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Hibiscus analalavensis M. Hanes & G. E. Schatz 2020, sp. nov.
No full textHibiscus analalavensis M. Hanes & G.E. Schatz, sp. nov. (Fig. 3A, 4A). Holotypus: MADAGASCAR. Reg. Atsinanana [Prov. Toamasina]: Analalava NP, 17°42'S 49°26'E, 16.XI.2007, fl., Andriamiarinoro 102 (MO-6562524!; iso-: P [P00847913]!, TAN). Hibiscus analalavensis M. Hanes & G.E. Schatz differs from H. vohipahensis M. Hanes & G.E. Schatz in having shorter petioles, 4–5 mm (vs. 6–10 mm), and shorter pedicels, 30–32 mm (vs. 45 mm). Shrub to small tree 3–5 m tall, 3–4 cm dbh; twigs moderately densely covered with golden stellate trichomes. Leaves 7.3–11.9 × 3.3–5.1 cm, narrowly obovate, the base cuneate and abruptly attenuate, the margin strongly revolute, the apex rounded to obtuse or slightly retuse, glabrous above, nearly completely glabrous below, venation brochidodromous with 6–7 secondary veins per side, midvein raised above and below, secondary veins slightly raised above and below, tertiary venation reticulate, slightly raised above and below; petiole 4 – 5 mm, c. 1 mm in diam., moderately densely covered with stellate trichomes. Flowers solitary, axillary, pendent; pedicels 30 –32 mm long, 1.4 mm in diam., densely covered with reddish-gold, erect simple, fasciculate and stellate trichomes; epicalyx with 5 lobes fused at the base for 2 mm, the lobes 15 × 6– 9 mm, broadly ovate, the apex acute, densely covered with rufous golden stellate trichomes on the outside, densely covered with golden appressed simple and fasciculate trichomes inside; calyx with 5 lobes, fused at the base for 6–9 mm (c. half of length), the lobes 20 × 18 mm, triangular, the apex triangular, very densely covered with golden stellate trichomes outside, very densely covered with appressed curly white trichomes inside; petals 5, 50 × 35– 37 mm, obovate, the apex rounded, white with a peach blush, meanwhile they turn from pink to white with age, densely covered with light golden stellate trichomes outside, glabrous inside with evident venation; staminal column c. 30 mm long, c. 2 mm in diam., white, glabrous, with a tuft of dense, golden, erect, simple trichomes at the apex; stamens c. 70, filaments 10 mm long, white, anthers c. 1.5 × 1.5 mm; styles 5, c. 16 mm long, white, slender, moderately densely covered with white, erect, simple and fasciculate trichomes, stigma c. 0.5 × 0.8 mm, bulbous. Fruit unknown. Vernacular names. – “Afopotsy” (Lehavana 426). Distribution and ecology. – Hibiscus analalavensis occurs in the lowland moist evergreen forest of Analalava near Foulpointe from 40 to 90 m in elevation. Conservation status. – Hibiscus analalavensis is known from 4 locations within the protected area of Analalava. Despite the very restricted extent of occurrence (EOO) (c. 1 km ²), there appear to be no current threats to H. analalavensis. Thus, the species is assigned a status of “Least Concern” [LC] using IUCN Red List Categories and Criteria (IUCN, 2012), with the caution that this status is highly dependent on continued effective protection. Notes. – Hibiscus analalavensis resembles H. vohipahensis in that they have whitish flowers but differs from the latter species by having shorter petioles, 4–5 mm (vs. 6–10 mm), shorter pedicels, 30–32 mm (vs. 45 mm), and smaller petals, 15 × 6–9 mm (vs. 12–17 × 12–18 mm). Paratypi. – MADAGASCAR. Reg. Atsinanana [Prov. Toamasina]: Foulpointe, Morarano, forêt d’Analalava, à 7 km SW de Foulpointe, 17°42'05"S 49°27'31"E, 87 m, 12.II.2007, fl., Lehavana 426 (MO, P, TAN); ibid. loco, 17°42'14"S 49°27'19"E, 40 m, 25.XI.2015, fl., Ratovoson 2092 (MO, P, TAN); ibid. loco, 19.XII.1967, fl., Service Forestier 28071 (P).Published as part of Hanes, Margaret M., Schatz, George E. & Callmander, Martin W., 2020, Transfer of the Malagasy genera Humbertianthus and Macrostelia to Hibiscus (Malvaceae) with description of four new species, pp. 193-202 in Candollea 75 (2) on pages 198-200, DOI: 10.15553/c2020v752a4, http://zenodo.org/record/568374
Hibiscus ankeranensis M. Hanes & G. E. Schatz 2020, sp. nov.
