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    Review of Taophila, a genus endemic to New Caledonia (Coleoptera: Chrysomelidae: Eumolpinae)

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    Samuelson, Allan (2010): Review of Taophila, a genus endemic to New Caledonia (Coleoptera: Chrysomelidae: Eumolpinae). Zootaxa 2621: 45-62, DOI: 10.5281/zenodo.19810

    Arsipoda Erichson 1842

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    Genus Arsipoda Erichson, 1842 Arsipoda Erichson, 1842: 235. Eratosthenes Clark, 1864: 261. (synonymized by Chapuis 1875)Published as part of Samuelson, Allan & Biondi, Maurizio, 2016, Taxonomic revision of the genus Arsipoda Erichson, 1842 (Coleoptera, Chrysomelidae) in New Caledonia, pp. 1-61 in European Journal of Taxonomy 230 on page 4, DOI: 10.5852/ejt.2016.230, http://zenodo.org/record/384040

    Fig. 16 in Taxonomic revision of the genus Arsipoda Erichson, 1842 (Coleoptera, Chrysomelidae) in New Caledonia

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    Fig. 16. Strict consensus tree with jackknife values (see text).Published as part of Samuelson, Allan & Biondi, Maurizio, 2016, Taxonomic revision of the genus Arsipoda Erichson, 1842 (Coleoptera, Chrysomelidae) in New Caledonia, pp. 1-61 in European Journal of Taxonomy 230 on page 54, DOI: 10.5852/ejt.2016.230, http://zenodo.org/record/384040

    Fig. 2. Habitus. A in Taxonomic revision of the genus Arsipoda Erichson, 1842 (Coleoptera, Chrysomelidae) in New Caledonia

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    Fig. 2. Habitus. A. Arsipoda longifrons sp. nov, Humboldt S track. B. A. montana sp. nov, Mt Humboldt. C. A. paniensis sp. nov, Mt Panié. D. A. povilaensis sp. nov, Pic d'Amoa. E. A. punctata sp. nov., Mueo. F. A. rostrata Gómez-Zurita, 2010, Koghi Mts. G. A. rutai sp. nov, Poro Plateau. H. A. shirleyae Samuelson, 1973, S Humboldt. I. A. transversa sp. nov., Mt Koghi.Published as part of Samuelson, Allan & Biondi, Maurizio, 2016, Taxonomic revision of the genus Arsipoda Erichson, 1842 (Coleoptera, Chrysomelidae) in New Caledonia, pp. 1-61 in European Journal of Taxonomy 230 on page 15, DOI: 10.5852/ejt.2016.230, http://zenodo.org/record/384040

    Fig. 6. A–C. Elytra. A in Taxonomic revision of the genus Arsipoda Erichson, 1842 (Coleoptera, Chrysomelidae) in New Caledonia

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    Fig. 6. A–C. Elytra. A. Arsipoda atra sp. nov., Aoupinié. B. A. geographica Gómez-Zurita, 2010, Dzumac Mts (base). C. A. shirleyae Samuelson, 1973, Chute de la Madeleine. D. Prosternum of A. punctata sp. nov., Mueo. E. Hind tibia and tarsus of A. communis sp. nov., Dzumac Mts (1) and A. shirleyae Samuelson, 1973, Pic du Pin (2). F. Metafemoral spring of A. communis sp. nov., Dzumac Mts. Abbreviations: ae = apical emargination with thickset spines; asp = apical spur of hind tibia; ba = basal angle of ventral lobe; dba = dorsal-basal angle of metafemoral spring; dl = dorsal lobe; dmv = dorsal margin of ventral lobe; ea = extended arm of dorsal lobe; fm = first metatarsomere; ht = hind tibia; pas = prosternal antecoxal space; pc = procoxal cavities posteriorly open (arrows); pip = prosternal intercoxal process; rf = recurve flange; sbe = spring basal edge; vl = ventral lobe.Published as part of Samuelson, Allan & Biondi, Maurizio, 2016, Taxonomic revision of the genus Arsipoda Erichson, 1842 (Coleoptera, Chrysomelidae) in New Caledonia, pp. 1-61 in European Journal of Taxonomy 230 on page 29, DOI: 10.5852/ejt.2016.230, http://zenodo.org/record/384040

