubescent. These setae on more anterior legs with broadly folioform main branch and rudimentary secondary branch. Tarsus of leg 9 (Fig. 26) tapering, (3.4–)3.5 times as long as the greatest width. Proximal seta tapering pointed, with depressed pubescence, 0.5 of the length of tarsus and 4.2 times as long as cylindrical blunt striate distal seta.</p> <p> <i>Pygidium</i> (Figs 27, 28). <i>Tergum</i> (Fig. 27). Posterior part with broad indention between <i>st</i>. Relative lengths of setae: <i>a</i> 1=10, <i>a</i> 2 =(14–)16, <i>a</i> 3=(15–))16, <i>st</i> =(7–)8. <i>a</i> -setae tapering pointed pubescent, <i>st</i> thin cylindrical glabrous (in one of the paratypes pubescent and cleft apically), <i>a</i> 1 straight, directed upwards-outwards, <i>a</i> 2, <i>a</i> 3 and <i>st</i> curved inwards, the latter also converging. Distance <i>a</i> 1 <i>-a</i> 1 2.1 times as long as <i>a</i> 1, distance <i>a</i> 1 <i>-a</i> 2 5.0(-8) times as long as distance <i>a</i> 2- <i>a</i> 3; distance <i>st-st</i> 2.7(-2.8) times as long as <i>st</i> and as long as distance <i>a</i> 1 <i>-a</i> 1. Cuticle glabrous except in the middle of posterior indention between <i>st</i>, there dense, erect pubescence.</p> <p> <i>Sternum</i> (Fig. 28). Posterior margin between <i>b</i> 1 straight. Relative lengths of setae (pygidial <i>a</i> 1=10): <i>b</i> 1=(24–)28(–30), <i>b</i> 3=(7–)8. Setae cylindrical, <i>b</i> 1 tapering glabrous, <i>b</i> 3 cylindrical pubescent; the latter placed near the posterior margin of sternum; <i>b</i> 1 1.1 times as long as interdistance, <i>b</i> 3 1.0(–1.1) times as long as interdistance. Sternum glabrous.</p> <p> <i>Anal plate</i> about as broad as long, subcircular, lateral margins convex, U-shaped posterior incision with depth about half of the length of plate, posterior corners each with a small swelling; pubescence sparse but distinct, most prominent on swellings.</p>Published as part of <i>Scheller, Ulf, 2011, Pauropoda (Myriapoda) from Great Smoky Mountains National Park, U. S. A., with descriptions of four new species, pp. 36-48 in Zootaxa 2962</i> on pages 42-45, DOI: <a href="http://zenodo.org/record/202418">10.5281/zenodo.202418</a>
Donzelotauropus tenuitarsus Scheller, 2011, n. sp.
<i>Donzelotauropus tenuitarsus</i> n. sp. <p>Figs 29–36</p> <p> <b>Type locality.</b> Tennessee, Sevier Co., Appalachian Trail, 350 m east of Road Prong trailhead, 3536.707N, 8328.033W, steep slope at log gate, under rotten log, 17 September 2002, U. Scheller.</p> <p> <b>Type material.</b> Holotype: ad. 9(Ƥ), data as above. Paratype: 1 ad. 9(Ƥ), same data as holotype.</p> <p> <b>Nontype material.</b> Tennessee, Sevier Co., Ramsey Cascades Trail, 35º37ʹ75"N, 83º51ʹ62"W, leaf litter, 1 ad. 9(Ƥ), 1 2002, leg. J.P. Gruber. – 3 specimens.</p> <p> <b>Diagnosis</b>. There are characters in the antennae, collum segment, legs, pygidial tergum and anal plate which connect <i>D. tenuitarsus</i> <b>n. sp.</b> to <i>D. diffisus</i> Scheller from Alaska (Scheller 1986). The following can be used for their distinction: calyces of antennal flagella glabrous in <i>D. tenuitarsus</i>, distincrly pubescent in <i>D. diffisus</i>, 4th antennal segment with 6 setae, not 4, antennal seta <i>p</i> longer than antennal branch <i>t,</i> not shorter than, caps of collum processes thin, not distinctly vaulted, tarsus of leg 9 at least 7 times longer than wide, not 5–6, and the proportion <i>b</i> 3/ <i>b</i> 3- <i>b</i> 3 1.1 times longer than distance <i>b</i> 3- <i>b</i> 3, not 0.5-0.8 of that distance.</p> <p> <b>Description</b>. <i>Head</i> (Fig. 29). Setae on tergal side subcylindrical, with oblique pubescence, <i>a</i> 3 in 2nd row and <i>l</i> - group thin pointed. Relative lengths of setae, 1st row (holotype only): <i>a</i> 1=10; <i>a</i> 2=12; 2nd row: <i>a</i> 1=?, <i>a</i> 2=16, <i>a</i> 3=8; 3rd row: <i>a</i> 1=11, <i>a</i> 2=14; 4th row: <i>a</i> 1=16, <i>a</i> 2=?, <i>a</i> 3=?, <i>a</i> 4=7; <i>l</i> 1= <i>l</i> 3=16, <i>l</i> 2=11. The ratio <i>a</i> 1 <i>/a</i> 1 <i>-a</i> 1 is in 1st row 1.4, in 2nd row?, in 3rd row 0.8, in 4th row 1.2. Temporal organs large, as long as their shortest interdistance; posterior pore not ascertained. Head cuticle glabrous.