No full textHibiscus ankeranensis M. Hanes & G.E. Schatz, sp. nov. (Fig. 3B, 4B). Holotypus: MADAGASCAR. Reg. Atsinanana [Prov. Toamasina]: Dist. Brickaville, Com. Maroseranana, Fkt. Ambodilendemy, Andrangato river, Ankerana, 18°26'52"S 48°46'28"E, 346 m, 13.III.2011, fl., Antilahimena 7544 (MO-2652132!; iso-: P [P06774026]!, TAN). Hibiscus ankeranensis M. Hanes & G.E. Schatz differs from H. calyculatus (Hochr.) M. Hanes, G.E. Schatz & Callm. by its epicalyces, 5 non-symmetrical, succulent in vivo lobes (vs. 4 small, reflexed lobes), calyx color, green (vs. red), and its nonreflexed petals. Tree c. 6 m tall, 12 cm dbh; twigs moderately densely covered with light, golden, semi-erect, simple, fascicled and stellate trichomes. Leaves 3.5– 9.8 × 1.9 –3.9 cm, elliptic to narrowly obovate, the base acute to obtuse, apex acuminate, the acumen to 1.2 cm, sometimes folded to one side, the tip rounded, glabrous and verruculose above, very sparsely covered with golden, stellate trichomes below, venation weakly brochidodromous with 7–10 secondary veins per side, with occasional intersecondary veins, midvein raised above and below, secondary veins slightly raised above and below, tertiary venation finely reticulate, slightly raised above and below; petiole 3–5 mm, 0.7–1 mm in diam., moderately densely covered with semi-appressed, curly light golden trichomes. Flowers solitary, axillary, pendent; pedicels 7–15 mm long, 0.2–0.7 mm in diam., densely covered with golden erect, simple and fasciculate trichomes; epicalyx with 5 free lobes, appearing somewhat succulent in vivo, the lobes 9 × 5 – 7 mm, ovate, the apex acute, sparsely covered with golden stellate trichomes on the outside, glabrous very sparsely covered with golden simple and stellate trichomes inside; calyx tubular with 4(–5?) lobes, fused for 5–7 mm, the lobes 2.5 × 2.5 mm, triangular, the apex triangular, moderately densely covered with golden stellate trichomes outside and inside; petals 5, 24 × 13 mm, obovate, the apex rounded to emarginate, red, sparsely covered with very light golden stellate trichomes outside, glabrous inside with evident venation; staminal column 20 mm long, 0.8 mm in diam., pinkish red, glabrous with very scattered minute trichomes; stamens 30–40(–50), filaments 8–10 mm, red, anthers 0.8 × 0.4 mm; styles 5, 8 mm, red, moderately densely covered with white trichomes, stigma c. 0.7 mm in diam., urceolate. Fruit unknown. Distribution and ecology. – Hibiscus ankeranensis is known only from a single collection near the Andrangato river at Ankerana in the eastern coast of Madagascar at c. 350 m in elevation in lowland moist evergreen forests (Fig. 2). [A: Ratovoson 2092; B: Antilahimena 7544; C: Razafitsalama 1409] [Photos: A: F. Ratovoson; B: P. Antilahimena; C: C. Birkinshaw] Conservation status. – Hibiscus ankeranensis is known only from a single collection from the newly designated Corridor Ankeniheny-Zahamena protected area. We lack information on population size but lowland moist evergreen forests (below 800 m) are fragmented and heavily deforested in the region (GAUTIER, 2018). Recent satellite imagery from Google earth [https://www.google.com/intl/en/earth] suggests that the collection site and the surrounding area is deforested or disturbed. Hibiscus ankeranensis is therefore assessed as “Critically Endangered” [CR B 2ab(iii)] using IUCN Red List Categories and Criteria (IUCN, 2012). Notes. – Hibiscus ankeranensis is similar to H. calyculatus in corolla color but differs by its epicalyces with 5 lobes, 9 mm long (vs. 4(–5) lobes, reflexed, 3–4 mm long), calyx with 4(–5) teeth, green, succulent in vivo (vs. 5 teeth, red, inflated) and petals not reflexed (vs. reflexed).Published as part of Hanes, Margaret M., Schatz, George E. & Callmander, Martin W., 2020, Transfer of the Malagasy genera Humbertianthus and Macrostelia to Hibiscus (Malvaceae) with description of four new species, pp. 193-202 in Candollea 75 (2) on pages 200-201, DOI: 10.15553/c2020v752a4, http://zenodo.org/record/568374
Hibiscus involucratus M. Hanes, G. E. Schatz & Callm. 2020, comb. nov.
No full textHibiscus involucratus (Hochr.) M. Hanes, G.E. Schatz & Callm., comb. nov. Macrostelia involucrata Hochr. in Notul. Syst. (Paris) 14: 230. 1952. Holotypus: MADAGASCAR. Reg. Atsimo-Atsinanana [Prov. Fianarantsoa]: Vondrozo, [22°49'S 47 °19'E], 11.IX.1926, fl., Decary 5257 (P [P00037126]!; iso-: G [G00014442]!, P [P00365083, P00037127]!, TEF). Distribution and ecology. – Hibiscus involucratus is known from the low elevation tropical evergreen forests around Vondrozo at c. 450 m elevation (Fig. 2). Conservation status. – Hibiscus involucratus is known only from a single location near Vondrozo, collected in 1926. It is possible that the collection has been made in what is now known as the Ambositra-Vondrozo protected area. With a single collection in the low elevation moist evergreen forests, which are highly threatened in Madagascar, this species is assigned a conservation status of “Critically Endangered” [CR B2ab(iii)] using the IUCN Red List Categories and Criteria (IUCN, 2012). Notes. – Hibiscus involucratus can be distinguished from all other pendent Hibiscus in Madagascar by its tubular epicalyx with 6 irregular teeth that completely hides the calyx. We consider the sheet [P00037126] as the holotype, as it is the only specimen with Hochreutiner’s handwriting in P.Published as part of Hanes, Margaret M., Schatz, George E. & Callmander, Martin W., 2020, Transfer of the Malagasy genera Humbertianthus and Macrostelia to Hibiscus (Malvaceae) with description of four new species, pp. 193-202 in Candollea 75 (2) on page 196, DOI: 10.15553/c2020v752a4, http://zenodo.org/record/568374
Hibiscus ambanitazensis M. Hanes & G. E. Schatz 2020, sp. nov.