    Arsipoda yiambiae Samuelson 1973

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    Arsipoda yiambiae Samuelson Figs 3B, 5F, 10C, 11R, 15A Arsipoda yiambiae Samuelson, 1973: 132. Arsipoda yiambiae – Gómez-Zurita et al. 2010: 2560. Type material examined NEW CALEDONIA: holotype, ³, Yiambi, NE, [-20,47008 164,70042], 500–700 m, 14 Oct. 1967 (BPBM). Taxonomic notes Arsipoda yiambiae and A. paniensis sp. nov. are the only species with a pronotum clearly convex laterally (Figs 4F, 5F), but they are clearly different in color and size: A. paniensis sp. nov. is dark green, metallic, and larger; A. yiambiae is yellow and smaller. Genitalia are also easily distinguishable (Figs 9D, 10C, 11K, R). Median lobe of aedeagus slender (Fig. 10C); parallel in ventral view, distinctly enlarged at basal third, with triangular apical part; ventral sulcus thin and shallow; in lateral view median lobe curved at basal third, then straight; dorsal ligula narrow, fusiform. Spermatheca (Fig. 11R) with rounded basal part; collum very elongate (as long as basal part); length of apical part about half as long as collum; ductus elongate, thickset, apically inserted. Biometry Male (holotype): LE = 1.65 mm; WE = 1.04 mm; LP = 0.48 mm; WP = 0.82 mm; LAN = 1.12 mm; LAED = 1.05 mm; LB = 2.15 mm; LE/LP = 3.44; WE/WP = 1.27; WP/LP = 1.71; WE/LE = 0.63; LAN/ LB = 0.52; LE/LAED = 1.58. Distribution Collected in Northern and Central Grande Terre (Fig. 15A).Published as part of Samuelson, Allan & Biondi, Maurizio, 2016, Taxonomic revision of the genus Arsipoda Erichson, 1842 (Coleoptera, Chrysomelidae) in New Caledonia, pp. 1-61 in European Journal of Taxonomy 230 on pages 49-50, DOI: 10.5852/ejt.2016.230, http://zenodo.org/record/384040

    Taophila millei Samuelson, new species

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    Taophila millei Samuelson, new species Fig. 2 f, 3 h Description. Female (Holotype). Head dark castaneous but paler on lower frons and central to upper vertex; prothorax dull dark castaneous with a slight brassy lustre; elytron orange-fuscous. Antenna, pterothorax, abdominal ventrites, and legs largely pitchy orange-fuscous. Dorsal surfaces: pronotum dulled by close puncturation; elytron subshining with slight isodiagramatic sculpture. Dorsal vestiture of conspicuous silvery setae. Head: frontal surfaces essentially flattened but interantennal region very weakly and gradually raised transversely; frons bearing small deep punctures, mostly about 1 x as large as interspaces; lower vertex with similar but larger punctures; punctures of upper vertex becoming elliptical at sides and above and median line of disc fine but distinct; eye rather small, with maximum diameter about 0.4 x as broad as interantennal space; gena about 0.55 x as deep as eye. Antenna extending beyond mid-elytron; pedicel exceeding half-length of segment 3; segments rather slender but with apical 5 distinctly heavier and longer. Prothorax roughly 0.7 x as broad as long, broadest across anterior third; side convex anteriorly, thence becoming straightened and narrowed to projecting posterior angle; basal antimarginal sulcus a deep narrow scratch; discal punctures small and ± elliptical, commonly 3 or 4 x as large as briefly raised interspaces, roughly tending to run in longitudinal directions; marginal punctures at side slightly larger and more circular. Elytron broadest across anterior third, the side rather evenly convex and gradually narrowed to apex; apex weakly declined in lateral view; humerus briefly swollen and followed by slightly swollen interstices; inner basal discal quadrant with punctures largest and confused; puncturation becoming regular apically and laterally; discal sculpture generally subgranulate-shining at most in inner discal quadrant, otherwise nearly smooth. Ventral surfaces: propleuron set with deep punctures mostly less than 1 x as large as smooth interspaces; metasternum subgranulate-smooth, with large punctures at side; abdominal ventrites 1 and 2 modified: ventrite 1 shield-like with apical margin bituberculate; ventrite 2 normally short but with apical margin bituberculate as in preceding; intercoxal piece of ventrite 1 fairly narrow at 11: 68 as broad as segment. Legs: metafemur rather slender but incrassate; metatibia ± straight, apex expanded. Genitalia: spermatheca with receptacle slender vlike, somewhat sinuate on one arm. Measurements: Body length 3.75 mm; body breadth 1.8 mm; head breadth 102 cmm; interantennal space 36 cmm; interocular space 68 cmm; eye 27 cmm; gena 15 cmm; pronotal length 102 cmm; pronotal breadth 140 cmm; elytral length 2.8 mm. Holotype Ψ (BPBM 17,188). NEW CALEDONIA. Col d’Amieu, 750 m, 3.iii. 1960, J.L. Gressitt. Remarks. A larger bicolorous species, with dorsal surfaces moderately setose. Differs from the smaller bicolorous species by having the elytral disc normally turgid and evenly convex instead of flattened with barely any granulation on the inner disc as compared to the other females. Differs further from T. scorpii, n. sp. by having a more evenly narrowed elytron and larger size. Unique female. The name honors Christian Mille of IAC in La Foa, provider of material critical to this study.Published as part of Samuelson, Allan, 2010, Review of Taophila, a genus endemic to New Caledonia (Coleoptera: Chrysomelidae: Eumolpinae), pp. 45-62 in Zootaxa 2621 on page 58, DOI: 10.5281/zenodo.19810