</p> <p> <i>Antennae</i> (Fig. 30). Segment 4 with 5 cylindrical setae, all blunt, obliquely pubescent. Relative lengths of setae: <i>p</i> =100, <i>p</i> ΄=(92–)95, <i>p"</i> =(25–)29, <i>p</i> ΄΄΄=20, <i>r</i> =17, <i>u=</i> 9. Tergal seta <i>p</i> as long as (–1.2 times as long as) the length of tergal branch <i>t</i>. The latter branch fusiform, 5.0 times as long as its greatest diameter and 1.2(–1.4) times as long as sternal branch <i>s,</i> that branch 2.9(–3.0) times as long as its greatest diameter and with its posterodistal corner deeply truncate. Seta <i>q</i> as <i>p</i> ʹ of 4th segment, almost as long as <i>s</i>. Relative lengths of flagella (with base segments included) and of base segments alone: <i>F</i> 1=100, <i>bs</i> 1=16, <i>F</i> 2=(74–)80, <i>bs</i> 2=16, <i>F</i> 3≈60, <i>bs</i> 3=18. The <i>F</i> 1 2.2(–2.4) times as long as <i>t</i>, <i>F</i> 2 and <i>F</i> 3 2.0 and 1.4 times as long as <i>s</i> respectively. Distal calyces glabrous, on <i>F</i> 1 conical, longer than wide, other calyces with rounded tips. Globulus <i>g</i> 3.3 times as long as its greatest diameter, stalk curved; ≈12 bracts, partly bifurcate; capsule subspherical. Diameter of <i>g</i> 0.9 of the greatest diameter of <i>t</i>. Antennae glabrous.</p> <p> <i>Trunk</i> (Figs 31, 32). Setae of collum segment (Fig. 31) narrowly leaf-shaped, distinctly pubescent, furcate with rudimentary blunt glabrous secondary branch; sublateral seta 1.2 times as long as submedian seta. Sternite process very broad anteriorly, there divided into two rounded lobes by a shallow median incision; appendages flattened low, caps thin, inconsiderably vaulted. Process with distinct pubescence, appendages glabrous.</p> <p> Setae on anterior tergites as submedian setae on tergal side of head, increasing in length posteriorly; 4+4 setae on tergite I, 6+6 setae on II–V, 4+2 on VI. Posterior setae on VI (Fig. 32) tapering pointed, with sparse depressed pubescence, 1.8 times as long as interdistance and 1.4 times as long as the length of pygidial setae <i>a</i> 1.</p> <p> <i>Bothriotricha</i>. Relative lengths (paratype): <i>T</i> 1=100, <i>T</i> 2=135, <i>T</i> 3=161, <i>T</i> 4=184, <i>T</i> 5=323. Axes thin simple straight, pubescence of simple straight hairs, oblique on most proximal parts, outwards erect on <i>T</i> 1 and <i>T</i> 2, sparse on <i>T</i> 5.</p> <p> <i>Legs</i> (Figs 33, 34, 36). Long slender, leg 9 (Fig. 34) at least twice longer than leg 1. Setae on tibiae and proximal seta on tarsi long outstanding. Setae on coxa and trochanter (Fig. 33) of leg 9 thickest at base, furcate, branches cylindrical, main branch thick, secondary branch thin, both distinctly pubescent. These setae on more anterior legs with broadly folioform main branch and rudimentary glabrous secondary branch. Tarsus of leg 9 (Fig. 34, 36) slen- der tapering, (6.1–)7.4 times as long as the greatest width. Proximal seta tapering pointed, with a few pubescence hairs only, 0.4 of the length of tarsus and 4.0(–4.5) times as long as cylindrical blunt striate distal seta.</p> <p> <b>FIGURES 29–36.</b> <i>Donzelotauropus tenuitarsus</i> <b>sp. n.</b>, holotype, Ƥ. 29, head, submedian and right part, tergal view. 30, right antenna, tergal view. 31, collum segment, median and left part, sternal view. 32, tergite VI, posteriomedian part. 33, seta on trochanter of leg 9. 34, leg 9, tibia and tarsus, anterior view. 35, pygidium, median and right part, sternal view. 36, posterior part of trunk with pygidium, leg 9 and bothriotrix <i>T</i> 5. Pubescence only partly drawn in 36. Scale a: Figure 36; b: Figure 34; c: Figures 31, 33; d: Figure 32; e: Figures 29, 30, 35, 36.</p> <p> <i>Pygidium</i> (Figs 34, 35, 36). <i>Tergum</i> (Fig. 35). Posterior part with broad indention between <i>st</i>. Relative lengths of setae: <i>a</i> 1=10, <i>a</i> 2 =(14–)16, <i>a</i> 3=(15–))16, <i>st</i> =(7–)8. <i>a</i> -setae tapering pointed pubescent, <i>st</i> thin cylindrical glabrous (in one of the paratypes pubescent and cleft apically), <i>a</i> 1 straight, directed upwards-outwards, <i>a</i> 2, <i>a</i> 3 and <i>st</i> curved inwards, the latter also converging. Distance <i>a</i> 1 <i>-a</i> 1 2.1 times as long as <i>a</i> 1, distance <i>a</i> 1 <i>-a</i> 2 5.0(–8) times as long as distance <i>a</i> 2- <i>a</i> 3; distance <i>st-st</i> 2.7(–2.8) times as long as <i>st</i> and as long as distance <i>a</i> 1 <i>-a</i> 1. Cuticle with short dense pubescence.