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Phytoseius khowaiensis Kar & Karmakar 2022, sp. nov.
No full textPhytoseius khowaiensis sp. nov. (Figs 21–25, 56–59) Diagnosis. The dorsum of the new species is strongly reticulated, most of the dorsal setae serrated except setae j4, j5, j6, J5, z2 and z4 which are smooth. The inner margin of setae s4, s6, Z4 and Z5 with distinct longitudinal slit. Ventrianal shield sole-shaped with three pairs of pre-anal setae; fixed digit of chelicerae with two teeth and pilus dentilis and movable digit with two teeth; spermatheca cup shaped with bifurcated atrium; major duct wide and distinct; minor duct visible; leg IV with three macrosetae with spatulated tips. Description Female (n= 8) Dorsum (Fig. 21). Dorsal shield 278 (275–284) long and 143 (140–146) wide, strongly reticulated, prodorsum wide, with six pairs of solenostomes (gd1, gd2, gd4, gd5, gd8 and gd9); 15 pairs of setae on the dorsal shield: j1 20 (19–22), j3 25 (23–26), j4 8 (7–9), j5 8 (7–9), j6 8 (7–9), J5 6 (5–8), z2 14 (13–16), z3 27 (25–28), z4 11 (10–12), z5 8 (7–9), Z4 46 (45–48), Z5 59 (58–62), s4 68 (67–69), s6 69 (67–71), r3 33 (32–34). Peritreme (Fig. 21). Extending up to the base of j1. Venter (Fig. 22). All shields are smooth. The sternal shield square shaped, posterior margin indistinct, 50 (48–53) long and 61 (59–63) wide at level of setae st1–st3 and st3–st3 respectively, with three pairs of setae and two pairs of poriods; one pair of setae (st4) 10 (10–12) long on metasternal platelet with distinctly visible poroids. Distances between st2–st2 51 (50–54), st5–st5 63 (60–65). The base of the genital shield straight. One pair of metapodal platelets present 26 (25–28) long and 3 (2–4) wide. Genital shield smooth, broad at the level of st5 and narrow at genital opening with irregular circular flap. Ventrianal shield sole-shaped 108 (105–108) long, 40 (39–41) wide at level of ZV2 and 52 (51–54) wide at anus level with three pairs of pre-anal setae, JV1 16 (16–18), JV2 16 (16–18), ZV2 16 (16–18) and one pair of pores gv3 12 (12–15) apart below JV2. Membrane surrounding ventrianal shield with three pairs of setae ZV1 16 (15–18), ZV3 6 (5–9) and JV5 43 (43–46); the later one is long and serrated. Chelicera (Fig. 23). Fixed digit 24 (22–25) long with two teeth and distinct pilus dentilis, movable digit 24 (22–25) long with two teeth. Spermatheca (Fig. 24). The proximal part of the calyx short tubular and the distal arms of calyx forms a funnel shaped appearance 18 (15–19) long, atrium bifurcated, major duct distinct broad, minor duct visible. Legs (Fig. 25). The genu, tibia and tarsus of leg IV with macrosetae with spatulated tips and of following lengths – genu 15 (14–17), tibia 36 (33–37) and tarsus 22 (18–23). Chaetotactic formula of genu II: 1 2/1 2/0 1 and genu III 1 2/1 2/01. Length of leg I 242 (240–245), leg II 218 (217–225), leg III 236 (235–242) and leg IV 389 (387–394). Type specimens. Holotype: female (Acarol.lab/ BCKV/8598 /2021) deposited in the NZC, Zoological Survey of India, Kolkata, collected from star fruit tree, (Averrhoa carambola) at Khowai: 24°2'24"N, 91°35'56"E, 34m above AMSL, Khowai, Tripura, on 28th August, 2019; 10 paratype females (Acarol.lab/ BCKV/8598–8600 /2021) collected from same host with same collection data as holotype on 4 th February, 2021 and deposited in the Acarology laboratory of BCKV, Mohanpur. Etymology. The specific name khowaiensis refers to the type locality “Khowai” of Tripura state from where the species was collected. Remarks. The new species Phytoseius khowaiensis sp. nov. belongs to horridus species group with setae J2 and R1 absent and ventral setal pattern JV-3,4: ZV. Following the keys of Kar & Karmakar, 2021b, the new species is found different from all the Phytoseius species described and recorded from India but is very close to P. aonlae Kar & Karmakar, 2021b, where setae s4, Z4 and Z5, StiIV are shorter in new species than P. aonlae species. However, the new species is close to P. crinitus Swirski & Shechter, 1961, P. neoferox Ehara & Bhandhufalck, 1977, P. litoralis Silva et al., 2013 and P. aonlae Kar & Karmakar, 2021b by having similar dorsal setal pattern, setal slit along the longitudinal line of setae s4, s6, Z4 and Z5 and macrosetae of leg IV. But the new species differs from these close species by the dorsal setal length; denticles in fixed and movable digit of chelicerae; shape of ventrianal shield; length of macrosetae in leg IV; and shape form of spermatheca. All the differences between the new species and its close species are described in Table 5.Published as part of Kar, Anamika & Karmakar, Krishna, 2022, Seven new species of phytoseiid mites (Acari: Mesostigmata) from Tripura, northeastern India, pp. 213-241 in Zootaxa 5120 (2) on pages 226-228, DOI: 10.11646/zootaxa.5120.2.3, http://zenodo.org/record/638917
The histology of the euselachian-type dermal denticles and ?Helodus sp. tooth [VU-ICH-KAR-012] from the Karpėnai quarry
No full text(A) VU-ICH-KAR-014 from the
KAR-7 sample. (A1) General view of the dermal denticle; (A2)
general view of the longitudinal section of the denticle; (A3–A5)
enameloid covering the top surface of the crown. (B) VU-ICH-KAR-016 from the KAR-11
sample. (B1) General view of the dermal denticle; (B2) general
view of the longitudinal section of the denticle; (B3) enameloid
covering the top surface of the crown. (C1) The entire isolated fossil in
lateral view; (C2) in occlusal view; (C3) the
longitudinal sectioned tooth with a well-differentiated cap of enameloid and a distinct
enameloid-dentine junction (EDJ); (C4) in visual detail, the
penetrating canals of the enameloid layer indicated by the white arrow pointers;
(C5) in visual detail, some randomly arranged enameloid
crystallites. The white rectangles represent the outer layer of the crown in
detail. </p
Phytoseius baramuracus Kar & Karmakar 2022, sp. nov.