No full text<p> <i>Hibiscus ambanitazensis</i> M. Hanes & G.E. Schatz, <b>sp. nov.</b> (Fig. 1C).</p> <p> <b>Holotypus: MADAGASCAR. Reg. Antsiranana [Prov. SAVA]:</b> Com. Ampahana, Fkt. Andrapengy, Ambanitaza, 14°41'02"S 50°11'14"E, 9.XII.2015, fl., <i>Razanatsoa</i>, <i>Bernard & Mashburn</i> 644 (MO-6710358!; iso-: P, TAN).</p> <p> <i>Hibiscus ambanitazensis M. Hanes & G.E. Schatz is distinct from all other reddish, pendent flowered Hibiscus from the east coast of Madagascar in its petal pigmentation (purple red), and the combination of a long pedicel (up to 42 mm), epicalyx bracts with 6 lobes (fused at the base), and calyx fused to at least half of length (fused for 7–11 mm).</i></p> <p> <i>Shrub</i> c. 3.5 m tall, sparsely branched; twigs covered with golden, semi-erect, simple, fascicled and stellate trichomes. <i>Leaves</i> 5.5–9.8 × 2.8–4.6 cm, elliptic to narrowly obovate, the base acute to obtuse, apex acuminate, the acumen to 0.8 cm long, the tip rounded, glabrous above, very sparsely covered with golden stellate trichomes below, venation weakly brochidodromous with 8–10 secondary veins per side, midvein raised above and below, secondary veins slightly raised above and below, tertiary venation finely reticulate, slightly raised above and below; petiole 8–11 mm, c. 0.8 mm in diam., moderately densely covered with semi-appressed, curly, light golden trichomes and occasional light golden stellate trichomes. <i>Flowers</i> solitary, axillary, pendent; pedicels 40–42 mm long, c. 0.8 mm in diam., densely covered with golden, erect, simple, fasciculate, and stellate trichomes; epicalyx with 6 lobes fused at the base for 3–4 mm of its length, the lobes 6 × 2.5–4 mm, ovate, the apex acute, densely covered with golden stellate trichomes on the outside, densely covered with golden appressed simple trichomes inside; calyx with 5 lobes, fused at the base for 7– 11 mm, the lobes c. 15 × 10 mm, ovate to triangular, the apex narrowly triangular, very densely covered with golden stellate trichomes outside and inside; petals 5, 30 × 14 mm, obovate, the apex rounded, purple-red, strongly reflexed, densely covered with white stellate trichomes outside, glabrous and verrucose inside with visible venation; staminal column c. 21 mm long, c. 1.8 mm in diam., red, sparsely to moderately densely covered with light golden stellate trichomes, with a tuft of dense golden erect simple trichomes at the apex; stamens c. 70, filaments 8–10 mm long, red, anthers c. 1.3 × 1.3 mm; styles 5, 12 – 15 mm long, white, sparsely covered with white trichomes, stigma c. 0.8 mm in diam., urceolate. <i>Fruit</i> unknown.</p> <p> <i>Distribution and ecology. – Hibiscus ambanitazensis</i> is known from one collection in a forest fragment on volcanic soil (“table basaltique”) (Fig. 2).</p> <p> <i>Conservation status. – Hibiscus ambanitazensis</i> is known only by a single location outside of the Protected Area Network in a small patch of forest north of Antalaha in close proximity to recent slash and burn activity. The “table basaltique” along the east coast of Masoala to north of Antalaha are highly threatened because their rich volcanic soils are highly suitable for agriculture. The new species is assigned a conservation status of “Critically Endangered” [CR B2ab(iii)] using the IUCN Red List Categories and Criteria (IUCN, 2012).</p> <p> <i>Notes</i>. – The new species differs from all other reddish, pendent flowered <i>Hibiscus</i> from the east coast of Madagascar in its petal pigmentation, purple red (vs. red). Epicalyx, calyx and style color further distinguish <i>H</i>. <i>ambanitazensis</i> from <i>H</i>. <i>ankeranensis</i>, <i>H. calyculatus</i> and <i>H. laurinusus</i> (see Table 1).</p>Published as part of <i>Hanes, Margaret M., Schatz, George E. & Callmander, Martin W., 2020, Transfer of the Malagasy genera Humbertianthus and Macrostelia to Hibiscus (Malvaceae) with description of four new species, pp. 193-202 in Candollea 75 (2)</i> on pages 197-198, DOI: 10.15553/c2020v752a4, <a href="http://zenodo.org/record/5683749">http://zenodo.org/record/5683749</a>
Hibiscus vohipahensis M. Hanes & G. E. Schatz. Each 2020, sp. nov.