</p> <p> <i>Sternum</i> (Fig. 35). Posterior margin between <i>b</i> 1 straight. Relative lengths of setae (pygidial <i>a</i> 1=10): <i>b</i> 1=(24–)28(–30), <i>b</i> 3=(7–)8. Setae cylindrical, <i>b</i> 1 tapering glabrous, <i>b</i> 3 cylindrical pubescent; the latter placed near the posterior margin of sternum; <i>b</i> 1 1.1 times as long as interdistance, <i>b</i> 3 1.0(–1.1) times as long as interdistance. Sternum. Cuticle with short dense pubescence.</p> <p> <i>Anal plate</i> (Fig. 35) about as broad as long, lateral margins concave, large U-shaped posterior incision, depth about half of the length of plate, posteriolateral corners lengthened into cylindrical blunt diverging appendages, length about as long as ¾ of the length of plate, these appendages also somewhat curved inwards; at each side of Ushaped incision a thinner cylindrical blunt appendage pointing backwards, length as long as plate; plate and appendages shortly pubescent, most distinctly on appendages.</p>Published as part of <i>Scheller, Ulf, 2011, Pauropoda (Myriapoda) from Great Smoky Mountains National Park, U. S. A., with descriptions of four new species, pp. 36-48 in Zootaxa 2962</i> on pages 45-47, DOI: <a href="http://zenodo.org/record/202418">10.5281/zenodo.202418</a>
Trachypauropus lusitanicus Scheller, 2014, n. sp.
Trachypauropus lusitanicus n. sp. Figs 61–70 Type specimen. Holotype: ad. 9 (♀), Portugal, W of Lisboa, Parque de Sintra, June 11, 2012, in litter under deciduous trees, W. Niedbala leg. Type specimen deposited in the Zoological Museum, University of Lund, Sweden. Non type specimens. Same data as holotype, 6 subad. 8 (♀), 7 juv. 6, 1 juv. 3. FIGURES 61–70. Trachypauropus lusitanicus n. sp., holotype ad. 9 (♀). 61. Left antenna sternal view. 62. Collum segment, median and left part, sternal view. 63. Tergite I, central part. 64. Tergite II, right posterior corner. 65. Tergite VI. Left posterior corner. 66. T 1. 67. T 3. 68. Seta of trochanter of leg 9. 69. Tarsus of leg 9. 70. Pygidium, sternal view. Scale: a: 66–69; b: 61. 62; c: 63 –65, 70. Diagnosis. The new species is well delimited by the occurrence of two anterior setae on the median process of the collum segment, the shape of the a -setae of the pygidial tergum, a 1 short, pointed, a 2 scutellate, a 3 long, pointed, and the shape of the anal plate, lateral appendages straight and directed posteriorly. The affinities of the species are not possible to trace from the present knowledge of the genus. Derivation of name. A Latinised adjective of Lusitania = Portugal, referring to the collecting site. Description. Length. 1.01 mm. Head. Setae not available for study. Antennae (Fig. 61). Chaetotaxy of segments 1-4: 2 / 2 / 3 / 4; segment 3 with distinct globulus g´. Relative lengths of setae of 4 th segment: p = 10, p´= 11, p´´ = 9, r = 6, all thin, pointed, annulated. Sternal branch s, anterior margin = 35, posterior margin = 29, diameter of base = 10, maximum diameter = 16, length of globulus g = 31, diameter = 9 µm, stalk cylindrical, number of bracts ≈ 12, lengths of bracts ≈ 7, capsule length = 4, diameter = 6, q = 27 µm. Anterior margin/length of g = 1.1, anterior margin/length of q 1.1, anterior margin/posterior margin 1.2, anterior margin/maximum diameter = 2.2, maximum diameter/diameter of base = 1.6. Tergal branch t subcylindrical, widest in distal half, length = 36, diameter of base = 8, maximum diameter = 10, distance base to pore not studied, length of t/ maximum diameter = 3.6 µm. Flagella, length: F 1 = 63, F 2 = 42, F 3 = 65, bs 1 = 9, bs 2 = 10, bs 3 = 7 µm. Calyces glabrous, those of F 1 helmet-shaped, those of F 2 and F 3 conical with distinct point. Trunk (Figs 62, 65). Setae of collum segment similar, short, simple, annulated. appendages short with rounded distal cap, glabrous, no collar; sternite process cut squarely