No full textPhytoseius baramuracus sp. nov. (Figs 26–30, 60–63) Diagnosis. The species is characterised by strongly reticulated dorsum; serrated dorsal setae except j4, j5, j6, z5 and J5; ventrianal shield vase shaped with three pairs of pre-anal setae; fixed digit of chelicerae with four teeth and movable digit with one tooth; leg IV with three macrosetae with bulbous tips. Description Female (n= 8). Dorsum (Fig. 26). Dorsal shield 304 (303–305) long and 128 (127–131) wide, rugose, prodorsum wide, with five pairs of solenostomes (gd1, gd4, gd5, gd8 and gd9); 15 pairs of setae on the dorsal shield: j1 24 (23–25), j3 18 (17–20), j4 8 (8–9), j5 8 (8–9), j6 10 (8–11), J5 8 (7–9), z2 10 (8–11), z3 21 (20–22), z4 25 (24–27), z5 10 (9–12), Z4 48 (47–50), Z5 50 (48–52), s4 43 (42–45), s6 58 (57–59), r3 28 (27–30). Most of the setae serrated except setae j4, j5, j6, z5, and J5 which are smooth. Peritreme (Fig. 26). Extending beyond the bases of j1. Venter (Fig. 27). All the shields are smooth. The sternal shield rectangular 54 (52–55) long and 63 (62–65) wide at level of setae st1–st3 and st3–st3 respectively, posterior margin indistinct, with three pairs of setae and two pairs of poroids; one pair of setae (st4) on metasternal platelet 16 (15–17) long. Distances between st2–st2 51 (50–53), st5–st5 60 (58–61). Base of genital shield broad, posteriorly truncated. One pair of metapodal platelets visible 28 (27–30) long. Genital shield smooth and narrow at genital opening with irregular circular flap. Ventrianal shield smooth, sole-shaped 93 (92–95) long, 48 (46–50) wide at level of ZV2 and 42 (40–44) wide at anus level with three pairs of pre-anal setae, JV1 15 (13–16), JV2 12 (11–13), ZV2 12 (11–13) and one pair of pre-anal pores gv3 26 (26–29) apart at the level of JV2. Membrane surrounding ventrianal shield with three pairs of setae ZV1 18 (16–19), ZV3 7 (6–10), and JV5 22 (20–23); seta JV5 long and smooth. Chelicera (Fig. 28). Fixed digit 23 (22–25) long with four teeth and a distinct pilus dentilis, movable digit 24 (22–25) long with one tooth. Spermatheca (Fig. 29). Arms of calyx 12 (12–15) long, bell shaped with nodular atrium, major duct flattened, long, minor duct clearly visible. Leg (Fig. 30). Leg IV with three bulbous macrosetae on tibia and tarsus and of following lengths: genu 7 (6–9); tibia 33 (31–35) and tarsus 20 (18–22). Chaetotactic formula of genu II 1 2/1 2/0 1 and genu III 1 2/1 2/01. Length of leg I 250 (248–254), leg II 216 (212–218), leg III 228 (225–232) and leg IV 374 (372–378). Etymology. The specific name baramuracus refers to the “Baramura Hill” of Tripura state from where this new species was collected. Sources of measurement: Original paper; Swirski&Shechter, 1961; Gupta, 2003; ‘–’: data not available Specimens collected. Holotype female (Acarol.lab/ BCKV/8595 /2021) collected from Bamboo, (Babusa sp.), at Subalsing, Baramura Hill: 24°0'20"N, 91°27'47"E, 59m AMSL, West Tripura, on 2 May, 2019; deposited in the NZC, Zoological Survey of India, Kolkata. Paratypes - nine females (Acarol.lab/ BCKV/8595–97 /2021) collected from same host and location and deposited in the Acarological laboratory, Department of Agricultural Entomology, BCKV, Mohanpur, Nadia, West Bengal, India. Remarks. The new species is distinguished from all the species of horridus species group by having three pairs of leg macrosetae and balloon shaped spermatheca. The new species is diffrerent from the species described and reported from India (Kar & Karmakar, 2021b) but close to P. brevicrinis Swirski & Shechter, 1961 by having almost similar type of spermatheca and P. jujuba Gupta, 1977b, P. meyerae Gupta, 1977b, P. neocorniger Gupta, 1977b by having similar dorsum. However, this new species differs from these close species by the dorsal setal length, number of teeth in fixed and movable digit of chelicera, form of setae, number and shape of metapodal shield, length and shape of leg macrosetae and shape of spermatheca. All the differences between the new species and the close species are presented in Table 6.Published as part of Kar, Anamika & Karmakar, Krishna, 2022, Seven new species of phytoseiid mites (Acari: Mesostigmata) from Tripura, northeastern India, pp. 213-241 in Zootaxa 5120 (2) on pages 229-231, DOI: 10.11646/zootaxa.5120.2.3, http://zenodo.org/record/638917
Amblyseius azaliae Kar & Karmakar 2021, sp. nov.