No full textHibiscus vohipahensis M. Hanes & G.E. Schatz, sp. nov. (Fig. 3C, 4C). Holotypus: MADAGASCAR. Reg. Atsimo-Atsinanana [Prov. Fianarantsoa]: Vangaindrano, Vohipaho, Ankara Bolava Forest, 23°31'34"S 47°30'01"E, 27 m, 26.VIII.2008, fl., Bussmann 15177 (MO-6152851!; iso-: G [G00341945]!, P, TAN). Hibiscus vohipahensis M. Hanes & G.E. Schatz differs from H. analalavensis M. Hanes & G.E. Schatz in having longer petioles and pedicels, large pure white petals that are widest at the apex with filaments about twice as long as the long staminal column. Tree 3–6 m tall, 6 cm dbh; twigs covered with light golden stellate trichomes. Leaves 7.8– 13.7 × 2.9– 4.5 cm, narrowly obovate, the base acute, the margin strongly revolute, the apex acuminate, glabrous above, nearly glabrous below with the occasional stellate trichome below, venation brochidodromous with 8–10 secondary veins per side, midvein raised above and below, secondary veins slightly raised above and below, tertiary venation reticulate, slightly raised above and below; petiole 6–10 mm, 1 mm in diam., covered with stellate trichomes. Flowers solitary, axillary, pendent; pedicels 45 mm, 1.2 mm in diam., densely covered with golden, erect, simple, fasciculate and stellate trichomes; epicalyx with 5 lobes fused at the base for 2 mm, the lobes 12–17 × 12–18 mm, broadly ovate, the apex acute to obtuse, densely covered with golden stellate trichomes on the outside, densely covered with light golden, erect simple and fasciculate trichomes inside; calyx with 5 lobes, fused at the base for 12 mm (about half of length), the lobes 20 × 18 mm, ovate, the apex acute, densely covered with golden stellate trichomes outside, densely covered with appressed curly beige trichomes inside; petals 5, c. 55 mm, obovate, the apex rounded, white, widest at apex, turning reddish with age, densely covered with light golden stellate trichomes outside, glabrous inside with evident venation; staminal column 50 mm long, 2 mm in diam., white, glabrous, with a tuft of dense golden erect simple trichomes at the apex; stamens c. 70, filaments 20 mm, white, anthers 1.5 × 2 mm; styles 5, 25 mm, white, slender, delicate, covered with white erect simple and fasciculate trichomes, stigma c. 1.7 mm, bulbous. Fruit unknown. Vernacular name. – “Hafatra-rora” (Birkinshaw 2040). Distribution and ecology. – Hibiscus vohipahensis has the most southern distribution of all species discussed here and is known from two protected patches of lowland moist evergreen forests near Vangaindrano from 10 to 135 m elevation. Conservation status. – Hibiscus vohipahensis is known from 6 locations from two protected areas (Agnakatrika and Ankarabolava). Despite the very restricted extent of occurrence (EOO) and area of occupancy (AOO) of c. 9 km ², there appear to be no current threats to H. vohipahensis. Thus, the species is assigned a status of “Least Concern” [LC] IUCN Red List Categories and Criteria (IUCN, 2012), with the caution that this status is highly dependent on continued effective protection. Notes. – Hibiscus vohipahensis differs from H. analalavensis in having longer petioles, 6 – 10 mm (vs. 4– 5 mm), longer pedicels, 45 mm (vs. 30–32 mm), and wider petals, 12–18 mm (vs. 6–9 mm) that are pure white (vs. white to pink blush) that are widest at the apex (vs. widest in the middle of the petal). The staminal column in H. vohipahensis is also longer than that in H. analalavensis, 50 mm (vs. 30 mm) with filaments about twice as long, 20 mm (vs. 10 mm). Paratypi. – MADAGASCAR. Reg. Atsimo-Atsinanana [Prov. Fianarantsoa]: Vangaindrano, Vohipaho, forêt d’Ankarabolava, 23°31'51"S 47°29'48"E, 72 m, 17.IX.2009, fl., Andriamihajarivo 1679 (MO, P, TAN, TEF); ibid. loco, 23°31'32"S 47°30'12"E, 13 m, 29.V.2014, bud, fl., Birkinshaw 2040 (MO, P, TAN); Marofototra, NAP Ankarabolava-Agnakatrika, 23°30'24"S 47°30'51"E, 65 m, 5.VII.2010, fl., Randrianarivony 164 (MO, P, TAN); c. 5 km au SE du village Sahavia, 23°26'21"S 47°30'34"E, 135 m, 19.X.2010, fl., Razafitsalama 1409 (MO, P, TAN); forêt de Vohipaho, 23°26'34"S 47°30'50"E, 80 m, 22.IX.2009, bud, Razanatsima 780 (MO, P, TAN).Published as part of Hanes, Margaret M., Schatz, George E. & Callmander, Martin W., 2020, Transfer of the Malagasy genera Humbertianthus and Macrostelia to Hibiscus (Malvaceae) with description of four new species, pp. 193-202 in Candollea 75 (2) on pages 201-202, DOI: 10.15553/c2020v752a4, http://zenodo.org/record/568374
Hibiscus calyculatus M. Hanes, G. E. Schatz & Callm. 2020, comb. nov.