anteriorly and with two short annulated setae protruding anteriorly from anterolateral corners. Tergites with two types of protuberances: 1. Large, curved, pointed spines pointing backwards from posterior part of oval glabrous areas, most lateral spines, largest, probably winged; 2. Numerous small, conical, blunt protuberances evenly distributed all over the surface. Bothriotricha (Figs 66–67). Bothriotricha T 1 (Fig. 66), T 2 and T 4 with very thin axes and sparse, short erect pubescence, in whorls distally, T 3 (Fig. 67) clavate, distal 1 / 3 thickened, pubescence distinct, hairs curved, oblique, T 5 proportionately thick and with oblique pubescence hairs not arranged in whorls, Legs (Figs 68, 69). Legs 1 and 9 5 -segmented, tarsi divided in intervening pairs, legs there 6 -segmented. Setae on coxa and trochanter (Fig. 68) of all legs furcate with rudimentary secondary branch, primary branch short, cylindrical, blunt, striate. Tarsus of leg 9 (Fig. 69) almost cylindrical, 4.1 times as long as the greatest diameter, only one tergal seta placed near the middle, seta pointed, glabrous, length 0.2 of the length of the tarsus. All legs with large main claw and small anterior secondary claw. Length of main claw of leg 1 0.2 of the length of the tarsus, 0.3 in leg 9. Pygidium (Fig. 70). Tergum: Posterior margin curved roundly but with low lobe between setae st; a -setae of different shape, all glabrous, a 1 and a 3 tapering pointed, the former short, the latter much longer and with thick base, a 2 scutellate, length a 1 = 5, a 2 = 13, a 3 = 20, st = 1, a 1 - a 1 = 20, a 2 - a 2 = 45, a 3 - a 3 = 66, st – st = 18 µm; st-st/a 1 - a 1 = 0.9, a 1 / a 1 - a 1 = 0.2, a 1 / a 1 - a 2 = 0.3, a 1 - a 1 / a 2 - a 3 = 0.3. Sternum: Low broad lobe with small median incision between setae b 1. These setae glabrous pointed, b 2 and b 3 pointed too and with sparse oblique pubescence, b 3 very thin; b 1 = 32, b 2 = b 3 = 12, b 1 - b 1 = 30, b 2 - b 2 = 51, b 3 - b 3 = 10 µm; b 1 - b 2 = 22, b 2 - b 3 = 18 µm; b 1 - b 1 / b 1 = 0.9, b 1 / b 2 = 3.7. Anal plate lyrate and divided into four straight posteriorly directed cylindrical branches: two thin lateral ones being shortest, 0.4 of the length of the plate, and two submedian ones, longer, thicker, widened a little at distal ends, there cut obliquely with inner side somewhat lengthened, each branch with a bladder-shaped and distinctly pubescent appendage with thin stalk proceeding from an excavation in the obliquely cut end of the branch, length of bladders 0.4 of the length of submedian branch.Published as part of Scheller, Ulf, 2014, New records of Pauropoda (Myriapoda) with descriptions of new taxa, pp. 301-332 in Zootaxa 3866 (3) on pages 325-327, DOI: 10.11646/zootaxa.3866.3.1, http://zenodo.org/record/22744
Allopauropus circulus Scheller, 2014, n. sp.
Allopauropus circulus n. sp. Figs 1–10 Type specimen s. Holotype: ad. 9 (♀), France, Var, Mt Vinaigre, loc. # 83-145, October 6, 1995, 81653, Y 43.5033, Berlese extraction of soil, depth 20–40 cm, February 24, 1995, A. Bedos & L. Deharveng leg. Paratype: 1 subad. 8 (♀), same data as holotype. Described below. Type specimens deposited in the Zoological Museum, University of Lund, Sweden. Diagnosis. The new species belongs to a group of species characterized by a short anal plate with two posterior appendages, short pygidial setae a 1 and st and very long setae a 2 and a 3. It may be closest to A. maoriorum Remy (Remy, 1956), a wide ranging species in the southern hemisphere (known from Australia, New Zealand, New Caledonia and Chile) but known also from Sabah in the Malaysian part of Borneo. The two species are very alike but can be distinguished by many obvious dissimilarities: the shape of the antennal globulus g, stalk short and wide in A. maoriorum, distinctly thin in A. circulus; the shape of the bothriotricha T 3, axes increasing in width from the base and outwards, then being cylindrical, pubescence short, straight, oblique, not decreasing in width outwards and with long branched pubescence hairs arranged in whorls; the shape of the anal plate, with lateral corners, not circular; the posterior appendages of the anal plate appendages almost parallel, not diverging; and the size of the median lobe of the posterior margin of the pygidial sternum, small, not large. Derivation of name. From the Latin circulus = circle, referring to the shape of the anal plate. Description. Length. 