No full textAmblyseius azaliae sp. nov. (Figs 1–5; 58–62) Following the classification of Chant & McMurtry (2004), the new species Amblyseius azaliae sp. nov. belongs to the obtusus species group with the ventrianal shield pentagonal, seta z4 short and J2 present; and sub-group pamperisi having spermatheca with calyx swollen basally, then narrowing and finally flaring distally. Female (n = 3) Diagnosis. The species is distinguished by dorsal shield smooth with seven pairs of pores, female idiosomal setal pattern: 10A: 9B/JV–3: ZV, peritreme extends beyond seta j1, sternal shield with three pairs of setae, posterior margin of sternal shield concave, fixed digit of chelicera with 11 teeth and movable digit with three, spermatheca tubular with constriction at the distal part of calyx and then flared, all macrosetae on leg IV are pointed with longest one on genu of leg IV. Dorsum (Fig. 1). Dorsal shield 400 (396–401) long and 275 (273–276) wide, smooth, with sigillum mostly on prodorsal area, dorsum with seven pairs of pores (gd1, gd2, gd4, gd5, gd6, gd8 and gd9); 17 pairs of setae on dorsal shield and two pairs of setae on unsclerotised cuticle: j1 38 (36–39), j3 52 (51–53), j4 7 (6–8), j5 5 (4–6), j6 6 (6–8), J2 5 (5–6), J5 5 (5–6), z2 8 (8–10), z4 8 (7–9), z5 5 (4–6), Z1 8 (8–9), Z4 113 (111–113), Z5 238 (237–239), s4 75 (74–76), S2 11 (10–12), S4 10 (9–11), S5 10 (9–11), r3 13 (12–14), R1 14 (13–15). All setae smooth, setae j1, j3, s4, and Z4 long, seta Z5 longest, all other setae medium to minute. Peritreme (Fig. 1). Extending beyond the bases of j1. Venter (Figs 2; 59). Sternal shield square shaped, posterior margin of sternal shield concave. Sternal shield 73 (72–74) long and 86 (84–87) wide at level of setae st1-st3 and st3-st3 respectively, with three pairs of setae and two pairs of poroids; one pair of setae (st4) on distinct metasternal plate 27 (26–28) long with conspicuous poroids. Distances between st2-st2 83 (82–85), st5-st5 72 (71–74). Genital shield smooth, broad at the base and narrower at genital opening with anterior irregular circular flap. Ventrianal shield smooth, pentagonal 123 (122–125) long, 83 (82–84) wide at level of ZV2 and 64 (62–65) wide at anus level with three pairs of pre-anal setae, JV1 15 (14–16), JV2 18 (17–19), ZV2 16 (15–18) and one pair of crescentic pores gv3 17 (16–19) apart at the level JV2. Membrane surrounding ventrianal shield with four pairs of setae ZV1 15 (14–16), ZV3 14 (12–15), JV4 12 (11–15) and JV5 88 (86–89) and smooth. Two pairs of metapodal shields present. Primary inguinal sigilla 26 (25–27) long and 6 (5–7) wide, secondary ones 12 (10–12) long. Chelicera (Figs 3; 58). Fixed digit 36 (35–38) long with 11 teeth and a distinct pillus dentilis; movable digit 40 (39–41) long with three teeth. Spermatheca (Figs 4; 61–62). Calyx tubular 19 (18–20) long, arms of the calyx flared distally at vesicle, with narrow constriction. Atrium covers almost width of calyx or incorporated into the basis of calyx; major duct clearly visible, minor duct indistinct. Legs (Figs 5; 60). The genu, tibia and tarsus of leg IV with pointed macrosetae of following lengths: genu 111 (109–114), tibia 75 (74–76) and tarsus 70 (69–71). Genu and tibia of leg I, II and III with macrosetae of the following length respectively, SgeI 40, StiI 30, SgeII 30, StiII 25, SgeIII 50, StiIII 30. Chaetotactic formula of genu II 1 2/1, 1/1 1 and genu III 1 1/1, 2/1 1. Length of leg I 375 (373–376), leg II 255 (253–256), leg III 278 (276–280) and leg IV 354 (351–355). Type specimens. Holotype: female (Acarol.lab/ BCKV/8371 /2019) collected from Azalea, (Rhododendron sp.), at Upper Shillong: 25°32'09"N, 91°49'29"E, 1589 m AMSL, Ri-Bhoi, Meghalaya on 2 nd May, 2019; deposited in the NZC, Zoological Survey of India, Kolkata. Two paratype females (Acarol.lab/ BCKV/8372–8373 /2019) with same collection data as holotype deposited in the Acarological laboratory, Department of Entomology, BCKV, Mohanpur, Nadia, West Bengal, India. Etymology. The specific name azaliae refers to the host plant azalea (Rhododendron sp.) of the new species. Remarks. Amblyseius azaliae is different from all species under pamperisi sub-group by having its unique type of spermatheca, distal part of of calyx is constricted followed by flared arms. However, this new species is very close to A. orientalis Ehara, 1959; A. kulini Gupta, 1978c; A. raoiellus Denmark & Muma, 1989; A. pamperisi Papadopoulis, 1997; and A. euterpes Gondim Jr. & Moraes, 2001; by having similar type of spermatheca, leg IV with three setaceous macrosetae. However, the new species differs from these close species by the length of dorsal setae, number of teeth in fixed and movable digit of chelicera, length of leg macrosetae and shape of spermatheca. All of them have tubular spermathecae, but spermateca of the new species is very specific with a constriction in calyx and this combination of all characters makes it unique from all the other species of this group. All the differences between the new species and its close species are presented in Table 2.Published as part of Kar, Anamika & Karmakar, Krishna, 2021, Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India, pp. 301-354 in Zootaxa 5068 (3) on pages 306-309, DOI: 10.11646/zootaxa.5068.3.1, http://zenodo.org/record/570688
Euseius spontaneum Kar & Karmakar 2022, sp. nov.