No full textHibiscus calyculatus (Hochr.) M. Hanes, G.E. Schatz & Callm., comb. nov. (Fig. 1A). Macrostelia calyculata Hochr. in Notul. Syst. (Paris) 14: 232. 1952. Holotypus: MADAGASCAR. Reg. Haute Matsiatra [Prov. Fianarantsoa]: vallée Ampamaherana, [21°19'S 47°19'E], [1300 m], 25.V.1949, fl., Service Forestier 2046 (P [P00037125]!; iso-: G [G00014441]!, TEF). Vernacular name. – “Tsilaitra” (Service Forestier 2046, 14420). Distribution and ecology. – Hibiscus calyculatus grows in medium elevation moist evergreen forests east of Fianarantsoa, in the massif of Andrambovato, in the Ampamaherana valley and near Kianjavato (D. Scherberich, pers. comm.) from 1155–1300 m in elevation (Fig. 2). Conservation status. – Hibiscus calyculatus is known from 3 locations all encompassed in the Protected Area Network (Corridor Forestier Ambositra-Vondrozo and Ranofanana). Despite the very restricted Extent of Occurrence (EOO) of 12 km ² and an Area of Occupancy (AOO) of 8 km ², there appear to be no current threats to H. calyculatus. Thus, the species is assigned a conservation status of “Least Concern” [LC] using the IUCN Red List Categories and Criteria (IUCN, 2012), with the caution that this status is highly dependent on continued effective protection. [B: Rasoandriana 89; C: J. Razanatsoa 644] [Photos: A: D. Scherberich; B: P. Antilahimena; C: B. Mashburn] Notes. – Hibiscus calyculatus is distinguished from all other pendent Hibiscus on Madagascar by its four small (3–4 mm), free, reflexed epicalyx bracts and a distinctive red, tubular calyx. Additional specimens examined. – MADAGASCAR. Reg. Vatovavy-Fitovinany [Prov. Fianarantsoa]: Massif d’Andrambovato, Tolongoina, Ikongo, 21°30'58"S 47°25'36"E, 1155 m, 28.V.1998, fl., Razafimandimbison 335 (G, MO, P, TAN, TEX); Fort Carnot, Andrambovato Station Forestier, Parcelle A-16, [21°31'S 47°25'E], 28.VII.1954, fl., Service Forestier 14420 (P, TEF).Published as part of Hanes, Margaret M., Schatz, George E. & Callmander, Martin W., 2020, Transfer of the Malagasy genera Humbertianthus and Macrostelia to Hibiscus (Malvaceae) with description of four new species, pp. 193-202 in Candollea 75 (2) on pages 194-196, DOI: 10.15553/c2020v752a4, http://zenodo.org/record/568374
Trichoribates scilierensis Bayartogtokh & Schatz, 2008, sp. nov.
No full textTrichoribates scilierensis sp. nov. (Figs. 1-3) Trichoribates longipilis (part.): Schatz 1989, p. 120. Latilamellobates cf. oxypterus: Fischer & Schatz 2007, p. 436. Diagnosis. Rostrum broadly rounded in dorsal view, with a pair of minute lateral dens and a nose-like protuberance dorsally; rostral, lamellar and interlamellar setae barbed; lamella wide, with long and thin translamella; lamellar cusp with long lateral and medial dens; sensillus short, with elongate oval head; tutorium broad, with four or five small dentations at the dorso-distal end; 10 pairs of long and barbed notogastral setae; porose areas Aa large and round, A1 divided into two parts, A2 and A3 smaller, oval; all ventral setae barbed; epimeral setal formula 3-1-3-2; setae l" of tibiae and genua I, II very thick, much thicker than other setae; genu I with streng ventro-distal projection. Measurements. Holotype: body length 532 µm, length of notogaster 446 µm, width of notogaster 364 µm; paratypes (n = 24): body length 494-562 (527) µm; length of notogaster 410-471 (460) µm; width of notogaster 338-387 (362) µm. Integument. Body colour dark brown to yellowish brown. Surface of body and leg segments with thick cerotegument, roughened by small granules. Faintly microtuberculate on cuticle of prodorsum, notogaster, ventral plate, leg segments and subcapitular mentum. Microtubercles forming longitudinal Striae on femora III and IV. Prodorsum (Figs. 1A, C, 2A-F, H-K). Rostrum broadly rounded in dorsal view, with a pair of minute lateral dens and with a nose-like protuberance dorsally (about 14-18 um elevated above prodorsal shield) (Figs. 1A, C, 2A-C, E). Rostral seta (ro) barbed, inserted laterally, underneath of ventro-distal end of tutorium, 72- 87 µm in length, extending beyond tip of rostrum, directed mediad. Lamella wide, with longitudinal striations, 112-127 µm in length, slightly converging, connected by moderately long and very thin translamella, slightly concave medially. Lamellar cusp with long lateral and medial dens subequal in length (10-18 µm), their size slightly variable, also depending on abrasion in old specimens (Figs. 1A, 2A, D-F, H). Lamellar seta (le) 91- 104 µm in length, ciliate, slightly thinner than rostral setae. Interlamellar seta (in) 99-150 µm in length, barbed, not reaching the anterior margin of rostrum, insertions concealed under anterior margin of notogaster. Sensillus short, exposed portion 43-48 µm in length, with squamose elongate oval head. Bothridium relatively small, irregularly funnel-shaped, almost completely covered by notogaster. Tutorium broad, 162-176 µm in length, with distinct cusp distally, bears four or five small dentations at the dorso-distal end (Figs. 1C, 2I-K). Notogaster (Figs. 1A, C, D, 2G). Oval, about 1.3 x longer than wide. Anterior margin markedly arched anteriad; lenticulus present. Pteromorph well developed, curved ventrally. Ten pairs of notogastral setae present, ciliate, seta c longest (length 65-90 µm), other notogastral setae nearly equal in length. Porose