0.47 (- 0.65) mm. Head (Fig. 1). Tergal setae of medium lengths, subcylindrical, annulated, blunt. Relative lengths of setae, 1 st row: a 1 = a 2 = 10; 2 nd row: a 1 = 13, a 2 = 19, a 3 = 17; 3 rd row: a 1 = 11, a 2 = 14; 4 th row: a 1 = 19, a 2 = 20, a 3 = 17, a 4 = 16; lateral group setae not studied. Ratio a 1 /a 1 – a 1 in 1 st and 3 rd rows 1.1, 2 nd 0.8, 4 th 1.3. Temporal organs in tergal view short, length 0.9 of shortest interdistance; short but wide pistil near middle, length well 0.1 of the length of the temporal organ. Head cuticle glabrous. Antennae (Fig. 2). No g´ on segment 3; segment 4 with 5 subcylindrical annulated setae, u thinnest, their relative lengths: p = 100, p' = 70 (- 76), p'' = 61 (- 67), p´´ ´= 33 (- 38), r = 47 (- 60), u = 3. Tergal seta p 1.1 (- 1.2) times as long as the length of tergal branch t. The latter (2.4 -) 2.5 times as long as its greatest diameter and 0.9 (-1.0) of the length of sternal branch s, that branch 2.1 times as long as its greatest diameter; anterodistal corner of s distinctly truncated. Seta q as p and p´ of 4 th segment, (1.1 -) 1.2 times as long as the length of s, insertion point of q just below the level of the insertion point of flagella F 2. Relative lengths of flagella (basal segments included) and basal segments: F 1 = 100, bs 1 = (9 -) 10 (- 11); F 2 = 42 (- 45), bs 2 = 7 (- 8); F 3 = 89 (- 91), bs 3 = 9 (- 11). The F 1 3.2 (- 3.3) times as long as t, F 2 and F 3 1.3 (- 1.5) and 2.8 (- 3.1) times as long as s respectively. Distal calyces of F 1 and F 3 large flattened, those of F 2 small, obliquely helmet-shaped, distal part of flagella axes almost not at all widened below calyces. Globulus g with short but distinct stalk, as wide as long, ≈ 9 bracts, capsule with flattened bottom, width of g almost as long as greatest with of t. Antennae glabrous. Trunk (Figs 3–4). Setae of collum segment (Fig. 3) similar, furcate, main branch cylindrical, pointed, annulated, secondary branch rudimentary, pointed; sternite process triangular, narrow and blunt anteriorly, appendages cylindrical with subspherical caps with distinct collar, process and appendages glabrous. Setae on anterior tergites similar to submedian setae of head but lengthening and also growing thinner posteriorly, on posterior tergites with short oblique pubescence; 4 + 4 setae on tergite I, 6 + 4 on II, 6 + 6 on III-IV, 6 + 4 on V, 4 + 2 on VI. Submedian posterior setae on tergite VI (Fig. 4) thin, pointed, somewhat curved inwards, length 1.3 times as long as interdistance and (4.1 -) 5.4 times as long as the length of pygidial setae a 1. Tergites glabrous. Genital papillae (Fig. 5) with subcylindrical proximal half, distal part rounded on outer sides, papillae 1.7 times as long as their greatest diameter; distal seta not fully 0.5 of the length of the papilla. Setae on coxa and trochanter of 2 nd pair of legs as on 9 th pair of legs. Bothriotricha (Fig. 6): Relative lengths: T 1 = T 2 = 100, T 3 = 82, T 4 = 65, T 5 = 129; axes simple, T 1 (Fig. 6) and T 2 very thin, proximal halves glabrous, distal halves with long sparse and partly branched hairs; posterior setae thicker, T 3 (Fig. 6) and T 5 with oblique simple pubescence hairs in proximal third, more outwards with longer and branched hairs in distinct whirls; T 5 with very delicate pubescence. FIGURES 1–10. Allopauropus circulus n. sp. 1 –4, 6– 10, holotype ad. 9 (♀), 5, paratype ad. 9 (♂). 1, Head, right half, tergal view. 2. Left antenna, tergal view. 3. Collum segment, median and left part, sternal view. 4. Posterior part of tergite VI. 5. Genital papillae, anterior view. 6. T 3. 7. Seta on trochanter of leg 9. 8. Tarsus of leg 9. 