No full textEuseius spontaneum sp. nov. (Figs 6–10, 41–44) Diagnosis. The new species is characterised by having the dorsum strongly reticulated; lateral margin at waist region irregular; R1 on the dorsal shield; peritreme extends extends up to base of j3; all ventral shields smooth, fixed digit of chelicerae with six distinct teeth and a pilus dentilis; spermatheca unique tubular, constricted at base of vesicle; leg IV with three macrosetae with blunt tips. Description Female (n=8). Dorsum (Fig. 6). Dorsal shield 298 (296–299) long and 198 (195–197) wide, strongly reticulated, with sigilla mostly on prodorsal area, prodorsum wide, irregular lateral margin at waist region with five pairs of solenostomes (gd2, gd5, gd6, gd8 and gd9); 18 pairs of setae on dorsal shield and one pair of setae on unsclerotised cuticle: j1 30 (28–30), j3 17 (15–18), j4 8 (7–9), j5 8 (7–9), j6 8 (7–9), J2 5 (3–6), J5 7 (5–7), z2 10 (9–11), z4 10 (9–12), z5 8 (7–9), Z1 8 (7–9), Z4 16 (15–18), Z5 56 (55–58), s4 20 (19–23), S2 19 (18–20), S4 22 (20–23), S5 26 (25–28), r3 13 (12–14), R1 10 (9–12). Seta R1 present on the dorsal shield. All setae smooth, setae j1, j3, Z4, Z5, s4, S2, S4 and S5 long, seta Z5 longest, all other setae medium to small. Peritreme (Fig. 6). Extending up to base of j3. Venter (Fig. 7). Sternal shield square shaped, posterior margin of sternal shield with loop at the posterior margin of sternal shield. Sternal shield 54 (53–56) long and 80 (79–82) wide at level of setae st1–st3 and st3–st3 respectively, with three pairs of setae and two pairs of poroids; one pair of setae (st4) 25 (24–26) long and conspicuous poroids on distinct metasternal platelets. Distances between st2–st2 75 (72–76), st5–st5 70 (70–73). Two pairs of metapodal platelets present. Primary metapodal platelet 25 (23–26) long and 3 (3–4) wide, secondary metapodal platelet 10 (9–10). Genital shield smooth, broad at base and narrow at genital opening with irregular circular flap touching sternal loop anteriorly. Ventrianal shield smooth, slightly sclerotised at anterior part 98 (97–101) long, 46 (46–49) wide at level of ZV2 and 68 (65–68) wide at anus level with three pairs of pre-anal setae, JV1 18 (16–19), JV2 20 (17–20), ZV2 22 (19–22) and one pair of large elliptical pre-anal pores gv3 just below JV2 29 (28–31) apart. Membrane surrounding ventrianal shield with four pairs of setae ZV1 13 (12–14), ZV3 8 (8–10), JV4 8 (8–10) and JV5 33 (32–35); seta JV5 long and smooth. Chelicera (Fig. 8) Fixed digit 22 (21–24) long with six teeth clubbed anteriorly and distinct pilus dentilis, movable digit 20 (20–21) long with one tooth. Spermatheca (Fig. 9). Calyx elongated 26 (24–27) and tubular, distinctly constricted at the distal part of calyx thereafter flared and gradually tapered at the proximal part towards the atrium with conspicuous major and minor duct. Arms of the calyx formed a funnel shaped appearance. Legs (Fig. 10). The genu, tibia and tarsus of leg IV with blunt hyaline tip macrosetae of following lengths: genu 40 (39–42), tibia 33 (32–34) and tarsus 58 (57–59). Chaetotactic formula of genu II 1 2/1 1/1 1 and genu III 1 2/1 0/2 1. Length of leg I 338 (332–340), leg II 275 (273–276), leg III 262 (260–265) and leg IV 325 (324–330). Leg III and leg II having macrosetae, Genu III 31 (29–32); genu II 18 (16–20). Type specimens. Holotype: female (Acarol.lab/ BCKV/8590 /2021) collected from Kash grass (Saccharum spontaneum), at Mungiakami, Atharamura hill: 23°53'19"N, 91°44'0"E, 118m AMSL, Tripura, on 28 th August, 2019 and deposited in the NZC, Zoological Survey of India, Kolkata; 3 paratype females (Acarol.lab/ BCKV/8591 /2021) with same collection data as holotype; 4 paratype females (Acarol.lab/ BCKV/8592 /2021) collected from unidentified shrub with same collection data as holotype; deposited in the Acarological laboratory, Department of Entomology, BCKV, Mohanpur, Nadia, West Bengal, India. Sources of measurement: Original paper; Gupta, 1978; Schicha and O’Dowd, 1993; Denmark et al., 1999; Ehara and Moraes, 1998 ‘–’: not available. Etymology. The specific name sponateum refers to the species name of the host plant, Saccharum spontaneum or Kash grass from which the type species was collected. Remarks. The new species is distinguished from all other species under the genus Euseius by its spermatheca, where calyx is tubular, distinctly constricted at the base of vesicle, flared distally and gradually tapered towards the atrium with conspicuous major and minor duct. This new species is also different from all the Euseius species described and reported from India (Kar & Karmakar, 2021b) but close to E. neococciniae Gupta, 1978; E. dowdi (Schicha, 1993 in Schicha &O’Dowd, 1993); E. lasalasi Denmark & Evans, 1998 in Denmark et al., 1999; E. inouei (Ehara & Moraes, 1998), by having similar type of spermatheca. However, the new species differs from these close species by the dorsal shield reticulated versus smooth, length of dorsal setae (long versus short), lateral margin of dorsal shield (regular versus irregular), position of seta R1 (on the shield versus off the shield), number of teeth in fixed and movable digit of chelicera and leg macrosetae variable. All the differences between the new species and its close species are presented in Table 2.Published as part of Kar, Anamika & Karmakar, Krishna, 2022, Seven new species of phytoseiid mites (Acari: Mesostigmata) from Tripura, northeastern India, pp. 213-241 in Zootaxa 5120 (2) on pages 217-220, DOI: 10.11646/zootaxa.5120.2.3, http://zenodo.org/record/638917
Euseius tripurii Kar & Karmakar 2022, sp. nov.