area Aa largest (diameter 15-25 µm), almost round in shape; A1 divided into two parts, lateral area large, medial small (Figs. 1A, C, 2G). Porose areas A2 and A3 round to oval in shape, the latter slightly larger than the former. Lyrifissure im clearly visible in dorsal view, ia, ih and ips visible in lateral view, ip visible in posterior view, situated laterad A3. Opisthosomal gland opening (gla) located posterior to lyrifissure im. Gnathosoma (Figs. 1B, 2L, M). Subcapitular mentum conspicuously wider than long. Hypostomal setae a, h and m distinctly barbed, length of a 20-22 µm, h, m 30-32 µm long. Chelicera with strongly sclerotized blunt teeth; setae cha and chb barbed, length 40-50 µm. Palp typical for family, palpal setation 0-2-1-3-10 including both ventral setae and solenidion omega of tarsus. Epimeral region (Fig. 1B). Epimeral setae (except 1c) of medium length (15-20 µm), all finely barbed, seta 1c much longer (45-50 µm). Epimeral setal formula: 3-1-3-2, seta 4c not evident. Custodium reaching posterior margin of pedotectum II, discidium well developed, conspicuously projected laterally. Pedotectum I large, surface smooth, without striation. Ano-genital region (Figs. 1B, D, 2N). Anal aperture larger than genital one, anal and genital plates smooth. All genito-anal setae of medium length, barbed (genital, aggenital setae 10-12 µm, anal setae 18-22 µm, adanal setae 22-25 µm in length); setal formula 6-1-2-3. Distance between bases of aggenital setae nearly equal to that between setae ad3-ad3. Seta ad3 situated in paranal position. Adanal lyrifissure iad short, situated at same level as setae an2, adjacent to anterolateral margin of anal aperture. Postanal porose area (Ap) narrowly elongate, longer than distance between bases of adanal setae ad2 (Fig. 1D). Legs (Fig. 3). Lateral claws thinner than medial claw, claws of leg IV slightly longer and more slender than those of other legs. Tibia I with a relatively small, but distinct anterodorsal apophysis, bearing solenidion phi2. Solenidion phi1 on tibia I very long (115-140 µm), length of phi2 36-49 µm. Genu I with strong ventro-distal projection; femur and trochanter IV with distinct ventral keel or blade, distally rounded. Most leg setae distinctly barbed, except setae (p) and (u) on tarsi I-IV, and v on trochanter and femur IV Setae l" on tibiae and genua I, II very thick, much thicker than other setae, barbed. Formula of leg setation (including famulus): 1(1-5-3-4-20); II (1-5-3-4-15); III (2-2-1-3-15); IV (1-2-2-3-12); formula of solenidia: I (1-2-2); II (1-1-2); III (1-1-0); IV (0-1-0). Material examined. Holotype: Italy, Provincia di Bolzano, Schlern/Sciliar massif, alpine meadows (46°30,56 ’ N, 11°33,92 ’ E, 2450 m a.s.l., 05 August 2007: female, leg. Irene Schatz).Paratypes: ibid., alpine meadow below rock face (46°30,68 ’ N, 11°35,31 ’ E, 2220 m a.s.l., 24 May 2007, leg. Irene Schatz: 2 males, 14 females),ibid. (01 July 2007: leg. H. Schatz). Type deposition: The alcohol preserved adult holotype and 2 adult paratypes will be deposited in: Muséum d'histoire naturelle, Genève, Switzerland, other specimens in Museo di Scienze Naturali Alto Adige, Bolzano, Italy, and in authors' collections. Remarks. The new species is distinguishable from most other known species of Trichoribates by the long lateral and medial dens of the lamellar cusps. Moreover, in T. scilierensis the rostrum bears a nose-like protuberance, and porose area A1 is divided into two parts. Both characters are very rare in Trichoribates species. The division of porose area A1 is constantly observed in all examined specimens, although the shape and size of the two parts were slightly variable. The combination of these characters can be considered as a diagnostic feature of T. scilierensis sp. nov. Trichoribates biarea Gjelstrup & Solhøy, 1994, described from Iceland, resembles the new species in some characters, namely in having large porose areas Aa, divided porose areas A1, long notogastral setae, and similar shape of the tutoria (our attempt was unsuccessful in getting the type specimens of T. biarea from the host museum for comparative study). However, according to the original description and figures (Gjelstrup & Solhøy 1994), in T. biarea the rostrum is concave without a nose-like protuberance, the medial dens on the lamellar cusps are minute, the translamella is considerably thicker, the head of sensilli are more swollen, the lamellar and interlamellar setae are much longer, and the epimeral and ano-genital setae are smooth. Gjelstrup and Solhøy (1994) described the species T. biarea from dry heat-lands and fell-fields in Iceland. The new species is also morphologically similar to Trichoribates strigatus Mahunka & Mahunka Papp, 2006, described from Ticino and Appenzell, Switzerland (Mahunka & Mahunka Papp 2006), in having almost equal dens on lamellar cusps, long notogastral setae, dentate tutoria, and almost round and large porose areas Aa. Trichoribates strigatus is larger in body size (length 590-608 µm) than T. scilierensis, the rostrum of T. strigatus is irregularly undulate or serrate, the cuspidal dens are shorter, the head of sensilli are more swollen, the lamellar and interlamellar setae are much longer, the latter reach the anterior margin of the rostrum, and porose areas A1 are undivided. The North