9. Pygidium, posteriomedian and left part, sternal view. 10. Anal plate, lateral view. Scale: a: 3–7; b: 1 –2, 8– 10. Legs (Figs 7, 8): Setae on coxa and trochanter (Fig. 7) of leg 9 simple striate blunt. Tarsus of leg 9 (Fig. 8) distinctly tapering, 4.7 times as long as its greatest diameter; proximal seta long, tapering, glabrous, distal seta short, cylindrical, blunt, with short pubescence, proximal seta 0.6 of the length of tarsus and 5.8 times as long as distal seta. Cuticle of tarsus with short pubescence. Pygidium (Figs 9, 10): Tergum: Posterior margin almost straight, only inconsiderable bulge between setae st. Relative lengths of setae: a 1 = 10, a 2 = (33 -) 35 (- 36), a 3 = (65 -) 83, st = (2 -) 3; all setae directed posteriorly, a -setae thin, tapering, a 1 almost straight, annulated, a 2 and a 3 very thin distally, the latter curved inwards, st very short, clavate, glabrous. Distance a 1 – a 1 2.3 times as long as interdistance; distance a 1 – a 2 as long as distance a 2 – a 3; distance st–st 9 times longer than st and (1.1 -) 1.2 times as long as distance a 1 – a 1. Cuticle glabrous. Sternum: Setae b 1 on small semi-circular lobes, margin in between straight with small median lobe having a broadly V-shaped median incision. Relative lengths of setae (pygidial a 1 = 10): b 1 = (43 -) 48, b 2 = (25 -) 29, b 3 = (12 -) 15. b 1 cylindrical, annulated, blunt, b 2 and b 3 tapering, pointed, particularly the former, b 1 0.9 of interdistance, b 2 1.5 times as long as distance b 1 – b 2, b 3 0.4 of interdistance. Cuticle glabrous. Anal plate (Figs 9, 10) pointing obliquely upwards, circular, glabrous and with two short clavate diverging appendages protruding backwards from distal part of sternal side, length of appendages (0.8 -) 0.9 of the length of the plate.Published as part of Scheller, Ulf, 2014, New records of Pauropoda (Myriapoda) with descriptions of new taxa, pp. 301-332 in Zootaxa 3866 (3) on pages 313-315, DOI: 10.11646/zootaxa.3866.3.1, http://zenodo.org/record/22744
Mojingapauropus biappendiculatus Scheller, 2014, n. sp.
Mojingapauropus biappendiculatus n. sp. Figs 47 –60 Type specimens. Holotype: ad. 9 (♀), Panama, near Ft. Sherman, Mojinga Swamp. in leaf litter, August 10, 1979, R.J. Snider leg. Paratype: ad. 9 (♀), same data as holotype. Type specimens deposited in the Zoological Museum, University of Lund, Sweden. Derivation of name. From the Latin bis = two and appendix = appendage, referring to the appendages of the posterior part of last tergite. Description. Length. (0.65) 0.84 mm. Head (Figs 47–49). Posterior part covered by tergite I, only most anterior and lateral margins visible from above. Setae of three types: 1, anteriomedian one cylindrical, blunt, striate; 2, submedian and anteromarginal ones lanceolate-irregular; 3, a few lateral ones cylindrical, pointed, with short pubescence or glabrous. Cuticle of head coarse. Only small parts of temporal organs visible from above (Fig. 47), organs broadest anteriorly (Figs 48-49), tergal margin even, anterior and lower margins with irregularities, broad lobe in the direction of the mouth, only posterior and lower margins partly and narrowly uplifted. Antennae (Fig. 50). Segment 4 longish, five thin, cylindrical, annulated setae, their relativ lengths: p = 100, p ' = (80) 81, p '' = 49 (69), r = 49 (41), u = 6 (9). Tergal seta p 0.7 (0.8) of the length of tergal branch t. The latter branch almost cylindrical, 3.0 (3.3) times as long as the greatest diameter, (1.1) 1.2 times as long as sternal branch s, that branch (2.1) 2.3 times as long as the greatest diameter, anterodistal corner distinctly truncate. Seta q as seta r of segment four, short, (0.4) 0.5 of the length of s. Relative lengths of flagella (with base segments included) and base segments: F 1 = 100, bs 1 = (7) 8, F 2 = (40) 42, bs 2 = 5 (6), F 3 = (88) 94, bs 3 = 4 (5). F 1 2.8 (3.3) times as long as t, F 2 and F 3 1.4 (1.5) and 3.1 (3.3) times as long as s respectively. Distal calyces of F 1 and F 2 more pointed than those of F 3, distal parts of flagella axes widened only just below calyces. Globulus g subspherical with distinct, thin stalk, 2.9 times as long as its greatest diameter; ≈ 11 bracts; capsule flattened. Diameter of g almost equal to the greatest diameter of t. Antennae glabrous Trunk (Figs 47, 51, 52 –55). Setae of collum segment (Fig. 51) furcate, primary branch