No full textEuseius tripurii sp. nov. (Figs 11–15, 45–50) Diagnosis. The dorsum of the new species is heavily reticulated with five pairs of solenostomes; peritreme extends until base of j3; sternal shield reticulated; other ventral plates smooth; fixed digit of chelicerae with five teeth and a pilus dentilis; spermatheca unique, tubular, half of calyx towards atrium is darkened, atrium broad and covers almost the total width of calyx, leg IV with three blunt macrosetae of hyaline tip. Description Female (n=13). Dorsum (Fig. 11). Dorsal shield 325 (322–300) long and 208 (205–210) wide, strongly reticulated, with sigilla mostly on prodorsal area, prodorsum wide with five pairs of solenostomes (gd2, gd4, gd5, gd8 and gd9); 17 pairs of setae on dorsal shield and two pairs of setae on unsclerotised cuticle: j1 28 (28–30), j3 24 (22–25), j4 11 (9–12), j5 11 (9–12), j6 12 (10–13), J2 12 (10–13), J5 5 (4–6), z2 15 (13–16), z4 20 (18–21), z5 10 (9–11), Z1 12 (11–14), Z4 15 (14–17), Z5 46 (43–46), s4 26 (24–27), S2 16 (15–18), S4 20 (19–21), S5 23 (22–25), r3 10 (9–12), R1 15 (13–16). All setae smooth, setae j1, j3, z4, Z4, Z5, s4, S2, S4 and S5 long, seta Z5 longest, all other setae medium to small. Peritreme (Fig. 11). Extending until base of j3. Venter (Fig. 12). Sternal shield square shaped, posterior margin of sternal shield indistinct and forms a groove at the lateral part of the posterior margin of sternal shield. Sternal shield 56 (55–58) long and 68 (66–69) wide at level of setae st1–st3 and st3–st3 respectively, with three pairs of setae and two pairs of poroids; one pair of setae (st4) 20 (19–22) long and conspicuous poroids on distinct metasternal plate. Distances between st2–st2 58 (56–59), st5–st5 72 (70–73). Two pairs of metapodal platelets present. Primary metapodal platelet 17 (15–18) long and 4 (3–5) wide, secondary metapodal platelet 8 (8–10). Genital shield smooth, broad at base and narrow at genital opening with irregular circular flap touching sternal loop anteriorly. Ventrianal shield smooth, slightly sclerotised at anterior part 91 (90–93) long, 45 (44–48) wide at level of ZV2 and 60 (58–62) wide at anus level with three pairs of pre-anal setae, JV1 24 (22–25), JV2 27 (26–28), ZV2 27 (26–28) and one pair of large elliptical pre-anal pores gv3 27 (26–28) apart below JV2. Membrane surrounding ventrianal shield with four pairs of setae ZV1 23 (22–24), ZV3 7 (6–9), JV4 9 (8–10) and JV5 33 (32–35); seta JV5 long and smooth. Chelicera (Fig. 13) Fixed digit 25 (23–26) long with five teeth clubbed anteriorly and distinct pilus dentilis, movable digit 22 (22–25) long with one tooth. Spermatheca (Fig. 14) Calyx, tubular, elongated 14 (12–15) long, proximal half portion of calyx towards atrium darkened, atrium broad, almost covers the total width of calyx, minor duct and major duct visible. Legs (Fig. 15) The genu, tibia and tarsus of leg IV with blunt with macrosetae of hyaline tip and following lengths: genu 33 (31–35), tibia 30 (30–33) and tarsus 60 (59–63). Chaetotactic formula of genu II 1 2/1 1/1 1 and genu III 1 1/1 2/1 1. Length of leg I 412 (410–418), leg II 270 (265–275), leg III 275 (272–280) and leg IV 385 (382–390). Leg II and III with macrosetae. Genu II 20 (19–22), tibia II 20 (19–22), tarsus II 25 (23–26); genu III 27 (25–28), tibia III 22 (20–24), tarsus III 22 (20–24). Type specimens. Holotype: female (Acarol.lab/ BCKV/8593 /2021) deposited in the NZC, Zoological Survey of India, Kolkata), collected from star fruit tree, (Averrhoa carambola) at Khowai: 24°2'24"N, 91°35'56"E, 34m AMSL, Khowai, Tripura on 6 th February, 2021; 2 paratype females (Acarol.lab/ BCKV/8594 /2021) with same collection data as holotype; 10 paratype females (Acarol.lab/ BCKV/8676–82 /2021) collected from tamarind (Tamarindus indica) from East Sikkim: 27°17'25"N, 88°35'36"E, 820m AMSL, East Skkim, Sikkim on 30 th March, 2021 and deposited in the Acarology laboratory of BCKV, Mohanpur. Etymology. The specific name tripurii refers to the name of the major tribal community of Tripura, called Tripuri. Remarks. The new species is different from all the species of Euseius by its unique spermatheca and reticulated sternal shield and also differs from all the Euseius species described and recorded from India (Kar & Karmakar, 2021). However, the new species is close to Euseius victoriensis (Womersley, 1954); E. magucii (Meyer &Rodrigues, 1966); E. noumeae (Schicha, 1979); and E. neomagucii Ueckermann, Moraes & Oliveira, 2001 in Moraes et al. (2001) by having almost similar type of spermatheca; but differs from these close species by the dorsal shield reticulated versus smooth; sternal shield reticulated versus smooth; length of dorsal setae; number of teeth in fixed and movable digit of chelicera; leg macrosetae and the shape of spermatheca. All the differences between the new species and its close species are given in Table 3. Sources of measurement: Original paper; Womersley, 1954; Meyer and Rodrigues, 1966; Schicha, 1979; Moraes et al., 2001 ‘–’: not available.Published as part of Kar, Anamika & Karmakar, Krishna, 2022, Seven new species of phytoseiid mites (Acari: Mesostigmata) from Tripura, northeastern India, pp. 213-241 in Zootaxa 5120 (2) on pages 220-222, DOI: 10.11646/zootaxa.5120.2.3, http://zenodo.org/record/638917
Phytoseius birbikrami Kar & Karmakar 2022, sp. nov.