    Taophila deimos Samuelson, new species

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    <i>Taophila deimos</i> Samuelson, new species <p>Fig. 3e</p> <p> <b>Description.</b> Male (Holotype). Dorsum pitchy castaneous with slight violaceous lustre, head and pronotum darkest, almost black. Antenna orange-testaceous on 1–5, 4–6 more reddish, 7–11 darkest, castaneous. Ventral surfaces and legs largely pitchy castaneous. Pronotum appearing granose being closely and deeply punctulate in contrast to a smoother elytron; elytral punctures small, deep and not too conspicuous between swollen, pustulate interspaces. Dorsal surfaces moderately setose, the silvery setae fine and erect.</p> <p>Head: frontal surfaces generally flat and continuous through interantennal region; frons rather deeply punctate with interspaces smooth, punctures a little larger above, though still small; vertex similarly punctate and with fairly distinct median line; eye with maximum diameter about 0.55 x as broad as interantennal space; gena about 0.4 x as deep as eye. Antenna extending to basal 2/5 of elytron, apical 5 segments heavier than preceding ones. Prothorax about 0.80 x as long as broad, broadest just before middle; side convex anteriorly, then evenly narrowed from widest convex point to produced posterior angle; base beaded with antimarginal impression; disc closely and densely punctulate, appearing rough at low magnifications, punctures small and deep with interspaces actually smooth-shining and raised at higher magnifications; surface densely setose. Elytron rather gradually and convexly narrowed to apex; humerus moderately produced; disc with basal quadrant confusedly punctate the punctures commonly 1 x as large as interspaces and 2– 3 x as large as pronotal ones, these punctures not prominent due to swollen, ± transversely pustulate interspaces; lateral punctures large and becoming more regular in lateral 3 rows; surfaces smooth-shining. Ventral surfaces: propleuron smooth-shining with deep punctures commonly 2 x as large as interspaces; metasternum shining and smiliarly punctate and with surface with suberect setae; abdominal ventrites shining but with fine microsculpture; intercoxal piece of ventrite 1 fairly narrow at 9:60 breadth of segment; ventrite 1 in lateral profile gradually inclined to metathorax; remaining ventrites normal. Legs: femora moderately swollen medially; metatibia rather straight and briefly expanded at apex. Genitalia: aedeagus: length including base 102 cmm; narrowest breadth 18 cmm; preapex normally expanded with microtruncate extremity. Measurements: Body length 2.8 mm; body breadth 1.4 mm; head breadth 84 cm; interantennal space 30 cmm; interocular space 54 cmm: eye 30 cmm: gena 12 cmm; pronotal length 90 cmm; pronotal breadth 110 cmm; elytral length 2.0 mm.</p> <p>Holotype ɗ (BPBM 17,184). NEW CALEDONIA. Yahoue, iii.1978, N.L.H. Krauss.</p> <p> <b>Remarks.</b> A smaller dark castaneous species with violaceous lustre. Differing from congeners by the overall dark coloration and erect dorsal pubescence. Further differs from the variant of <i>T. subsericea</i> Heller with pustulate discal elytral interspaces by having the elytral apex evenly and convexly narrowed. Unique male. The name commemorates a satellite of planet Mars.</p>Published as part of <i>Samuelson, Allan, 2010, Review of Taophila, a genus endemic to New Caledonia (Coleoptera: Chrysomelidae: Eumolpinae), pp. 45-62 in Zootaxa 2621</i> on page 53, DOI: <a href="http://zenodo.org/record/198100">10.5281/zenodo.198100</a&gt