cylindrical, blunt, annulated, secondary branch rudimentary, pointed, glabrous, sublateral and submedian setae of the same length. Sternite process with deep anterior, narrowly U-shaped incision; appendages ovoid with short pubescence, caps distinct, glabrous. Tergites (Figs 47, 53– 55). All entire, strongly sclerotized, 3 rd (Fig. 53) and 4 th ones broadest. Setae in irregular rows, scutellate, inner ones largest. Total number of setae on tergites (holotype only): I (Fig. 47) with 14, II with 36. III (Fig. 53) with 60, IV with 64, V with 46, VI (Fig. 54) with 18. All tergites except I with two longitudinal rows of clear spots (Fig. 53): II with 5 ones in each row, III-V with 7, VI with 2. Cuticle of tergites coarse, distinctly grainy. Bothriotricha (Fig. 57). Relative lengths (holotype only): T 1 = 100, T 2 = 75 and 76, T 3 = 34 and 36, T 4 = 70 and 75, T 5 = 75. Axes simple, proximal halves glabrous, distal halves very thin except in T 3, with thin and dense erect pubescence. T 3 (Fig. 57) clavate distally, almost glabrous, thickest, most distinct in proximal ¾. Legs (Figs 58-60). All legs 5 -segmented. Seta on coxa of leg 9 (Fig. 58) simple, cylindrical, blunt, annulated, seta on trochanter (Fig. 59) furcate, branches as on coxal seta but secondary branch thin and short, not half of the length of main branch. More anteriorly these setae as seta on trochanter of leg 9 but with rudimentary secondary branch. Tarsus of leg 9 (Fig. 60) short, tapering, (2.1) 2.3 times as long as greatest diameter. Setae curved, proximal one thin, tapering, pointed, with sparse oblique pubescence on distal half; distal seta cylindrical, blunt, annulated; proximal seta not fully 0.4 of the length of tarsus and as long as distal seta. Cuticle glabrous. Pygidium (Figs 52, 54– 56). Much narrower than tergite VI (Figs 52, 54). Tergum (Fig. 55). Hind margin straight with very small V-shaped incision between st. Relative lengths of setae: a 1 = 10, a 2 = 10 (11), a 3 = 20, st = 2; a 1 scutellate, pointed, in tergal view ovoid, point directed downwards, a 2 lanceolate with very short pubescence, a 3 long, tapering, pointed, glabrous, curved inwards, st small, clavate, shortly pubescent, all directed posteriorly: distance a 1 -a 1 1.3 times as long as a 1 and 0.8 of distance a 1 -a 2, that distance 3 times longer than distance a 2 - a 3; distance st-st 10 times longer than st and (1.4) 1.7 times as long as distance a 1 -a 1. Cuticle glabrous. Sternum (Figs 52, 56). Posterior margin with deep, broadly V-shaped indentation between setae b 1 and a large median posteriorly directed lobe with posteromedian incision below anal plate. Setae b 1 projecting backwards from posterior part of large triangular lobes. Relative lengths of setae (pygidial a 1 = 10): b 1 = 33 (35); b 3 = 7. These setae cylindrical, b 1 tapering, glabrous, b 3 thin, cylindrical, with short oblique pubescence. b 1 as long as (1.2 times as long as) interdistance, b 3 0.3 of interdistance. Anal plate (Fig. 56) very like some occurring in the Eurypauropodidae genera Trachypauropus, Acopauropus and Samarangopus. Narrowest at base, then widening into four posteriorly directed appendages, two shorter, tapering lateral ones reaching to half of the length of the plate, and two, cylindrical, straight, submedian ones cut obliquely at ends and each provided with a bladder-shaped appendage with very short pubescence, length of appendages 0.3 of the length of the plate. FIGURES 53–60. Mojingapauropus biappendiculatus n. sp., holotype, ad. 9 (♀). 53. Tergite III, median and right part, surface structures only partly drawn. 54. Last tergite, right half, and pygidium, tergal view, surface structures only partly drawn. 55. Pygidium, median and right half, tergal view. 56. Pygidium, sternal view. 57. T 3. 58. Seta on coxa of leg 9. 59. Seta on trochanter of leg 9. 60. Tarsus of leg 9. Scale a: 53, 54; b: 55–57; c: 58–60.Published as part of Scheller, Ulf, 2014, New records of Pauropoda (Myriapoda) with descriptions of new taxa, pp. 301-332 in Zootaxa 3866 (3) on pages 323-325, DOI: 10.11646/zootaxa.3866.3.1, http://zenodo.org/record/22744
Decapauropus pyriformis Scheller, 2014, n. sp.