No full textPhytoseius birbikrami sp. nov. (Figs 16–20, 51–55) Diagnosis. The new species is characterised by dorsum smooth, anteriorly striated with a distinct incision at level r3, dorsum without conspicuous large solenostome at the level z5, most of the dorsal setae serrated, setae j4, j5, j6, J5, z2 and z4 are smooth; ventrianal shield with three pairs of pre-anal setae; fixed digit of chelicerae with three teeth and movable digit single toothed, leg IV with three macrosetae. Description Female (n= 5). Dorsum (Fig. 16). Dorsal shield 263 (260–265) long and 120 (118–123) wide, smooth, anteriorly striated with five pairs of solenostomes (gd2, gd4, gd5, gd8 and gd9); 15 pairs of setae on the dorsal shield: j1 23 (21–25), j3 35 (33–36), j4 5 (4–7), j5 5 (4–7), j6 8 (7–9), J5 5 (4–7), z2 7 (6–9), z3 20 (19–21), z4 9 (8–10), z5 5 (4–7), Z4 56 (54–58), Z5 63 (62–66), s4 76 (75–78), s6 63 (61–65), r3 35 (33–36), R1 9 (8–11). Most of the setae serrated, setae j4, j5, j6, J5, z2, z4, z5 and R1 smooth. Peritreme (Fig. 16). Extending up to the bases of j1. Venter (Fig. 17). All shields are smooth. The sternal shield wider than its length, posterior margin indistinct. Sternal shield 65 (65–67) long and 84 (82–85) wide at level of setae st1–st3 and st3–st3 respectively, with three pairs of setae and two pairs of poroids; one pair of setae (st4) on metasternal platelet 23 (21–24) long. Distances between st2–st2 70 (68–72), st5–st5 61 (60–65). The base of the genital shield broad, concave posteriorly. one pair of metapodal platelet present, 19 (17–20) long and 3 (3–5) wide, Genital shield smooth, broad at the base and narrow at genital opening with irregular circular flap. Ventrianal shield smooth, sole-shaped 90 (89–92) long, 43 (42–45) wide at level of ZV2 and 40 (39–42) wide at anus level with three pairs of pre-anal setae, JV1 15 (14–17), JV2 15 (14–17), ZV2 15 (14–17) and one pair of small pre-anal pores gv3 24 (22–25) apart below seta JV2. Membrane surrounding ventrianal shield with three pairs of setae ZV1 20 (18–21), ZV3 7 (5–9) and JV5 40 (38–42); the later one is long and slightly serrated. Chelicera (Fig. 18) Fixed digit 24 (22–25) long with three teeth and distinct pilus dentilis, movable digit 25 (24–27) long with single tooth. Spermatheca (Fig. 19). Calyx elongated, funnel shaped 13 (11–15) long with conspicuous atrium, minor duct and major duct visible. Sources of measurement: Chant and Yoshida, 1992a; ‘–’: not available Leg (Fig. 20) The genu and tibia of leg IV with macrosetae of bilobed tip, macroseta of tarsus bulbous and of following lengths: genu 15 (14–17), tibia 30 (28–32) and tarsus 17 (15–18). Chaetotactic formula of genu II: 1 2/1 2/0 1 and genu III 1 2/1 1/1 1. Length of leg I 312 (310–315), leg II 234 (233–235), leg III 235 (235–238) and leg IV 340 (338–342). Type specimens. Holotype: female (Acarol.lab/ BCKV/8601 /2021) deposited in the NZC, Zoological Survey of India, Kolkata, collected from unidentified shrub at Laxmipur, Atharamura hill: 23°51'18"N, 91°42'9"E, 91m AMSL, Tripura, on 26 th August, 2019; 6 paratype females (Acarol.lab/ BCKV/8601–8602 /2019) collected from same collection data as holotype. Etymology. The new species Phytoseius birbikrami has been dedicated to the famous king of Tripura, Maharaja Birbikram Kishore Manikya Bahadur. Remarks. The new species P. birbikrami sp. nov. is different from all the nine species under the species group purseglovei (Chant &Yoshida, 1992) by having short leg macroseta with bilobed tips. However, the new species is closed to P. ikeharai Ehara, 1967, P. subtilis Wu & Li, 1984a and P. incises Wu & Li, 1984b by having similar dorsum without large conspicuous solenostomes at the level of seta z5 but differs from these close species by the presence or absence of notch at podo-scutum at the level r3 and position of seta r3, length of dorsal setae, shape of ventrianal shield, number of teeth in fixed and movable digit of chelicera, leg macrosetae and shape of spermatheca. All the differences between the new species and closed species are presented in Table 4.Published as part of Kar, Anamika & Karmakar, Krishna, 2022, Seven new species of phytoseiid mites (Acari: Mesostigmata) from Tripura, northeastern India, pp. 213-241 in Zootaxa 5120 (2) on pages 223-226, DOI: 10.11646/zootaxa.5120.2.3, http://zenodo.org/record/638917
Kar Çiçekleri
No full textEbûbekir Hâzım Tepeyran's Kar Çiçekleri (Snow Flowers) is a remarkable work emanating from one of the notable figures of Turkish literature, reflecting the social and cultural atmosphere of its era. This volume skillfully blends the author's observations from his official life and his personal experiences with a literary style, offering a profound insight into the lifestyle of the populace, their beliefs, and their social relationships. Kar Çiçekleri is more than just a literary text; it embodies Tepeyran's sensitive approach to societal issues and the human spirit, taking the reader on an emotional and intellectual journey through a collection of stories. This work is a valuable resource for readers interested in the nuances of Turkish Literature and the author's potent observational skills.Ebûbekir Hâzım Tepeyran'ın Kar Çiçekleri adlı eseri, Türk edebiyatının önemli isimlerinden birinin kaleminden çıkan, dönemin sosyal ve kültürel atmosferini yansıtan dikkate değer bir çalışmadır. Yazarın memuriyet hayatındaki gözlemlerini ve kişisel deneyimlerini edebi bir dille harmanladığı bu eser, özellikle halkın yaşam tarzı, inançları ve toplumsal ilişkileri üzerine derinlemesine bir bakış sunar. Kar Çiçekleri, sadece bir edebi metin olmanın ötesinde, Tepeyran'ın toplumsal sorunlara ve insan ruhuna dair hassas yaklaşımını sergileyen, okuyucuyu duygusal ve düşünsel bir yolculuğa çıkaran bir hikâyeler bütünüdür. Bu eser, Türk Edebiyatı'nın inceliklerini ve yazarın güçlü gözlem yeteneğini merak eden okurlar için değerli bir kaynaktır
Okiseius jainticus Kar & Karmakar 2021, sp. nov.