    Taophila scorpii Samuelson, new species

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    Taophila scorpii Samuelson, new species Fig. 2 e, 3 h, 4 f Description. Female (Holotype). Head pitchy fuscous becoming darker on sides of vertex; prothorax pitchy castaneous; elytron orange-testaceous. Antenna testaceous, intermediate segments fuscescent but without much contrast. Ventral surfaces and legs largely orange-testaceous but propleuron dark. Dorsal surfaces subshining granulate (isodiametric). Vestiture: pronotal disc with fine silvery setae; elytron with longer setae, these most conspicuous along side and apical region. Head: frontal surfaces generally raised transversely across interantennal region; frons deeply punctulate; lower vertex with larger punctures, commonly smaller than interspaces; upper vertex with punctures at sides more elliptical; discal median line obscure; eye with maximum diameter roughly 0.6 x as broad as interocular space; gena nearly 0.5 x as deep as eye. Antenna just exceeding mid-elytron; segments slender but apical 5 heavier; pedicel ovate and distinctly exceeding half-length of segment 3. Prothorax about 0.85 x as long as broad, broadest just before middle; side rather strongly convex anteriorly thence narrowed and ±straight to projecting posterior angle; base sulcate antimarginally; discal punctures small and deep, mostly about 1 x as large as interspaces. Elytron relatively robust, parallel-sided along basal 2 / 5, thence narrowed to fairly broad and sinuate preapex; apex slightly declined in lateral view; humerus slightly produced and followed by 2 weak costae, with the inner costa obliquely delimiting disc; disc with inner basal quadrant confusedly punctate, the punctures large; apical punctures smaller in ±irregular rows; apical part of lateral humeral costa swollen at preapex making apical area appear relatively broad with median emargination in dorsal view. Ventral surfaces: propleuron with deep punctures mostly smaller than interspaces, surface with slight granulation; metasternum with large shallow punctures laterally; abdominal ventrite 1 strongly modified, elevated over its length and exceeding half-length of abdomen, surface flattened to shallowly concave and extending to bituberculate apical margin; intercoxal piece fairly narrow at 11: 66; remaining ventrites appearing normal. Legs: metafemur slender, gradually thickened to about middle; metatibia rather straight, apex expanded. Genitalia: spermatheca with receptacle slender, v-shaped. Measurements: Body length 3.6 mm; body breadth 2.6 mm; head breadth 92 cmm; interantennal space 29 cmm; interocular space 52 cmm; eye 32 cmm; gena 15 cmm; pronotal length 102 cmm; pronotal breadth 122 cmm; elytral length 2.65 mm. Male (Allotype). Similar to holotype but pronotal surface a little smoother and elytral surface much smoother and shinier; elytral preapex normally closing to apex without sinuation or enlarged humeral costa above; ventrite 1 not overtly modified. Genitalia: aedeagal apex as in Fig. 3 h. Measurements: Body length 2.8 mm; body breadth 1.3 mm; head breadth 81 cmm; interantennal space 24 cmm; interocular space 48 cmm; eye 30 cmm; gena 12 cmm; pronotal length 90 cmm; pronotal breadth 98 cmm; elytral length 2.0 mm. Holotype Ψ (BPBM 17,190), NEW CALEDONIA. Col des Roussettes, 350–450 m, 3.ii. 1971, N.L.H. Krauss; Allotype ɗ (BPBM), same loc. but 450–550 m, 4–6.ii. 1963, J.L. Gressitt. Paratypes: same data as holotype (1 Ψ); same data as allotype but G. Kuschel (1 Ψ, 1 ɗ); Plateau de Dogny, 29.iii. 1968, sweeping, T.C. Maa (1 ɗ) and Maa & Gressitt (2 Ψ BPBM, MNHN); Col d’Amieu, 650 m, 31.iii. 1968, Gressitt (1 Ψ). Remarks. A smaller bicolorous species generally fitting with the smaller bicolorous species: T. corvi, n. sp. and T. subsericea Heller. The highly modified bituberculate female ventrite 1 and simple ventrite 2 separate this species from its close allies. Both sexes treated. Named for the constellation Scorpius.Published as part of Samuelson, Allan, 2010, Review of Taophila, a genus endemic to New Caledonia (Coleoptera: Chrysomelidae: Eumolpinae), pp. 45-62 in Zootaxa 2621 on pages 59-60, DOI: 10.5281/zenodo.19810