American species, T. spatulasetosus Reeves, 1967, has equal sized lateral and medial dens of lamellar cusps, but, T. spatulasetosus differs from T scilierensis in the densely barbed and strongly swollen head of sensilli; the very long and thick lamellar and interlamellar setae; strongly concave anterior margin of translamella; undivided porose area A1, and a smaller body size (length of females 472-495 µm, males 443- 450 µm, according to Reeves 1967). Trichoribates furcatus Schweizer, 1956, described from high alpine areas in the Swiss National Park, Grisons, Switzerland, resembles the new species by the presence of long dens on lamellar cusps, but in T. furcatus the lamellar cusps and the dens are extremely long, whereas the medial dens are longer than the lateral dens (Schweizer 1956). Also, in T. furcatus the interlamellar setae as well as the anterior notogastral setae are much longer than those in T. scilierensis. The body size of the Swiss species is larger (length 585-620 µm) than that of the new species. Schweizer (1956) also provided a figure of T. oxypterus, described by Berlese (1910) from Italy. This species also has relatively long dens on the lamellar cusps as in T. scilierensis, but in T. oxypterus the lateral dens of the cusps are much larger than the medial ones, while in the new species the size of medial and lateral dens is equal. Moreover, the interlamellar setae in the species studied by Schweizer (1956) are much longer (extending beyond the tip of rostrum) than those of the new species. It should be noted that T. oxypterus sensu Schweizer seems not to be the same species as T. oxypterus (Berlese, 1910). Mahunka and Mahunka-Papp (1995) examined the type specimens of T. oxypterus in the Berlese collection, giving a short diagnostic description and a figure of the prodorsum. The lateral dens of the lamellar cusps are very long, nearly as long as the basal part of cusps (much longer than that in Schweizer’s drawing), but the medial dens are absent; the lamellae and especially the translamella are much wider, the interlamellar and notogastral setae are relatively shorter, and the head of sensilli are shorter and more swollen compared to the clavate sensilli in Schweizer’s specimen. Shaldybina (1975) provided a figure of T. oxypterus, which is identical with that in the work of Schweizer (1956), but Mahunka and Mahunka-Papp (1995) noted that Shaldybina’s species is not identical with Berlese’s. Thus, it is obvious that Schweizer misidentified another species under the name of T. oxypterus. To clarify the status of T. oxypterus sensu Schweizer, the specimens in Schweizer’s collection will have to be studied. Another problem is the validity of T. oxypterus. Subías (2004) listed T. oxypterus as a junior synonym of Viracochiella (Latilamellobates) incisella, described by Kramer (1897). According to the diagnostic characteristics and the figures given by Mahunka and Mahunka-Papp (1995), it is obvious that T. oxypterus is not a junior synonym of Kramer’s species. Therefore, we reject the synonymy by Subías (2004) and retain the validity of T. oxypterus. Etymology. The name of the new species is derived from the term "Scilier", the name of the beautiful Schlern/Sciliar massif in the ladin language (Rhaeto-Romance), spoken by about 35000 people in the Dolomites area. Distribution. Trichoribates scilierensis was found in large numbers in the Schlern/Sciliar area in the Southern Alps in subalpine and alpine meadows between 2200 and 2650 m a.s.l. (summit of Mount Petz) (Schatz 2008). The species was also found on Plattkofel/Sasso Piatto by Fischer and Schatz (2007) at 2200 m a.s.l., under the name “ Latilamellobates cf. oxypterus ”.This species was also recently recorded in Italy, Trentino, Mezzano: Lago Giarine southeast of the Schlern/Sciliar massif (2126 m a.s.l., 21 September 2005, leg. R. Gerecke: 1 ad.). In material collected previously by one of us (H. S.) it became apparent that this species also occurs in other areas in the Central Alps: Austria, East Tyrol, Kalser Dorfertal – Daberklamm, with loose gravel and cushion plant vegetation (1520 m a.s.l., 17 July 1988, leg. H. Schatz, 6 ad.), sub Trichoribates longipilis (Schatz 1989); Austria, Tyrol, Ischgl – Idalpe, 2570 m a.s.l., in Caricetum curvulae (24 July 1991, leg. H. Schatz). By checking specimens from the collection of the Institute of Zoology, University of Innsbruck, we found two slides with Trichoribates scilierensis sp. nov., labeled as "slide 47: K. Schmölzer, Innsbruck near D8 leg. Janetschek ": 5 females, 4 males, and "slide 66 Schmölzer near D8 leg. Janetschek ": 8 females, 3 males. Unfortunately, we could not find records of these labels. Both Janetschek and Schmölzer collected in different parts of the Alps (e.g. Central Alps – Stubai and Zillertal mountains, Southern Alps – Dolomite mountains). These records indicate that Trichoribates scilierensis sp. nov. seems to have a distribution in a wider area of the Central and Southern Alps, but it is restricted to high altitudes. The specimens from East Tyrol in the comparatively low Daberklamm (1520 m) were found in an isolated site with alpine pioneer vegetation (Schatz 1989).Published as part of Bayartogtokh, B. & Schatz, H., 2008, Trichoribates and Jugatala (Acari: Oribatida: Ceratozetidae) from the Central and Southern Alps, with notes on their distribution, pp. 1-35 in Zootaxa 1948 on pages 2-