Decapauropus pyriformis n. sp. Figs 20–29 Type specimens. Holotype: ad. 9 (♀), Italy, Veneto, C.A. Deskalb-Chiarano, TU 1, April 1, 1980. M.G. Paoletti leg. Paratype: ad. 9 (♂), same data as holotype. Type specimens deposited in the Zoological Museum, University of Lund, Sweden. Diagnosis. Many species in the genus have proportionately large antennal globulus g but none of them has a similar set of bothriotricha and anal plate. At present no plausible relationships can be traced Derivation of name. From the Latin pirum = pear and – formis in the shape of, referring to the shape of the antennal globulus g. Description. Length. (0.62) 0.65 mm. Head (Fig. 20). Setae on tergal side of short-medium length, thin cylindrical striate. Relative lengths of setae (holotype only), 1 st row: a 1 = a 2 = 12; 2 nd row: a 1 = a 3 = 12, a 2 = 15, 3rd row: a 1 = 12, a 2 = 16; 4 th row: a 1 = 13, a 2 = a 3 = 20. a 4 =?, l- group setae not studied. The ratio a 1 /a 1 - a 1 in 1 st row 0.7, 2 nd row 0.3, 3 rd row 0.8, 4 th row 1.3. Temporal organs in tergal view broadest in posterior half, their length 0.7 of shortest interdistance. Pore not ascertained. Head cuticle glabrous. Antennae (Fig. 21). Segment 4 with four cylindrical, striate setae, r and p´´ thinnest, their relative lengths: p = 100. p ´= 42, p´´ = 20, r = 41. Tergal seta p 1.6 times as long as tergal branch t. The latter widest in distal half, 2.5 times as long as its greatest diameter and 1.1 times as long as sternal branch s, there anterodistal corner distinctly truncate. Seta q cylindrical, striate, blunt, almost as long as the length of s. Relative lengths of flagella (basal segments included) and basal segments alone: F 1 = 100, bs 1 = 12, F 2 = 47, bs 2 = 6, F 3 = 91, bs 3 = 11. F 1 as long as t, F 2 and F 3 0.5 and 0.9 of the length of s respectively. Distal part of flagella not studied. Globulus g proportionally large, 1.7 times as long as its greatest diameter; ≈ 11 bracts; capsule longer than wide. Diameter of g 1.7 times as long as its greatest diameter and 1.1 times as long as the greatest diameter of t. Antennae glabrous. Trunk (Fig. 22). Collum segment not available for study. Seta on tergites as setae of the head. Two transversal rows, 4 + 4 setae on tergite I,? 4 + 6 on II and III, 6 + 6 on IV and V, 4 + 2 on VI. Submedian posterior setae on VI (Fig. 22) cylindrical, striate, blunt, curved inwards, 0.6 of interdistance and 1.6 times as long as pygidial setae a 1. Bothriotricha (Figs 23–24). Relative lengths: T 1 = 100, T 2 = T 3 = 108, T 4 = 137, T 5 = 167. Axes simple, straight, in all but T 3 very thin, with erect pubescence in whirls on T 1 (Fig. 23), T 2 and T 4. T 3 (Fig. 24) with thickened axes, particularly in distal halves and with very short pubescence of simple hairs, partly arranged in whirls. T 5 with shortest pubescence, on distal part oblique. Legs (Figs 25–27). Seta on coxa (Fig. 25) and trochanter (Fig. 26) of leg 9 furcate, both branches cylindrical, blunt, striate, secondary branch distinctly shorter than primary one. Tarsus of leg 9 (Fig. 27) tapering, glabrous, 3.2 times as long as its greatest diameter, setae cylindrical, blunt, annulated, proximal seta 0.2 of the length of tarsus and 1.3 times as long as distal seta. Pygidium (Figs 28–29). Tergum. Posterior margin between insertion areas of setae a 3 rounded with low semicircular posterior lobe between setae st. Relative lengths of setae: a 1 = 10, a 2 = 5, a 3 = 15, st = 4. Setae subsimilar, a 2 an st cylindrical blunt, a 1 and a 3 tapering, a 3 pointed, all curved inwards and with short pubescence. Distance a 1 - a 1 0.7 of the length of a 1, distance a 1 -a 2 3.0 times as long as distance a 2 - a 3, distance st-st 2.3 times as long as st and 1.3 times as long as distance a 1 - a 1. Sternum. Posterior margin between insertion areas of setae b 1 indented shallowly. Relative lengths of setae (pygidial a 1 = 10): b 1 = 72, b 2 = 20. These setae tapering pointed, annulated distally, b 1 1.2 times as long as interdistance, b 2 0.9 of distance b 1 - b 2. Anal plate (Figs 28–29) nearly square with a posteromedian V-shaped incision reaching the middle of the plate. Two short appendages protruding upwards from the distal parts of the posterior branches. Plate glabrous. FIGURES 20–29. Decapauropus pyriformis n. sp., holotype ad. 9 (♀). 20. Head, right half, tergal view. 21. Right antenna, sternal view. 22. Posteriomedian part and right posterior corner of tergite VI. 23. T 1. 24. T 3. 25. Seta on coxa of leg 9. 26. Seta on trochanter of leg 9. 27. Tarsus of leg 9. 28. Pygidium, posteriomedian and left part, sternal view. 29. Anal plate, lateral view. Scale: a: 21–24; b: 20, 25– 29.Published as part of Scheller, Ulf, 2014, New records of Pauropoda (Myriapoda) with descriptions of new taxa, pp. 301-332 in Zootaxa 3866 (3) on pages 317-318, DOI: 10.11646/zootaxa.3866.3.1, http://zenodo.org/record/22744
Rabaudauropus andinus Remy 1962
Rabaudauropus andinus (Remy, 1962) Chile, Valdivia Prov., Lago Calafquén, moist temperate forest, 1 ad. 9 (♀), Mars 14, 1965, F. di Castri leg.Published as part of Scheller, Ulf, 2014, New records of Pauropoda (Myriapoda) with descriptions of new taxa, pp. 301-332 in Zootaxa 3866 (3) on page 310, DOI: 10.11646/zootaxa.3866.3.1, http://zenodo.org/record/22744
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