No full textOkiseius jainticus sp. nov. (Figs 21–25; 75–78) Female (n=7). Diagnosis. The species Okiseius jainticus sp. nov. belongs to the subgroup maritimus, by having R1 off the dorsal shield. This new species is distiguished from all the six species under this subgroup (Chant & McMurtry, 2003) by having all setae serrated except j4, j5, j6, J5, and z5 which are smooth and all the setae arise from distinct tubercles, seta S5 serrated, both the setae r3 and R1 are off the dorsal shield and each with a distinct depression at the lateral margin of dorsal shield, one pair of long narrow metapodal shields, fixed digit of chelicerae with six teeth, movable digit with three teeth, leg IV with three distinct macrosetae with bulbous tips. In addition, the genu of leg IV having all setae rod-like except macroseseta which has bulbous tip, tibia and basitarsus of leg IV having rod-like setae but ventral setae are pointed. Dorsum (Figs 21; 75). Dorsal shield 333 (332–335) long and 193 (192–195) wide, strongly reticulated, prodorsum wide, smooth lateral margin with seven pairs of pores (gd1, gd2, gd4, gd5, gd6, gd8 and gd9); 15 pairs of setae on dorsal shield and two pairs of setae on unsclerotised cuticle: j1 26 (25–28), j3 38 (38–40), j4 9 (8–10), j5 9 (8–10), j6 25 (24–25), J5 5 (4–5), z2 34 (33–35), z4 43 (42–45), z5 13 (13–15), Z1 38 (38–41), Z4 50 (49–53), Z5 70 (69–72), s4 48 (46–49), S2 49 (48–50), S5 15 (14–17), r3 41 (39–42), R1 28 (26–29). All setae serrated, except j4, j5, j6, J5 and z5 which are smooth. Peritreme (Fig. 21). Extending beyond the bases of j1. Venter (Figs 22; 78). All shields smooth. Sternal shield longer than width, posterior margin of sternal shield wavy. Sternal shield 66 (63–67) long and 76 (74–77) wide at level of setae st1 - st3 and st3-st3 respectively, with three pairs of setae and two pairs of poroids; one pair of setae (st4) on unsclerotised membrane 17 (15–18) long and conspicuous poroids on distinct metasternal plate. Distances between st2-st2 59 (58–60), st5-st5 65 (64–68). Genital shield smooth, broad at base and narrow at genital opening with irregular circular flap and almost straight posteriorly. Secondary ones fused anteried form a single pair 42 (40–44) long. Ventrianal shield smooth, pentagonal 105 (103–105) long, 56 (54–57) wide at level of ZV2 and 60 (59–62) wide at anus level with three pairs of pre-anal setae, JV1 14 (13–15), JV2 14 (13–15), ZV2 13 (12–14) and one pair of pre-anal pores gv3 26 (24–27) apart at the level JV2. Membrane surrounding ventrianal shield with four pairs of setae ZV1 18 19 (18–20), ZV3 10 (9–10), JV4 15 (13–15) and JV5 49 (48–50); seta JV5 long and serrated. Chelicera (Fig. 23). Fixed digit 25 (24–25) long with six teeth and a distinct pilus dentilis, movable digit 25 (24–25) long with three teeth. Spermatheca (Figs 24; 77). Calyx short 10 (7–11) long, arms diverging distally, atrium nodular, minor duct and major duct distinctly visible. Legs (Figs 25; 76). The genu, tibia and tarsus of leg IV with bulbous macrosetae of following lengths: genu 29 (28–30), tibia 24 (23–25) and tarsus 51 (50–53). Chaetotactic formula of genu II 1 2/1. 1/1 1 and genu III 1 1/1. 2/1 1. Length of leg I 282 (280–284), leg II 250 (246–250), leg III 236 (235–238) and leg IV 342 (341–345). Type specimens. Holotype: female (Acarol.lab/ BCKV/8384 /20) collected from Golden Himalayan raspberry (Rubus elypticus), at West Jaintia Hills, Meghalaya 25°21'44"N, 91°52' 40"E, 1933 m AMSL on 1 st May, 2019; and deposited in the NZC, Zoological Survey of India, Kolkata. Six paratype females (Acarol.lab/ BCKV/8385- 8387 /2019 ) collected from undetermined climber at Mawkdok: 25°24'30"N, 91°46'44"E, 1806 m AMSL, East Khasi Hills, with same data as holotype and deposited in the Acarological Laboratory of BCKV, Mohanpur, Nadia, West Bengal, India. Etymology. The specific name jainticus refers to the Jaintia Hills, the habitat of the new species. Remarks. Okiseius jainticus is close to O. maritimus (Ehara, 1967a); O. alniseius Wainstein & Beglyarov, 1972; O. juglandis Wang & Xu, 1985; O. tibetagramins Wu, 1987; O. tribulation Walter, 1999 and Okiseius wongi Denmark & Kolodochka, 1996, by the similar type of dorsal and ventral pattern, presence of depression at level r3, R1 and S5. The new species differs from all these close species by the length of dorsal setae, serration of dorsal setae, number of metapodal shields, denticles on fixed and movable digit of chelicerae, number of rod-like setae on leg IV etc. All the differences between the new species and its close species are presented in Table 6.Published as part of Kar, Anamika & Karmakar, Krishna, 2021, Description of eleven new species of phytoseiid mites (Acari: Mesostigmata) from Meghalaya state, north eastern India, pp. 301-354 in Zootaxa 5068 (3) on pages 320-322, DOI: 10.11646/zootaxa.5068.3.1, http://zenodo.org/record/570688
Paraamblyseius ranipoolensis Kar & Karmakar 2022, sp. nov.