    Taophila mantillerii Jolivet, Verma

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    Taophila mantillerii Jolivet, Verma, and Mille Fig. 1 b, 3 b, 4 b Taophila mantillerii Jolivet et al. (2007) Revue française d’Entomologie (N.S.), 29 (1): 43–44, fig. 26 (aedeagus). A larger orange-fulvous species with brassy lustre; described from male but sexes rather similar dorsally, with surfaces smooth-granulate and generally punctate, the pronotal punctures much smaller and closer than the elytral ones. Vestiture inconspicuous except for fringing setae along elytral apex. Legs very slender, of weak appearance. Female (nov.). Similar to male. Abdominal ventrite 1 broadly concave and densely setose. Spermatheca as figured. Measurements: Body length 4.6 mm; body breadth 2.2 mm; head breadth 120 cmm; interantennal space 44 cmm; interocular space 78 cmm; eye 42 cmm; gena 22 cmm; pronotal length 120 cmm; pronotal breadth 140 cmm; elytral length 3.6 mm. Variation. Elytral coloration sometimes with a fuscescent darker streak extending longitudinally from the base. Material examined. NEW CALEDONIA Col des Roussettes, 450–550 m, 4–6.ii. 1963, C.M. Yoshimoto (5 Ψ, 1 ɗ); same data but 450–500 m, 5–6.ii. 1963 (2 Ψ, 2 ɗ); same loc., Piste Est, 600 m, 24.xi. 2002, beating Cyathea, A. Mantilleri & E.-A. Legun (1 ɗ paratype, JOLVET); Plateau de Dogny, 1.ii. 1963, N.L.H. Krauss (3 Ψ, 2 ɗ); same loc., 700 m, 1.ii. 1963, Yoshimoto (1 Ψ, 2 ɗ); Col d’Amieu, 500–600 m, 28.xii. 1976, J.L. Gressitt (1 Ψ); Sarramea/Col d’Amieu, 2–25 / 11 /05, S. Cazeres, C. Mille, J.P. Kataoui [Red label: Jolivet 2006] (1 Ψ IAC); 9.2 km NE of Col d’Amieu, slopes of Mt Rembai, 375–600 m, 23.ix. 1979, ferns, G.A. Samuelson (Ψ nov. MNHN); same data (2 ɗ); same data but 375–675 m, W.C. Gagne, G.M. Nishida & Samuelson (1 Ψ, 3 ɗ); Ponerihouen/Aoupinie, S 21.1038 E 165.1707, 945 m, 11 / 10 /06, Cazeres (49 mixed sexes, IAC); same data but Mille (42 mixed sexes, IAC). Parts of IAC Ponerihouen/Aoupinie series for deposit in BPBM, ISNB, JOLIVET, MNHN, NMNH, ONNC, and SMTD. Plant associates. Treeferns: Cyathea, also ferns at mid-montane elevations. Remarks. Unique in body form among congeners, the body breadth distinctively expanded across the mid-length of the elytra. The build of abdominal ventrite 1 in the female is distinctive, as is the c-shape of the spermatheca. In the male, the termination of the aedeagus is bifid and unique in the genus. Both sexes treated. PLATE 1. Dorsal view— Taophila, named species. a. subsericea Heller Ψ—body length = 3.55 mm; b. mantillerii Jolivet et al. ♂ —body length = 4.5 mm; c. nigrans Jolivet et al. Ψ—body length = 3.8 mm;. nigrans Jolivet et al. ♂— body length = 3.05 mm.Published as part of Samuelson, Allan, 2010, Review of Taophila, a genus endemic to New Caledonia (Coleoptera: Chrysomelidae: Eumolpinae), pp. 45-62 in Zootaxa 2621 on pages 47-48, DOI: 10.5281/zenodo.19810
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