Fig. 4 in Transfer of the Malagasy genera Humbertianthus and Macrostelia to Hibiscus (Malvaceae) with description of four new species
No full textFig. 4. – Photographs of epicalyx and calyx of Hibiscus L. A. Hibiscus analalavensis M. Hanes & G.E. Schatz; B. Hibiscus ankeranensis M. Hanes & G.E. Schatz; C. Hibiscus vohipahensis M. Hanes & G.E. Schatz.Published as part of Hanes, Margaret M., Schatz, George E. & Callmander, Martin W., 2020, Transfer of the Malagasy genera Humbertianthus and Macrostelia to Hibiscus (Malvaceae) with description of four new species, pp. 193-202 in Candollea 75 (2) on page 201, DOI: 10.15553/c2020v752a4, http://zenodo.org/record/568374
Fig. 3 in Transfer of the Malagasy genera Humbertianthus and Macrostelia to Hibiscus (Malvaceae) with description of four new species
No full textFig. 3. – Photographs of flowers of Hibiscus L. A. Hibiscus analalavensis M. Hanes & G.E. Schatz; B. Hibiscus ankeranensis M. Hanes & G.E. Schatz; C. Hibiscus vohipahensis M. Hanes & G.E. Schatz.Published as part of Hanes, Margaret M., Schatz, George E. & Callmander, Martin W., 2020, Transfer of the Malagasy genera Humbertianthus and Macrostelia to Hibiscus (Malvaceae) with description of four new species, pp. 193-202 in Candollea 75 (2) on page 199, DOI: 10.15553/c2020v752a4, http://zenodo.org/record/568374
Beguea borealis G.E. Schatz & Lowry, spec. nova
No full text6. Beguea borealis G.E. Schatz & Lowry, spec. nova (Fig. 8). Typus: MADAGASCAR . Prov. Antsiranana: DIANA Region, Montagne des Français, forêt d’Ampitiliantsambo, à 3 h. de marche à pied, à l’E d’Andranomanitra, 12̊23’13’’S 49̊23’04’’E, 205 m, 14.I.2005, fl., Randrianaivo et al. 1163 (holo-: P [P04765740]!; iso-: MO, TAN!). Haec species a congeneris caulibus petiolis petiolulisque indumento aureo dense tomentulosis atque foliolis conduplicatis 4 ad 6 venis secundariis in quoque latere 7 ad 10 distinguitur. Tree c. 12 m tall, bole c. 15 cm in diam.; stems densely golden tomentulose, glabrescent. Petioles 1.7-3.6 cm, densely golden tomentulose. Leaves with 4-6 subopposite to opposite leaflets; rachis 0.2-2.1 cm, not keeled, densely golden tomentulose; petiolule 2-4 mm, light gray-tan tomentulose; leaflet blade 3.3-4.6 × 1.6-2.4 cm, elliptic (occasionally very slightly ovate or obovate), subcoriaceous, nearly always conduplicate and folded along the midvein in pressed material, glabrous and glossy above, glabrous below except very sparsely tomentulose at the base, base symmetrical, cuneate to acute, margins flat, minutely thickened but not revolute, apex acuminate, the acumen rounded, midvein flat above, sparsely tomentulose, raised below, sparsely tomentulose towards the base, secondary veins 7-10 per side, slightly raised above and below, tertiary venation raised on both surfaces. Inflorescence axis 1.5-9.7 cm long, c. 1 mm in diam. at base, unbranched, sparsely tan sericeous; bracts c. 0.2 mm, triangular, adaxially concave; pedicels usually paired on a common peduncle to 1 mm, sometimes solitary and epedunculate, 0.5-1.5 mm, moderately to densely tan sericeous. Male flowers with a cupulate, 5- to 7-lobed calyx, the lobes 0.2-0.3 × 0.3-0.6 mm, broadly triangular, sparsely tan sericeous outside; disc glabrous to sparsely white granular-farinose; stamens 8 (or 9), filaments c. 2 mm, with sparse spreading trichomes along basal third, anthers 0.7-0.8 × 0.3-0.4 mm, oblong; pistillode 0.2-0.3 mm high, 0.2-0.3 mm in diam., hemispherical, covered with short appressed trichomes. Female flowers not seen. Fruit not seen. Etymology. – The epithet borealis refers to the northernmost occurrence of the genus Beguea in Madagascar. Vernacular name. – “Gavoala”. Conservation status. – While Beguea borealis occurs within the recently decreed Montagne des Français (Ambohitr’Antsingy) protected area, it was collected adjacent to a clearing used as pasture and subject to potential burning, threats that may continue despite the new protected status. Therefore, with a restricted AOO and plausible threats that could rapidly push the species to Critically Endangered or Extinct status, B. borealis warrants a preliminary assessment of “Vulnerable” [VU D2] using the IUCN Red List Criteria (IUCN, 2012). Fig. 8. – Begueo boreolis G.E. Schatz & Lowry. A. Flowering branch; B. Leaflet (lower surface); C. Portion of inflorescence axis; D. Two flowers (one with filaments only partially expanded); E. Detail of flower at anthesis. [Rondrionoivo et ol. 1163, P]. [Drawing: R.L. Andriamiarisoa] Notes. – Beguea borealis is known only from the type specimen from the Forêt d’Ampitiliantsambo within the Montagne des Français massif (Fig. 4). It can be distinguished from B. apetala, with which it shares conduplicate leaflets that are usually folded along the midvein in pressed material, by its leaflets with a symmetrical base, the margins flat and minutely thickened but not revolute (vs leaflets with a distinctly asymmetrical base and strongly revolute margins) (Fig. 8).Published as part of George E. Schatz, Roy E. Gereau & Porter P. Lowry Ii, 2017, A revision of the endemic Malagasy genus Beguea (Sapindaceae), pp. 45-65 in Candollea 72 (1) on pages 56-58, DOI: 10.15553/c2017v721a6, http://zenodo.org/record/88817
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