No full textParaamblyseius ranipoolensis sp. nov. (Figs. 26–30; 69 –74) Diagnosis. Female dorsal shield well sclerotized, posteriorly reticulated; all the prodorsal setae are shorter than the opisthosomal setae, Z4 and Z5 are long and almost in same length, sternal shield wider than the length, reticulated; genital shield reticulated, posterior margin rounded and narrower than ventrianal shield. Ventrianal shield massive with four pairs of preanal setae, seta ZV3 absent; leg IV without macrosetae. Description. Female (n = 2). Dorsum (Fig. 26). Dorsal idiosomal chaetotaxy 10A:9B. Dorsal shield 470 (465–475) long and 220 (218–228) wide, well sclerotized, reticulated posteriorly, with marks of sigilla mostly on prodorsal area, prodorsum wide, with seven pairs of solenostomes (gd1, gd2, gd4, gd5, gd6, gd8 and gd9); 17 pairs of setae on dorsal shield and two pairs of setae on unsclerotised cuticle: j1 15 (14–17), j3 24 (23–26), j4 18 (16–19), j5 18 (16–19), j6 20 (19–22), J2 32 (30–33), J5 13 (12–15), z2 28 (28–30), z4 38 (35–39), z5 14 (13–16), Z1 40 (38–42), Z4 48 (46–50), Z5 41 (39–42), s4 40 (38–41), S2 33 (32–35), S4 22 (20–23), S5 20 (18–23), r3 18 (15–18), R1 20 (19–22). All setae smooth except seta Z4, seta Z5 is longest, all other setae short to medium. Peritreme (Fig. 26). Extending near base of j 1 . Venter (Fig. 27). Sternal shield reticulated, 56 (55–58) long, 82 (82–85) wide at level of setae st1-st3 and st3-st3 respectively, with three pairs of setae and two pairs of lyrifissures; one pair of setae (st4) 12 (11–14) on unsclerotized kidney shaped membrane; posterior margin of sternal shield concave, lateral margin strongly indented. Distances between st2-st2 65 (65–68), st5-st5 85 (83–86). Genital shield strongly reticulated, wider at posterior part and posterior margin straight. Two pairs of metapodal shields present; primary metapodal shield 42 (40–44) long, 5 (4–6) wide, secondary metapodal shield 13 (12–15) long. Ventrianal shield very broad, strongly reticulated, anterior part of triangular ventrianal plate much wider than posterior part, with four pairs of setae, ZV1 within the ventrianal shield and seta ZV3 absent, ventrianal shield 125 (120–130) long, 185 (185–190) wide at level of ZV2 setae and 68 (66–70) wide at anus level with four pairs of pre-anal setae, ZV1 25 (23–26), JV1 21 (20–22), JV2 14 (13–15), ZV2 14 (13–15) and one pair of pre-anal pores (gv3) 27 (26–28) apart. Membrane surrounding ventrianal shield with two pairs of setae JV4 11 (10–13) and JV5 26 (25–28); JV5 long and smooth. Chelicera (Fig. 28). Fixed digit 25 (24–26) long with six teeth and distinct pilus dentilis, movable digit 23 (22–24) long with two backwardly directed teeth. Spermatheca (Fig. 29).Calyx tubular, 25 (24–26) long, wider distally and gradually tapering towards conspicuous atrium, atrium covers almost the total width of the calyx, minor duct and major duct clearly visible. Leg (Fig. 30). Length of leg I: 320 (315–325), leg II 220 (218–230), leg III 230 (225–235) and leg IV 335 (335–338). Chaetotactic formula of Sge II 1-2/1 1/1-1 and Sge III 1-2/1 1/1-1. Leg IV without macrosetae. Type specimens. Holotype: female (Acarol.lab/ BCKV/8639 /2021) (deposited in the NZC, Zoological Survey of India, Kolkata), collected from debdaru or false ashoka (Monoon longifolium),at Ranipool:27°17’25”N, 88°35’36”E, 820m AMSL, East Sikkim, Sikkim on 30 th March, 2021; 1 paratype female (Acarol.lab/ BCKV/8640 /2021) from same host with same collection data as holotype. Etymology. The specific name ranipoolensis refers to the type locality “Ranipool” of east Sikkim from where this new species was collected. Remarks. Paraamblyseius ranipoolensis sp. nov. is different from all the 9 species described under the species group (Denmark, 1988) by having unique type of reticulation; ZV1 within the ventrianal shield; spermatheca tubular. However, this new species is close to P. lunatus Muma 1962; P. formosanus (Ehara, 1970); P. dinghuensis (Wu & Qian, 1982); P. foliatus Corpuz-Raros, 1994 all by having ventrianal shield with 4 pairs of setae but differs from these close species by the length of dorsal setae, width of genital shield, length and width of ventrianal shield, number of teeth on fixed and movable digit of chelicera and shape of the spermatheca. All the differences are presented in Table 6.Published as part of Kar, Anamika & Karmakar, Krishna, 2022, Description of nine new species of phytoseiid mites (Acari: Mesostigmata) from Sikkim, a north eastern States of India, pp. 201-237 in Zootaxa 5182 (3) on page 218, DOI: 10.11646/zootaxa.5182.3.1, http://zenodo.org/record/705293
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